Checklist of the Collembola: Nomina Collembola Belgica

http://www.collembola.org/publicat/belgium/collbelg.htm - Last updated on 2003.03.06 by Frans Janssens

Checklist of the Collembola: Nomina Collembola Belgica, iteratio 15

Frans Janssens, Department of Biology, University of Antwerp (RUCA), Antwerp, B-2020, Belgium

Abstract

Samenvatting

Deze voorlopige uitgave van de naamlijst van de Collembola (Hexapoda) van België is gebaseerd op literatuur onderzoek. De lijst omvat de referenties naar de originele en latere vermeldingen voor alle 195 soorten van ons land en koppelingen naar 10*10 UTM verspreidingskaarten.

Résumé Une édition provisoire de la liste des Collembola (Hexapoda) de Belgique basée sur la littérature est fournie. Cette liste inclut les références aux citations originales et à certaines citations subséquentes des 195 espèces de notre pays ainsi qu'un lien à leurs cartes de distribution sur un quadrillage UTM 10*10 km.

Keywords: Collembola, checklist, distribution maps, Belgium

Preamble

This article is a translation and a revision of
Janssens, F. 1996. Nomina Collembola Belgica. Antwerpse Vereniging voor Entomologie, Entomo-Info, Jaargang 7, 1996, nr 3, p.79-90.
Since the first edition, 82 new species have been added to the list.

Contineously being revised.

100 years of Belgian Springtails research

This provisional edition is based on a literature survey and does not yet take into account all our own observations.
The following publications give a short overview of the history of 100 year of Belgian Collembola research :

Lameere (1895) : 45 species
Marlier (1942) : 84 species (+ 5 subspecific forms)
Berbiers & Mertens (1989) : 53 species
Janssens (1996) : 113 species

The current provisional list contains 192 valid species and 3 undefined species.

Systematic species list

A provisional edition of the checklist of Collembola (Hexapoda) from Belgium based on a literature survey is provided. The list includes the references to the original and several subsequent quotations for all 195 species from our country and a link to their distribution maps on a 10*10 km UTM grid.
The checklist is a systematic list of valid taxon names with reference to the original authority, followed by the revision authority, if applicable. Each name follows the original orthography and mentions the original genus. The valid taxon names are ordered phylogenetically up to the rank of genus. The species valid names are ordered alphabethically within each genus. However, the references to the unidentified species are listed first. In each species, the references to the quotations are ordered chronologically, according to the traditional presentation of zoological catalogs (i.e. the name in this original spelling and combination; the cases of original invalid use of homonyms or incorrect spellings are indicated respectively by nec and sic). Synonyms, homonyms (cf. 'nec'), incorrect spellings (cf. 'sic') are ordered chronologically for each species. Whenever the original publication was verified, the page reference is mentioned following the publication date. (cf. 'yyyy:pp'; e.g. '1902:8'). In case the genus was not explicitly mentioned in the referenced publication, it is indicated by '...'. Publication dates are not always mentioned in such case.

Abbreviations used in the species list

?	tentative species determination
cf	confer species
e	erroneous species determination
gr	group species
h	homonym
ios	incorrect original spelling
iss	incorrect subsequent spelling
sp	undefined species

A preliminary distribution map is provided for each species. The maps record the presence of all species in a 10 x 10 km UTM grid. Records published before 1950 are indicated with open symbols, records published after 1950 are indicated with solid symbols.
The maps are produced with the mapping module of the modular software package Inventar 1.3, intended to keep track of data collected by nature monitoring projects and data of collections of natural objects. Inventar is developed by Jan de Boer and it is distributed by Mistletoe Information, Maastricht, the Netherlands.
Click on the species name to open the associated species distribution map. Note that the available distribution data is rather sparse for most species and that several records can be added still.
Click on the Collembola taxon to open a map that gives an overview of all localities.

Classis Collembola Lubbock, 1870

Ordo Poduromorpha Börner, 1913, sensu Moen, P et Ellis, WN, 1984:203

Familia Hypogastruridae Börner, 1906

Genus Hypogastrura Bourlet, 1839

Hypogastruridae Hypogastrura Dethier, M et Hubart, J-M, 2000:27 ^sp

Species assimilis (Krausbauer, 1898) Handschin, E in Dahl, F, 1929:25

Hypogastrura manubrialis ssp. assimilis Delamare Deboutteville, C, 1948:136
Hypogastrura manubrialis ssp. assimilis Marlier, G in Pasteels, 1948:59

Species burkilli (Bagnall, 1940) Weidner, 1953

Hypogastrura burkilli Dethier, M, 1998:55
Hypogastrura burkilli Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Hypogastrura burkilli Dethier, M et Hubart, J-M, 2000:27
Hypogastrura burkilli Dethier, M et Hubart, J-M, 2000:27 ^cf

Species manubrialis (Tullberg, 1869:9) Linnaniemi, 1912

Hypogastrura manubrialis var. neglecta Marlier, GJ, 1942:8
Hypogastrura manubrialis Marlier, GJ, 1942:8
Hypogastrura manubrialis Heungens, A et Van Daele, E, 1973:138
Hypogastrura manubrialis Grégoire-Wibo, C, 1981:632

Species maynardi Christiansen & Bellinger, 1980:102

Hypogastrura montana Maynard, EA, 1951:42 ^h

Species purpurescens (Lubbock, 1867) Linnaniemi, 1911

Achorutes purpurascens(sic) Willem, V, 1902:278 ^iss
Hypogastrura purpurascens(sic) Marlier, GJ, 1942:8 ^iss
Hypogastrura purpurascens(sic) Delhez, F et Houssa, M, 1969:207 ^iss
Hypogastrura purpurescens Delhez, F et Kersmaekers, M, 1973:8
Hypogastrura purpurescens Heungens, A et Van Daele, E, 1973:138
Hypogastrura purpurescens Thibaud, J-M et Massoud, Z, 1977:6
Hypogastrura purpurascens(sic) Hubart, JM, 1982:343 ^iss
Hypogastrura purpurescens Dethier, M, 1998:55
Achorutes purpurascens(sic) var. pallens Willem, V, 1902:278
Hypogastrura (Hypogastrura) purpurascens(sic) var. pallens Bonet, F, 1931:374
Hypogastrura (Hypogastrura) purpurascens(sic) var. pallescens(sic) Leruth, R, 1939:196 ^iss
Hypogastrura purpurascens(sic) var. pallescens(sic) Marlier, GJ, 1942:8
Hypogastrura purpurescens Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Hypogastrura purpurescens Dethier, M et Hubart, J-M, 2000:27

Species sahlbergi (Reuter, 1895) Handschin, 1924:91

Hypogastrura sahlbergi Marlier, GJ, 1942:8

Species vernalis (Carl, 1901) Handschin, 1924:91

Hypogastrura vernalis Heungens, A et Van Daele, E, 1973:136

Species viatica (Tullberg, 1872:50) Linnaniemi, 1911

Achorutes viaticus Lameere, A, 1895:594
Achorutes viaticus Willem, V, 1900:19
Achorutes viaticus Lameere, 1940:8
Hypogastrura viatica Marlier, GJ, 1942:8
Achorutes viaticus Geyskens, J, 1945:15
Hypogastrura viatica Heungens, A et Van Daele, E, 1973:140
Hypogastrura viatica Mertens, J et Bourgoignie, R, 1975:369
Hypogastrura viatica Mertens, J et Bourgoignie, R, 1977:41
Hypogastrura viatica Mertens, J, Coessens, R et Blancquaert, J-P, 1983:567
Hypogastrura viatica Berbiers, P et Mertens J, 1989:236
Hypogastrura viatica Janssens, F, Jacobs, W et De Boey, B, 1998:47
Hypogastrura viatica f. inermis Marlier, GJ, 1942:8

Genus Ceratophysella Börner in Brohmer, 1932

Hypogastruridae Ceratophysella Dethier, M et Hubart, J-M, 2000:27 ^sp

Species armata (Nicolet, H, 1842:57) Börner, 1932

Achorutes armatus Lameere, A, 1895:594
Achorutes armatus Willem, V, 1900:6
Achorutes armatus Willem, V, 1902:278
Hypogastrura (Hypogastrura) armata Bonet, F, 1931:373
Hypogastrura (Hypogastrura) armata Leruth, R, 1939:196
Hypogastrura armata Marlier, GJ, 1942:8
Achorutes armatus Geyskens, J, 1945:16
Ceratophysella armata Verlinden, A, 1977:114

Species bengtssoni (Ågren, 1904:2) Stach, J, 1949:142

Hypogastrura (Hypogastrura) stygia Leruth, R, 1939:196
Hypogastrura stygia Marlier, GJ, 1942:8
Hypogastrura bengtssoni Dethier, M, 1998:55
Hypogastrura bengtssoni Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Ceratophysella bengtssoni Dethier, M et Hubart, J-M, 2000:27

Species denticulata (Bagnall, 1941) Cassagnau & Rapoport, 1962

Hypogastrura denticulata Heungens, A et Van Daele, E, 1973:136
Ceratophysella denticulata Jacobs, W, 1997:71
Ceratophysella denticulata Jacobs, W, De Bruyn, L et Lust, N, 1999:57
Ceratophysella denticulata Ponge, J-F, 2000:510
Ceratophysella denticulata Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20

Genus Schaefferia Absolon, 1900

Species willemi (Bonet, 1930) Gisin, H, 1960:55

Hypogastrura (Schaefferia) willemi Bonet, 1930
Hypogastrura (Schötella(sic)) willemi Bonet, F, 1931:272
Hypogastrura (Schaefferia) willemi Bonet, F, 1931:374
Hypogastrura (Schäfferia(sic)) Willemi(sic) Denis, JR, 1937:2 ^iss
Hypogastrura (Schaefferia) Willemi(sic) Leruth, R, 1939:197 ^iss
Octomma longispina Willem, 1902:279, nec Tullberg, 1876 ^h
Schäfferia(sic) willemi Marlier, GJ, 1942:8
Schaefferia willemi Thibaud, J-M et Massoud, Z, 1977:6
Schaefferia willemi Hubart, JM, 1982:343
Hypogastrura willemi Dethier, M, 1998:38
Schaefferia willemi Dethier, M, 1998:55
Schaefferia willemi Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Schaefferia willemi Hubart, J-M et Dethier, M, 1999:167
Schaefferia willemi Dethier, M et Hubart, J-M, 2000:28
Schaefferia willemi Ponge, J-F, 2000:510

Genus Mesogastrura Bonet, 1930

Species sp. Leruth, R, 1939:197

Hypogastrura (Mesogastrura) sp. nov. Leruth, R, 1939:197

Genus Mesachorutes Absolon, 1900

Species quadriocellatus Absolon, 1900

Hypogastrura (Mesachorutes) quadriocellata Leleup, N, 1947:322
Hypogastrura (Mesachorutes) quadriocellata Marlier, G in Brien, P, 1947:158
Hypogastrura (Mesachorutes) quadriocellata Marlier, G, 1947:162
Mesachorutes marlieri Delamare Deboutteville, 1947

Genus Xenylla Tullberg, 1869

Species brevicauda Tullberg, 1869

Xenylla brevicauda Jacquemart, S, 1967:5

Species brevisimilis Stach, J, 1949:232

Xenylla brevisimilis André, H, 1977:215
Xenylla brevisimilis ssp. brevisimilis André, H, 1977:222
Xenylla brevisimilis André, HM, 1983:274

Species grisea Axelson, 1900:108

Xenylla grisea Ponge, J-F, 2000:510

Species maritima Tullberg, 1869

Xenylla maritima Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20

Species tullbergi Börner, 1903

Xenylla tullbergi André, HM, 1983:274
Xenylla tullbergi Ponge, J-F, 2000:510

Species welchi Folsom, JW, 1916:497

Xenylla welchi Delamare Deboutteville, C, 1948:133
Xenylla Welchi(sic) Marlier, G in Pasteels, 1948:59 ^iss

Genus Willemia Börner, 1901

Species anophthalma Börner, 1901

Willemia anophtalma(sic) Marlier, 1942:4 ^iss
Willemia anophthalma Ponge, J-F, 2000:510

Species aspinata Stach, 1949

Willemia aspinata Ponge, J-F, 2000:510

Genus Bonetogastrura Thibaud, 1975

Species cavicola (Börner, 1901) Christian & Schaller, 1982

Hypogastrura cavicola Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Hypogastrura cavicola Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Hypogastrura cavicola Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Hypogastrura cavicola Pollet, M et Desender, K, 1985:583
Hypogastrura cavicola Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Hypogastrura cavicola Berbiers, P et Mertens J, 1989:236
Schaefferia cavicola Dethier, M, 1998:55

Familia Odontellidae Massoud, 1967

Genus Xenyllodes Axelson, 1903

Species armatus Axelson, 1903

Xenyllodes armata(sic) Berbiers, P et Segers, H, 1991:81 ^iss
Xennylodes(sic) armata(sic) Jacobs, W, 1997:73 ^iss
Xenyllodes armata(sic) Jacobs, W, De Bruyn, L et Lust, N, 1999:59 ^iss
Xenyllodes armatus Ponge, J-F, 2000:510

Familia Brachystomellidae Stach, 1949

Genus Brachystomella Ågren, 1903

Species parvula (Schäffer, 1896:176) Stach, 1926

Brachystomella parvula Marlier, GJ, 1942:8
Brachystomella parvula Heungens, A et Van Daele, E, 1973:140
Brachystomella parvula Grégoire-Wibo, C, 1981:632

Species quadrituberculata (Stach, 1964, nec Becker, 1905) Massoud, 1967:92

Brachystomella quadrituberculata Stach, J, 1964:6 ^h

Familia Neanuridae Börner, 1901

Genus Friesea von Dalla Torre, 1895 , 1976

Species mirabilis (Tullberg, 1871) Willem, V, 1901:260

... mirabilis Lameere
Frisea(sic) mirabilis Marlier, GJ, 1942:8
Friesea mirabilis Heungens, A et Van Daele, E, 1973:138
Friesea mirabilis Jacobs, W, 1997:71
Friesea mirabilis Jacobs, W, De Bruyn, L et Lust, N, 1999:58

Species truncata Cassagnau, 1958

Friesea truncata Ponge, J-F, 2000:510

Genus Gisinea Massoud, Z, 1965:431

Species delhezi Massoud, Z, 1965:432

Gisinea delhezi Delhez, F, 1965:11
Gisinea delhezi Massoud, Z, 1967:141
Gisinea delhezi Delhez, F, Gilson, R et Hubart, J-M, 1973:411
Gizinea(sic) delhezi Delhez, F, Gilson, R et Hubart, J-M, 1973:411
Gisinea delhezi Thibaud, J-M et Massoud, Z, 1977:6
Gisinea delhesi(sic) Hubart, JM, 1982:343 ^iss
Gisinea delhezi Dethier, M, 1998:33
Gisinea delhezi Delhez, F(), Dethier, M et Hubart, J-M, 1999:35
Gisinea delhezi Hubart, J-M et Dethier, M, 1999:167
Gisinea delhezi Delhez, Dethier & Hubart, 2000 in prep

Genus Pseudachorutes Tullberg, 1871

Species parvulus Börner, 1901

Pseudachotures(sic) parvulus Verlinden, 1977:114

Species subcrassus Tullberg, 1871

Pseudachorutes subcrassus Jacquemart, S, 1967:5

Genus Micranurida Börner, 1901

Species forsslundi Gisin, 1949

Micranurida forsslundi Ponge, J-F, 2000:510

Species papillosa Axelson, 1902

Micranurida papillosa Leleup, N, 1948:10

Species pygmaea Börner, 1901:341

Anurida pygmaea Heungens, A et Van Daele, E, 1973:136
Micranurida pygmaea Ponge, J-F, 2000:510

Genus Anurida Laboulbène, 1865

Species granaria (Nicolet, H, 1847:387) Tullberg, 1869

Anurida granaria Lameere, A, 1895:595
Aphoromma granaria Willem, V, 1902:279
Anurida granaria Bonet, F, 1931:376
Anurida granaria Leruth, R, 1939:197
Anurida granaria Marlier, GJ, 1942:8
Anurida granaria Delhez, F et Kersmaekers, M, 1973:8
Anurida granaria Dethier, M, 1998:55
Anurida granaria Delhez, F(), Dethier, M et Hubart, J-M, 1999:34

Species granulata Agrell, 1943

Anurida granulata Ponge, J-F, 2000:510

Species maritima (Guérin-Méneville, 1836) Laboulbène, 1865

Anurida maritima Willem, V, 1900:31
Anurida maritima Willem, 1925
Anurida maritima Marlier, GJ, 1942:8
Anurida maritima Hansson, HG, 1998:11

Genus Neanura MacGillivray, 1893 , 1956

Species muscorum (Templeton, 1835:97) Carl, J, 1899:279

Anura muscorum Lameere, A, 1895:595
Anura muscorum Willem, V, 1900:34
Neanura muscorum Willem, V, 1902:280
Achorutes muscorum Bonet, F, 1931:376
Achorutes muscorum Leruth, R, 1939:197
Anura muscorum Lameere, A, 1940:8
Achorutes muscorum Marlier, GJ, 1942:8
Achorutes muscorum Leleup, N, 1948:10
Neanura muscorum Heungens, A et Van Daele, E, 1973:136
Neanura muscorum Verlinden, 1977:113
Neanura muscorum Berbiers, P et Mertens J, 1989:236
Neanura muscorum Berbiers, P et Segers, H, 1991:83
Neanura muscorum Jacobs, W, 1997:72
Neanura muscorum Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Neanura muscorum Dethier, M et Hubart, J-M, 2000:28 ^cf
Neanura muscorum Dethier, M et Hubart, J-M, 2000:28
Neanura muscorum Ponge, J-F, 2000:510
Neanura muscorum Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20

Familia Onychiuridae Börner, 1901

Genus Onychiuroides Bagnall, 1948

Species pseudogranulosus (Delamare Deboutteville & Gisin, 1951) Weiner, WM, 1996:186

Onychiurus pseudogranulosus Thibaud, J-M et Massoud, Z, 1977:6

Genus Deharvengiurus Weiner, WM, 1996:183

Species argus (Denis, 1924) Weiner, WM, 1996:183

Onychiurus argus Leruth, R, 1939:198
Onychiurus argus Marlier, GJ, 1942:8
Onychiurus (Onychiurus) argus Gisin, H, 1960:136
Onychiurus argus Thibaud, J-M et Massoud, Z, 1977:6
Onychiurus argus Hubart, JM, 1982:343
Deharvengiurus argus Dethier, M et Hubart, J-M, 2000:28 ^gr

Species severini (Willem, V, 1902:277) Pomorski, RJ in Christiansen, K, 2001:1

Aphorura severini Willem, V, 1902:277
Onychiurus severini Bonet, F, 1931:378
Onychiurus Severini(sic) Leruth, R, 1939:198 ^iss
Onychiurus severini Marlier, GJ, 1942:8
Onychiurus severini Stach, 1954
Onychiurus severini Godissart, J et Delhez, F, 1969:38
Onychiurus severini Delhez, Gilson & Hubart, 1973
Onychiurus severini Dethier, M, 1998:56
Onychiurus severini Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Onychiurus severini Hubart, J-M et Dethier, M, 1999:167
Protaphorura severini Dethier, M et Hubart, J-M, 2000:28 ^cf

Genus Deuteraphorura Absolon, 1901

Species inermis (Tullberg, 1871) Bellinger, PF, 2000:1

Lipura inermis Willem, V, 1900:6
Lipura fimetaria Lameere, A, 1895:595
Onychiurus fimetarius Leruth, R, 1939:198
Aphorura spelea(sic) Willem, V, 1902:278 ^iss
Onychiurus fimetarius Marlier, GJ, 1942:8
Onychiurus fimetarius Delhez, F(), Dethier, M et Hubart, J-M, 1999:34

Genus Onychiurus Gervais, 1841

Onychiuridae Onychiurus Berbiers, P et Segers, H, 1991:83 ^sp
Onychiuridae Onychiurus Jacobs, W, 1997:72 ^sp

Species ambulans (Linnæus, C, 1758:609) Gervais, 1841

Lipura ambulans Lameere, A, 1895:595
Aphorura ambulans Willem, V, 1902:277
Onychiurus ambulans Bonet, F, 1931:378
Onychiurus fimetarius Bonet, F, 1931:272 :
Onychiurus ambulans Leruth, R, 1939:198
Onychiurus ambulans Marlier, GJ, 1942:8
Aphorura ambulans Geyskens, J, 1945:15
Onychiurus circulans Dethier, M, 1998:56
Onychiurus circulans Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Onychiurus ambulans Thibaud, J-M et Massoud, Z, 1977:6
Onychiurus ambulans Hubart, JM, 1982:343
Onychiurus ambulans Delhez, F(), Dethier, M et Hubart, J-M, 1999:34

Species arans Gisin, 1952

Onychiurus arans Dethier, M et Hubart, J-M, 2000:28 ^cf

Species dissimulans Gisin, 1952

Onychiurus dissimulans Dethier, M et Hubart, J-M, 2000:28

Species sp. Leruth, R, 1939:198

Onychiurus sp. nov. Leruth, R, 1939:198

Species sublegans Gisin, 1960

Onychiurus sublegans Delhez, F et Houssa, M, 1969:206
Onychiurus sublegans Delhez, Gilson & Hubart, 1973
Onychiurus sublegans Dethier, M, 1998:56
Onychiurus sublegans Delhez, F(), Dethier, M et Hubart, J-M, 1999:34

Genus Thalassaphorura Bagnall, 1949

Species encarpata (Denis, 1931) Fjellberg, A, 1998:111

Onychiurus hortensis Heungens, A, 1970:721
Onychiurus hortensis Heungens, A et Van Daele, E, 1973:136

Genus Bionychiurus Pomorski, RJ, 1996:92

Species normalis (Gisin, 1949) Pomorski, RJ, 1996:92

Onychiurus normalis Heungens, A et Van Daele, E, 1973:140

Genus Micraphorura Bagnall, 1949

Species absoloni (Börner, 1901) Pomorski, 1996:35

Archaphorura absoloni Ponge, J-F, 2000:510

Genus Protaphorura Absolon, 1901

Species armata (Tullberg, 1869) Börner, 1909

Lipura armata Lameere, 1895:595
Lipura armata Willem, V, 1900:24
Onychiurus armatus Marlier, GJ, 1942:8
Onychiurus armatus Leleup, N, 1948:10
Onychiurus campatus Heungens, A et Van Daele, E, 1973:136
Protaphorura subuliginata Jacobs, W, 1997:73
Protaphorura subuliginata Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Onychiurus armatus Heungens, A et Van Daele, E, 1973:140
Onychiurus armatus Verlinden, 1977:113
Protaphorura armata Jacobs, W, 1997:73
Protaphorura armata Jacobs, W, De Bruyn, L et Lust, N, 1999:59

Species eichhorni (Gisin, 1954) Salmon, JT, 1964:171

Onychiurus eichhorni Heungens, A et Van Daele, E, 1973:136
Protaphorura eichhorni Ponge, J-F, 2000:510

Species octopunctata (Tullberg, 1876) Kloet, GS & Hincks, WD, 1945:5

Onychiurus octopunctatus Marlier, GJ, 1942:8
Onychiurus octopunctatus Jacquemart, S, 1967:2
Onychiurus octopuntata(sic) Jacquemart, S, 1967:4 ^iss

Species orthacantha (Handschin, 1920) Salmon, JT, 1964:173

Onychiurus orthacanthus Leleup, N, 1948:10

Species quadriocellata (Gisin, 1943) Salmon, JT, 1964:170

Onychiurus quadriocellatus Heungens, A et Van Daele, E, 1973:136

Genus Supraphorura Stach, 1954

Species furcifera (Börner, 1901) Pomorski, 1996:27

Protaphorura furcifer(sic) Thibaud, J-M et Massoud, Z, 1977:6 ^iss
Onychiurus furcifer(sic) Dethier, M, 1998:56 ^iss
Onychiurus subfurcifer Jacquemart, S, 1967:1
Onychiurus furcifer(sic) Delhez, F(), Dethier, M et Hubart, J-M, 1999:34 ^iss
Kalaphorura furcifera Ponge, J-F, 2000:510

Genus Kalaphorura Absolon, 1901

Species burmeisteri (Lubbock, J, 1873:190) Handschin, 1920

Kalaphorura Burmeisteri(sic) Leruth, R, 1939:198 ^iss
Kalaphorura burmeisteri Marlier, GJ, 1942:8
Kalaphorura burmeisteri Dethier, M et Hubart, J-M, 2000:28

Species paradoxa (Schäffer, 1900) Ellis, WN & Bellinger, PF, 1973:26

Onychiurus paradoxus Godissart, J et Delhez, F, 1969:17
Onychiurus paradoxus Dethier, M, 1998:56
Onychiurus paradoxus Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Onychiurus paradoxus Hubart, J-M et Dethier, M, 1999:168

Genus Hymenaphorura Bagnall, 1948

Species sibirica (Tullberg, 1876 tt) Bagnall, 1948:637

Hymenaphorura sibirica Ponge, J-F, 2000:510

Familia Tullbergiidae Bagnall, 1935

Genus Paratullbergia Womersley, 1930

Species callipygos (Börner, 1903) Salmon, 1964:147

Tullbergia callipygos Heungens, A, 1970:721
Tullbergia callipygas(sic) Heungens, A et Van Daele, E, 1973:136 ^iss
Paratullbergia callipygos Jacobs, W, 1997:73
Paratullbergia callipygos Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Paratullbergia callipygos Ponge, J-F, 2000:510

Genus Neotullbergia Bagnall, 1935

Species crassicuspis (Gisin, 1943) Salmon, 1964:148

Neotullbergia crassicuspis Jacobs, W, 1997:72
Neotullbergia crassicuspis Jacobs, W, De Bruyn, L et Lust, N, 1999:59

Genus Mesaphorura Börner, 1901

Species betschi Rusek, 1979

Mesaphorura betschi Ponge, J-F, 2000:510

Species hylophila Rusek, J, 1982:14

Mesaphorura hylophila Ponge, J-F, 2000:510

Species isochaeta Arbea, JI et Jordana, R, 1989:8

Mesaphorura leitzaensis Ponge, J-F, 2000:510

Species italica (Rusek, J, 1971:203) Ellis, 1974:109

Mesaphorura italica Ponge, J-F, 2000:510

Species jevanica Rusek, J, 1996:208

Mesaphorura jevanica Ponge, J-F, 2000:510

Species krausbaueri Börner, 1901

Tullbergia krausbaueri Jacquemart, S, 1967:5
Tullbergia krausbaueri Heungens, A et Van Daele, E, 1969:1106
Tullbergia krausbaueri Heungens, A, 1970:721
Tullbergia krausbaueri Heungens, A et Van Daele, E, 1973:136

Species macrochaeta Rusek, 1976

Mesaphorura macrochaeta Jacobs, W, 1997:72
Mesaphorura macrochaeta Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Mesaphorura macrochaeta Ponge, J-F, 2000:510

Species pongei Rusek, 1982

Mesaphorura pongei Ponge, J-F, 2000:510

Species sensibilis Rusek, J, 1973:79

Mesaphorura tenuisensillata Ponge, J-F, 2000:510

Species yosiii (Rusek, J, 1967:191) Rusek, 1973

Mesaphorura yosiii Dethier, M et Hubart, J-M, 2000:28 ^cf
Mesaphorura yosii(sic) Ponge, J-F, 2000:510 ^iss

Genus Stenaphorura Absolon, 1900

Species denisi Bagnall, 1935

Tullbergia denisi Heungens, A et Van Daele, E, 1973:140

Species quadrispina Börner, 1901

Stenaphorura quadrispina Jacobs, W, 1997:73
Stenaphorura quadrispina Jacobs, W, De Bruyn, L et Lust, N, 1999:59

Ordo Metaxypleona Salmon, 1964, sensu Moen, P et Ellis, WN, 1984:203

Familia Poduridae Latreille, 1804

Genus Podura Linnæus, C, 1758:608

Species aquatica Linnæus, C, 1758:609

Podura aquatica Lameere, A, 1895:594
Podura aquatica Willem, V, 1900:9
Podura aquatica Lameere, 1940:8
Podura aquatica Marlier, GJ, 1942:8
Podura aquatica Geyskens, J, 1945:15
Podura aquatica Janssens, A, 1945:10

Ordo Neelipleona Massoud Z, 1971, sensu Moen, P et Ellis, WN, 1984:203

Familia Neelidae Folsom, 1896

Genus Neelus Folsom, 1896

Species murinus Folsom, 1896

Neelus murinus Willem, V, 1902:282
Neelus murinus Bonet, F, 1931:390
Neelus murinus Leruth, R, 1939:201
Neelus murinus Marlier, GJ, 1942:10
Neelus sp. Dethier, M et Hubart, J-M, 2000:29 ^sp

Genus Megalothorax Willem, V, 1900:7

Species minimus Willem, V, 1900:9

Megalothorax minimus Willem, V, 1900:9
Megalothorax minimus Willem, V, 1900:65
Megalothorax minimus Marlier, GJ, 1942:10
Megalothorax minimus Barlet, J, 1951:142
Neelus minimus Heungens, A et Van Daele, E, 1973:136
Megalothorax minimus Blancquaert, J-P et Mertens, J, 1979:125
Megalothorax minimus Blancquaert, J-P, Coessens, R et Mertens, J, 1981:116
Neelus minimus Berbiers, P et Mertens J, 1989:237
Megalothorax minimus Jacobs, W, 1997:72
Neelus minimus Dethier, M, 1998:57
Neelus minimus Delhez, F(), Dethier, M et Hubart, J-M, 1999:35
Megalothorax minimus Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Megalothorax sp. Dethier, M et Hubart, J-M, 2000:29
Megalothorax minimus Ponge, J-F, 2000:510

Ordo Entomobryomorpha Börner, 1913, sensu Moen, P et Ellis, WN, 1984:203

Familia Isotomidae Schäffer, 1896

Genus Anurophorus Nicolet, 1842

Species laricis Nicolet, 1842:53

Anurophorus corticinus Lameere, A, 1895:595
Anurophorus Laricis(sic) Lameere, 1895:595 ^iss
Anurophorus laricis Willem, V, 1900:28
Anurophorus laricis Lameere, A, 1940:8
Anurophorus laricis Marlier, GJ, 1942:9
Anurophorus laricis André, HM, 1983:274

Genus Pseudanurophorus Stach, 1922

Species binoculatus (Kseneman, 1934) Christiansen, 1951:134

Pseudanurophorus binoculatus Ponge, J-F, 2000:510

Genus Isotomodes Axelson, 1907

Species productus (Axelson, 1906:6) Handschin, E in Dahl, F, 1929:57

Isotomodes productus Heungens, A et Van Daele, E, 1973:140

Genus Folsomides Stach, 1922

Species angularis (Axelson, 1905) Gisin, 1942

Folsomides angularis Jacquemart, S, 1967:5

Genus Cryptopygus Willem, 1901

Species bipunctatus (Axelson, 1903) Dunger, 1977:184

Isotomina bipunctata Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Isotomina bipunctata Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Isotomina bipunctata Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Isotomina bipunctata Berbiers, P et Mertens J, 1989:236
Isotomina bipunctata Berbiers, P et Segers, H, 1991:83

Species thermophilus (Axelson, 1900:114) Massoud & Rapoport, 1968

Cryptopygus thermophilus Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20

Genus Folsomia Willem, 1902

Isotomidae Folsomia Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20 ^sp

Species candida Willem, V, 1902:280

Folsomia fimetaria Bonet, F, 1931:380 :
Folsomia candida Delhez, F et Houssa, M, 1969:207
Folsomia candida Delhez, Gilson Hubart, 1973
Folsomia candida Thibaud, J-M et Massoud, Z, 1977:6
Folsomia candida Hubart, JM, 1982:343
Folsomia candida Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Folsomia candida Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Folsomia candida Berbiers, P et Mertens J, 1989:236
Folsomia candida Dethier, M, 1998:56
Folsomia candida Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Folsomia candida Dethier, M et Hubart, J-M, 2000:19
Desoria(sic) candida Dethier, M et Hubart, J-M, 2000:28

Species diplophthalma (Axelson, 1902) Axelson, 1907

Folsomia diplophtalma(sic) Jacobs, W, 1997:71 ^iss

Species fimetaria (Burmeister, 1835) Willem, 1925:626

Folsomia fimetaria Leruth, R, 1939:199
Folsomia fimetaria Marlier, GJ, 1942:9
Folsomia fimetaria Leleup, N, 1948:10
Folsomia fimetaria Heungens, A, 1970:721
Folsomia fimetaria Heungens, A et Van Daele, E, 1973:136

Species fimetarioides (Axelson, 1903) Handschin, 1929:59

Folsomia fimetarioides Heungens, A et Van Daele, E, 1973:136
Folsomia fimetarioides Jacobs, W, 1997:71
Folsomia fimetarioides Jacobs, W, De Bruyn, L et Lust, N, 1999:57

Species manolachei Bagnall, 1939

Folsomia nana Heungens, A et Van Daele, E, 1973:140
Folsomia manolachei Dethier, M et Hubart, J-M, 2000:28
Folsomia manolachei Ponge, J-F, 2000:510

Species penicula Bagnall, 1939

Folsomia multiseta Heungens, A, 1970:721
Folsomia multiseta Heungens, A et Van Daele, E, 1973:136

Species quadrioculata (Tullberg, 1871) Handschin, 1929:58

... quadrioculata Willem
Folsomia quadrioculata Leruth, R, 1939:199
Folsomia quadrioculata Marlier, GJ, 1942:9
Folsomia quadrioculata Jacquemart, S, 1967:4
Folsomia quadrioculata Heungens, A et Van Daele, E, 1973:136
Folsomia quadrioculata Grégoire-Wibo, C, 1974:207
Folsomia quadrioculata Grégoire-Wibo, C, 1980:43
Folsomia quadrioculata Berbiers, P et Segers, H, 1991:83
Folsomia quadrioculata Jacobs, W, 1997:71
Folsomia quadrioculata Jacobs, W, De Bruyn, L et Lust, N, 1999:58
Folsomia quadrioculata Dethier, M et Hubart, J-M, 2000:28
Folsomia quadrioculata Ponge, J-F, 2000:510
Folsomia quadrioculata Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Folsomia quadrioculata De Boey, B, 2002:28

Species sexoculata (Tullberg, 1871) Handschin, 1929:57

Isotoma sexoculata Willem, 1900
Folsomia sexoculata Marlier, GJ, 1942:9

Species similis Bagnall, 1939:57

Folsomia similis Heungens, A et Van Daele, E, 1973:140

Species spinosa Kseneman, 1936

Folsomia spinosa Jacobs, W, 1997:71
Folsomia spinosa Jacobs, W, De Bruyn, L et Lust, N, 1999:58

Genus Folsomina Denis, 1931

Species onychiurina Denis, 1931

Folsomia(sic) onychiurina Heungens, A et Van Daele, E, 1973:140

Genus Proisotoma Börner, 1901

Species minima (Absolon, 1901) Handschin, 1929:62

Proisotoma minima Heungens, A et Van Daele, E, 1973:136
Proisotoma minima Ponge, J-F, 2000:510

Species minuta (Tullberg, 1871) Börner, 1903

Proisotoma minuta Marlier, GJ, 1942:9
Proistoma(sic) minuta Leleup, N, 1948:10
Isotoma stagnalis Willem, V, 1900:28
Isotoma stagnalis Willem, V, 1900:6
Isotoma stagnalis Willem, V, 1901:193
Isotoma stagnalis var? tenebricola Willem, V, 1900:29
Isotoma tenebricola Willem, V, 1900:29
Isotoma stagnalis var. tenebricola Willem, V, 1901:195
Proisotoma minuta Heungens, A et Van Daele, E, 1969:1106
Proisotoma minuta Heungens, A, 1970:721
Proisotoma minuta Heungens, A et Van Daele, E, 1973:136
Proisotoma minuta Verlinden, A, 1977:113

Genus Isotomurus Börner, 1903

Isotomidae Isotomurus De Boey, B, 2002:56 ^sp

Species maculatus (Schäffer, 1896) Ågren, 1903

Isotomurus palustris f. maculata Verlinden, A, 1977:114

Species palustris (Müller OF, 1776) Börner, 1903

Isotoma palustris Lameere, A, 1895:594
Isotomurus palustris Marlier, GJ, 1942:9
Isotoma palustris Geyskens, 1945:16
Isotomurus palustris Heungens, A et Van Daele, E, 1973:136
Isotomurus palustis(sic) Heungens, A et Van Daele, E, 1973:141 ^iss
Isotomurus palustris Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Isotomurus palustris Grégoire-Wibo, C, 1983:41
Isotomurus palustris Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Isotoma(sic) palustris Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Isotomurus palustris Pollet, M et Desender, K, 1986:962
Isotomurus palustris Pollet, M et Desender, K, 1988:1120
Isotomurus palustris Berbiers, P et Mertens, J, 1989:280
Isotomurus palustris Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Isotomurus palustris Berbiers, P et Mertens J, 1989:236
Isotomurus palustris Berbiers, P et Segers, H, 1991:83
Isotomurus palustris Jacobs, W, 1997:72
Isotomurus palustris Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Isotomurus sp. Dethier, M et Hubart, J-M, 2000:29
Isotomurus palustris De Boey, B, 2002:29

Species plumosus Bagnall, 1940

Isotomurus plumosus De Boey, B, 2002:29

Genus Isotoma Bourlet, 1839

Species intermedia Schött, 1902

Isotoma intermedia Marlier, GJ, 1942:5

Species viridis Bourlet, 1839 Opinion 291

Isotoma viridis Lameere, A, 1895:594
Isotoma viridis Willem, V, 1900:39
Isotoma (Isotoma) viridis Leruth, R, 1939:199
Isotoma viridis Marlier, GJ, 1942:9
Isotoma viridis Geyskens, J, 1945:16
Isotoma viridis Jacquemart, S, 1967:5
Isotoma viridis Heungens, A et Van Daele, E, 1973:138
Isotoma viridis Verlinden, 1977:114
Isotoma viridis Grégoire-Wibo, C, 1981:632
Isotoma viridis Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Isotoma viridis Grégoire-Wibo, C, 1983:41
Isotoma viridis Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Isotoma viridis Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Isotoma viridis Pollet, M et Desender, K, 1986:962
Isotoma viridis Pollet, M et Desender, K, 1988:1120
Isotoma viridis Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Isotoma viridis Berbiers, P et Mertens J, 1989:236
Isotoma viridis Berbiers, P et Segers, H, 1991:83
Isotoma viridis Viskens, G et Bruers, J, 1994:29
Isotoma viridis Janssens, F, 1995:19
Isotoma viridis Jacobs, W, 1997:72
Isotoma viridis Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Isotoma viridis Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Isotoma viridis De Boey, B, 2002:28
Isotoma viridis var. riparia Marlier, GJ, 1942:3

Genus Isotomiella Bagnall, 1939

Isotomidae Isotoma Leleup, N, 1948:10 ^sp

Species minor (Schäffer, 1896) Yosii, 1939

Isotoma (Isotoma) minor Leruth, R, 1939:199
Isotomiella minor Heungens, A, 1970:721
Isotomiella minor Heungens, A et Van Daele, E, 1973:136
Isotomiella minor Jacobs, W, 1997:72
Isotomiella minor Dethier, M, 1998:56
Isotomiella minor Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Isotomiella minor Ponge, J-F, 2000:510

Genus Agrenia Börner, 1906

Species bidenticulata (Tullberg, 1876) Börner, 1906

Agrenia bidenticulata Marlier, GJ, 1942:9

Genus Desoria Nicolet in Desor, 1841

Species olivacea (Tullberg, 1871)

Isotoma olivacea Marlier, GJ, 1942:9
Isotoma olivacea Heungens, A et Van Daele, E, 1973:138
Isotoma olivacea Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Isotoma olivacea Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Isotoma olivacea Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Isotoma olivacea Berbiers, P et Mertens, J, 1989:280
Isotoma olivacea Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Isotoma olivacea Berbiers, P et Mertens J, 1989:236

Species tigrina Nicolet, 1842:59

Desoria tigrina Dethier, M et Hubart, J-M, 2000:28
Isotoma tigrina De Boey, B, 2002:28

Species violacea (Tullberg, 1876)

Isotoma violacea Marlier, GJ, 1942:9
Isotoma violacea Heungens, A et Van Daele, E, 1973:138
Isotoma violacea Dethier, M et Hubart, J-M, 2000:29

Genus Paranurophorus Denis, 1929

Species simplex Denis, 1929

Coloburella simplex Heungens, A et Van Daele, E, 1973:140

Genus Parisotoma Bagnall, 1940

Species agrelli (Delamare Deboutteville, 1950)

Isotoma agrelli Jacobs, W, 1997:71
Isotoma agrelli Jacobs, W, De Bruyn, L et Lust, N, 1999:58

Species notabilis (Schäffer, 1896) Bagnall, 1940

Isotoma notabilis Marlier, GJ, 1942:9
Isotoma notabilis Jacquemart, S, 1967:5
Isotoma notabilis Heungens, A et Van Daele, E, 1969:1106
Isotoma notabilis Heungens, A, 1970:721
Isotoma notabilis Heungens, A et Van Daele, E, 1973:136
Isotoma notabilis Grégoire-Wibo, C, 1983:41
Parisotoma notabilis Dethier, M et Hubart, J-M, 2000:29
Parisotoma notabilis Ponge, J-F, 2000:510
Parisotoma notabilis Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20

Species tenuicornis (Axelson, 1903) Kloet, GS & Hincks, WD, 1945:7

Isotoma tenuicornis Marlier, GJ, 1942:4

Genus Pseudisotoma Handschin, 1924

Species monochaeta (Kos, 1942) Jacquemart, S, 1967:4

Pseudisotoma monochaeta Jacquemart, S, 1967:4

Species sensibilis (Tullberg, 1876) Agrell, 1939

Pseudisotoma sensibilis Marlier, GJ, 1942:9
Pseudisotoma sensibilis Jacquemart, S, 1967:4
Isotoma (Pseudisotoma) sensibilis Heungens, A et Van Daele, E, 1973:136
Pseudisotoma sensibilis Janssens, F in Bruers, J et Viskens, G, 2000:15
Pseudisotoma sensibilis Ponge, J-F, 2000:510

Genus Vertagopus Börner, 1906

Species arboreus (Linnæus, C, 1758:609) Ågren, 1903

Vertagopus arborea(sic) André, HM, 1983:274 ^iss

Species cinereus (Nicolet, 1842:60) Marlier, GJ, 1942:9

Isotoma cinerea Lameere, A, 1895:594
Isotoma cinerea Willem, 1900
Vertagopus cinerea(sic) Marlier, GJ, 1942:9 ^iss

Familia Actaletidae Börner, 1902

Genus Actaletes Giard, 1889

Species neptuni Giard, 1889

Actaletes Neptuni(sic) Willem, V, 1900:6 ^iss
Actaletes neptuni Willem, V, 1900:50
Actaletes Neptuni(sic) Willem, V, 1901:474 ^iss
Actaletes neptuni Marlier, GJ, 1942:10

Familia Entomobryidae Schött, 1891

Genus Orchesella Templeton, 1835

Species bifasciata Nicolet, 1842:80

Orchesella bifasciata Berbiers, P et Mertens J, 1989:236

Species cincta (Linnæus, C, 1758:609) Templeton, 1835

Orchesella cincta Lameere, A, 1895:593
Orchesella cincta Willem, V, 1900:49
Orchesella cincta Willem, V, 1901:654
Orchesella cincta Marlier, GJ, 1942:10
Orchesella cincta Geyskens, J, 1945:16
Orchesella cincta Thibaud, J-M et Massoud, Z, 1977:6
Orchesella cincta Mertens, J et Blancquaert, J-P, 1980:301
Orchesella cincta Mertens, Coessens & Blancquaert, 1982:9
Orchesella cincta André, HM, 1983:274
Orchesella cincta Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Orchesella cincta van Straalen, NM, 1986:71
Orchesella cincta Berbiers, P et Mertens, J, 1989:280
Orchesella cincta Berbiers, P et Mertens J, 1989:236
Orchesella cincta Berbiers, P et Segers, H, 1991:82
Orchesella cincta Janssens, F, 1995:19
Orchesella cincta Janssens, F, 1995:140
Orchesella cincta Janssens, F, 1996:53
Orchesella cincta Janssens, F in Bruers, J et Viskens, G, 2000:15
Orchesella cincta Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Orchesella cincta De Boey, B, 2002:30

Species flavescens (Bourlet, 1839) Nicolet, 1847:365

Orchesella rufescens Lameere, 1895:593
Orchesella rufescens Willem, V, 1900:49
Orchesella rufescens Willem, V, 1901:654,
Orchesella flavescens Marlier, GJ, 1942:10
Orchesella flavescens Berbiers, P et Mertens J, 1989:236
Orchesella flavescens De Boey, B, 2002:30
Orchesella flavescens var. pallida Marlier, GJ, 1942:5

Species quinquefasciata (Bourlet, 1842) Bourlet, 1843

Orchesella quinquefasciata Marlier, GJ, 1942:10
Orchesella quinquefasciata Berbiers, P et Mertens J, 1989:236

Species villosa (Linnæus, C, 1767:1014) Nicolet, 1842:78

Orchesella villosa Lameere, A, 1895:593
Orchesella villosa Willem, V, 1900:49
Orchesella villosa Willem, 1901:654
Orchesella villosa Lameere, A, 1940:8
Orchesella villosa Marlier, GJ, 1942:10
Orchesella villosa Verlinden, A, 1977:114
Orchesella villosa Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Orchesella villosa Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Orchesella villosa Berbiers, P et Mertens J, 1989:236
Orchesella villosa Berbiers, P et Segers, H, 1991:83
Orchesella villosa Viskens, G et Bruers, J, 1994:29
Orchesella villosa Janssens, F, 1996:53
Orchesella villosa Dethier, M et Hubart, J-M, 2000:29
Orchesella villosa Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Orchesella villosa De Boey, B, 2002:30

Genus Heteromurus Wankel, 1860 , 1976

Species nitidus (Templeton, 1835) Handschin, 1929:93

Templetonia crystallina Lameere, 1895:593
Heteromurus (Heteromurus) nitidus Bonet, F, 1931:387
Heteromurus (Heteromurus) nitidus Leruth, R, 1939:200
Heteromurus margaritarius Willem, V, 1902:281
Heteromurus nitidus Marlier, GJ, 1942:10
Heteromurus nitidus Leleup, N, 1948:10
Heteromurus nitidus Delhez, F et Houssa, M, 1969:207
Heteromurus nididus(sic) Godissart, J et Delhez, F, 1969:38 ^iss
Heteromurus nitidus Delhez, F et Kersmaekers, M, 1973:8
Heteromurus nitidus Delhez, Gilson & Hubart, 1973
Heteromurus nitidus Heungens, A et Van Daele, E, 1973:140
Heteromurus nitidus Thibaud, J-M et Massoud, Z, 1977:6
Heteromerus(sic) nitidus Hubart, JM, 1982:343
Heteromurus nitidus Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Heteromurus nitidus Berbiers, P et Mertens J, 1989:236
Heteromurus nitidus Jacobs, W, 1997:71
Heteromurus nitidus Dethier, M, 1998:56
Heteromurus nitidus Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Heteromurus nitidus Jacobs, W, De Bruyn, L et Lust, N, 1999:58
Heteromurus nitidus Dethier, M et Hubart, J-M, 2000:29
Heteromurus nitidus De Boey, B, 2002:31

Genus Entomobrya Rondani, 1861

Species albocincta (Burmeister, 1835) Rondani, 1861

Entomobrya albocincta Lameere, A, 1895:592
Entomobrya albocincta Marlier, GJ, 1942:9
Entomobrya albocincta Berbiers, P et Mertens, J, 1989:280
Entomobrya albocincta Berbiers, P et Mertens J, 1989:236
Entomobrya albocincta Berbiers, P et Segers, H, 1991:82

Species arborea (Tullberg, 1871) Lameere, 1895:592

Entomobrya arborea Lameere, 1895:592
Entomobrya arborea Marlier, GJ, 1942:9

Species corticalis (Nicolet, H, 1842:72) Rondani, 1861

Entomobrya corticalis Lameere, 1895:592
Entomobrya corticalis Willem, V, 1900:50
Entomobrya corticalis Marlier, GJ, 1942:9
Entomobrya corticalis Berbiers, P et Mertens, J, 1989:280
Entomobrya corticalis Berbiers, P et Mertens J, 1989:236
Entomobrya corticalis Berbiers, P et Segers, H, 1991:82

Species lanuginosa (Nicolet, 1842:74) Rondani, 1861

Entomobrya lanuginosa Lameere, A, 1895:592
Entomobrya lanuginosa Heungens, A et Van Daele, E, 1973:136
Entomobrya lanuginosa Dethier, M et Hubart, J-M, 2000:29

Species marginata (Tullberg, 1871) Schött, H, 1891:17

Entomobrya marginata Marlier, GJ, 1942:9
Entomobrya marginata Heungens, A et Van Daele, E, 1973:140

Species multifasciata (Tullberg, 1871) Brook, 1883

Entomobrya decemfasciata Marlier, GJ, 1942:9
Entomobrya multifasciata Lameere, A, 1895:592
Entomobrya multifasciata Willem, V, 1900:50
Entomobrya multifasciata Marlier, GJ, 1942:9
Entomobrya multifasciata Heungens, A et Van Daele, E, 1973:138
Entomobrya multifasciata André, H, 1977:217
Entomobrya multifasciata Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Entomobrya multifasciata Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Entomobrya multifasciata Berbiers, P et Mertens, J, 1989:280
Entomobrya multifasciata Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Entomobrya multifasciata Berbiers, P et Mertens J, 1989:236
Entomobrya multifasciata Janssens, F, 1995:19

Species muscorum (Nicolet, H, 1842:75) Rondani, 1861

Entomobrya muscorum Lameere, 1895:592
Entomobrya muscorum Marlier, GJ, 1942:9
Entomobrya muscorum Berbiers, P et Mertens J, 1989:236
Entomobrya muscorum De Boey, B, 2002:29

Species nicoleti (Lubbock, 1868) Brook, 1884

Entomobrya nicoleti Berbiers, P et Mertens J, 1989:236
Entomobrya nicoleti Janssens, F, 1995:19

Species nivalis (Linnæus, C, 1758:609) Rondani, 1861

Entomobrya nivalis Lameere, A, 1895:592
Entomobrya nivalis Marlier, GJ, 1942:9
Entomobrya nivalis Geyskens, J, 1945:16
Entomobrya nivalis Heungens, A et Van Daele, E, 1973:140
Entomobrya nivalis Verlinden, A, 1977:114
Entomobrya nivalis André, HM, 1983:274
Entomobrya nivalis Berbiers & Mertens, 1989:234
Entomobrya nivalis Berbiers, P et Mertens, J, 1989:280
Entomobrya nivalis Berbiers, P et Mertens J, 1989:236
Entomobrya nivalis Viskens, G et Bruers, J, 1994:29
Entomobrya nivalis Janssens, F, 1996:53
Entomobrya nivalis Ponge, J-F, 2000:510
Entomobrya nivalis Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Entomobrya nivalis De Boey, B, 2002:30

Species quinquelineata Börner, C, 1901:69

Entomobrya quinquelineata Marlier, GJ, 1942:9

Species spectabilis Reuter, 1890

Entomobrya spectabilis Berbiers, P et Mertens J, 1989:236

Genus Sinella Brook, 1882

Species tenebricosa Folsom, JW, 1902:365

Sinella caeca Marlier, GJ, 1942:9
Sinella tenebricosa Bellinger, PF, 2000:1

Genus Willowsia Shoebotham, 1917

Species buski (Lubbock, 1869) Paclt, 1962:56

Seira Buski(sic) Lameere, A, 1895:593 ^iss
Sira(sic) buski Marlier, GJ, 1942:9
Willowsia buski Heungens, A, 1970:721
Willowsia buski Heungens, A et Van Daele, E, 1973:136
Willowsia buski Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20

Species platani (Nicolet, 1842:72) Palissa, 1959

Willowsia platani André, HM, 1983:274
Willowsia platani Berbiers, P et Mertens J, 1989:236

Genus Drepanosira Bonet, 1942

Species pallida Brown, JM, 1925:209

Seira pallida Brown, JM, 1925:209

Genus Seira Lubbock, 1869

Species domestica (Nicolet, H, 1842:76) Lubbock, 1873:144

... domesticus Willem
Seira elongata Lameere, A, 1895:593
Sira(sic) elongata Willem, V, 1900:47
Lepidocyrtinus domesticus Marlier, GJ, 1942:9
Lepidocyrtinus domesticus van den Bruel, WE in Collart, A, 1944:210
Sira(sic) domestica Geyskens, J, 1945:16
Lepidocyrtinus domesticus van den Bruel, WE, 1945:36
Seira domestica Heungens, A et Van Daele, E, 1969:1106
Seira domestica Heungens, A, 1970:721
Seira domestica Heungens, A et Van Daele, E, 1973:136
Seira domestica Berbiers, P et Mertens J, 1989:236
Seira domestica Dethier, M, 1998:57
Seira domestica Delhez, F(), Dethier, M et Hubart, J-M, 1999:35

Genus Lepidocyrtus Bourlet, 1839

Entomobryidae Lepidocyrtus Grégoire-Wibo, C, 1981:631 ^sp
Entomobryidae Lepidocyrtus Dethier, M et Hubart, J-M, 2000:29 ^sp

Species curvicollis (Bourlet, 1839) Bourlet, 1841

Lepidocyrtus curvicollis Lameere, A, 1895:591
Lepidocyrtus curvicollis Leruth, R, 1939:199
Lepidocyrtus Börneri(sic) Willem, V, 1902:281 ^ios
Lepidocyrtus sp. var. Börneri(sic) Willem, V, 1902:282 ^?
Lepidocyrtus börneri(sic) Bonet, F, 1931:383 ^iss
Lepidocyrtus curvicollis Marlier, GJ, 1942:9
Lepidocyrtus (Lepidocyrtus) curvicollis Yosii, 1959
Lepidocyrtus curvicollis Delhez, F et Kersmaekers, M, 1973:8
Lepidocyrtus curvicolis(sic) Thibaud, J-M et Massoud, Z, 1977:6 ^iss
Lepidocyrtus curvicollis Berbiers, P et Segers, H, 1991:83
Lepidocyrtus curvicollis Jacobs, W, 1997:72
Lepidocyrtus curvicollis Dethier, M, 1998:56
Lepidocyrtus curvicollis Delhez, F(), Dethier, M et Hubart, J-M, 1999:35
Lepidocyrtus curvicollis Dethier, M et Hubart, J-M, 2000:29

Species cyaneus Tullberg, 1871

Lepidocyrtus cyaneus Leruth, R, 1939:199
Lepidocyrtus cyaneus Marlier, GJ, 1942:9
Lepidocyrtus cyaneus Jacquemart, S, 1967:5
Lepidocyrtus cyaneus Heungens, A et Van Daele, E, 1973:136
Lepidocyrtus cyaneus Verlinden, 1977:114
Lepidocyrtus cyaneus Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Lepidocyrtus cyaneus Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Lepidocyrtus cyaneus Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Lepidocyrtus cyaneus Berbiers, P et Mertens, J, 1989:280
Lepidocyrtus cyaneus Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Lepidocyrtus cyaneus Berbiers, P et Mertens J, 1989:236

Species lignorum (Fabricius O, 1775) Gervais, 1844

Lepidocyrtus lignorum Willem, V, 1900:48
Lepidocyrtus lignorum Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Lepidocyrtis(sic) lignorum Janssens, F in Bruers, J et Viskens, G, 2000:15 ^iss
Lepidocyrtus lignorum Ponge, J-F, 2000:510
Lepidocyrtus lignorum Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Lepidocyrtus lignorum De Boey, B, 2002:31

Species pallidus Reuter, 1890

Lepidocyrtus pallidus Heungens, A, 1967:243
Lepidocyrtus cyaneus Heungens, A, 1967:243 :
Lepidocyrtus pallidus Heungens, A et Van Daele, E, 1973:140

Species paradoxus Uzel, 1890

Lepidocyrtus paradoxus Heungens, A et Van Daele, E, 1973:140

Species violaceus (Fourcroy, 1785) Lubbock, J, 1873:154

Lepidocyrtus violaceus Lameere, 1895:591
Lepidocyrtus violaceus van den Bruel, WE in de Jonghe d'Ardoye, E, 1937:190
Lepidocyrtus cyanus(sic) van den Bruel, WE in de Jonghe d'Ardoye, E, 1937:190 ^iss
Lepidocyrtus cyanus(sic) van den Bruel, WE in de Jonghe d'Ardoye, E, 1937:321 ^iss
Lepidocyrtus violaceus Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20 ^cf
Lepidocyrtus cyaneus Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:22 :

Species lanuginosus Gmelin, 1788

Lepidocyrtus lanuginosus Lameere, 1895:591
Lepidocyrtus lanuginosus Leruth, R, 1939:200
Lepidocyrtus lanuginosus Marlier, GJ, 1942:9
Lepidocyrtus lanuginosus Jacquemart, S, 1967:5
Lepidocyrtus lanuginosus Heungens, A, 1970:721
Lepidocyrtus lanuginosus Heungens, A et Van Daele, E, 1973:136
Lepidocyrtus lanuginosus Verlinden, 1977:114
Lepidocyrtus lanuginosus Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Lepidocyrtus lanuginosus Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Lepidocyrtus lanuginosus Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Lepidocyrtus lanuginosus Berbiers, P et Mertens J, 1989:236
Lepidocyrtus lanuginosus Viskens, G et Bruers, J, 1994:29
Lepidocyrtus lanuginosus Jacobs, W, 1997:72
Lepidocyrtus lanuginosus Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Lepidocyrtus lanuginosus Ponge, J-F, 2000:510

Genus Pseudosinella Schäffer, 1897

Entomobryidae Pseudopsinella(sic) Leleup, N, 1948:10 ^sp
Entomobryidae Pseudosinella Dethier, M et Hubart, J-M, 2000:29 ^sp

Species alba (Packard, 1873) Handschin, 1929:90

Pseudosinella alba Marlier, GJ, 1942:10
Pseudopsinella(sic) alba Leleup, N, 1948:10
Pseudosinella alba Jacobs, W, 1997:73
Pseudosinella alba Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Pseudosinella alba Ponge, J-F, 2000:510

Species decipiens Denis, 1924

Pseudosinella decipiens Dethier, M et Hubart, J-M, 2000:29 ^cf

Species duodecimocellata Handschin, 1928

Pseudosinella duodecimoculata(sic) Marlier, GJ, 1942:9 ^iss

Species insubrica Gisin, H et da Gama, MM, 1969:292

Pseudosinella monoculata Thibaud, J-M et Massoud, Z, 1977:7

Species ksenemani Gisin, 1944

Pseudosinella ksenemani Jacobs, W, 1997:73

Species mauli Stomp, N, 1972:279

Pseudosinella mauli Ponge, J-F, 2000:510

Species octopunctata Börner, 1901

Pseudosinella octopunctata Marlier, GJ, 1942:10
Pseudosinella octopunctata Jacobs, W, 1997:73

Species petterseni Börner, 1901

Pseudosinella petterseni Heungens, A et Van Daele, E, 1973:140

Species vandeli Denis, 1923

Pseudosinella vandeli Thibaud, J-M et Massoud, Z, 1977:6
Pseudosinella vandeli Hubart, JM, 1982:343
Pseudosinella vandeli Dethier, M, 1998:38
Pseudosinella vandeli Delhez, F(), Dethier, M et Hubart, J-M, 1999:35
Pseudosinella vandeli Hubart, J-M et Dethier, M, 1999:167

Familia Cyphoderidae Börner, 1913

Genus Cyphoderus Nicolet, [1842]

Species albinus (Nicolet, 1842:67) Nicolet, 1847:383

Cyphoderus albinos(sic) Lameere, 1895:591 ^iss
Cyphoderus albinus Willem, V, 1900:49
Cyphoderus albinus Lameere, A, 1940:8
Cyphoderus albinus Marlier, GJ, 1942:10
Cyphoderus albinos(sic) Geyskens, J, 1945:16
Cyphoderus albinus Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Cyphoderus albinus Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Cyphoderus albinus Berbiers, P et Mertens J, 1989:236
Cyphoderus albinus Janssens, F, 1995:80

Familia Oncopoduridae Carl & Lebedinsky, 1905

Genus Oncopodura Carl & Lebedinsky, 1905

Entomobryidae Oncopodura Delhez, F(), Dethier, M et Hubart, J-M, 1999:44
Oncopoduridae Oncopodura Hubart, J-M et Dethier, M, 1999:168 ^sp
Oncopoduridae Oncopodura Dethier, M et Hubart, J-M, 2000:29 ^sp

Species crassicornis Shoebotham, 1911

Oncopodura crassicornis Heungens, A et Van Daele, E, 1973:140

Species reyersdorfensis Stach, 1936

Oncopodura reyersdorfensis Delhez, F(), Dethier, M et Hubart, J-M, 1999:35
Oncopodura reyersdorfensis Hubart, J-M et Dethier, M, 1999:168
Oncopodura reyersdorfensis Delhez, Dethier & Hubart, 2000 in prep

Species sp. (Leruth, R, 1939:201) Marlier, GJ, 1942:10

Oncopodura sp. nov. Leruth, R, 1939:201
Oncopodura sp. Marlier, GJ, 1942:10

Familia Tomoceridae Schäffer, 1896

Genus Tomocerus Nicolet, [1842], n.c. , 1954

Tomoceridae Tomocerus Dethier, M et Hubart, J-M, 2000:29 ^sp

Species minor (Lubbock, 1862) Schäffer, 1900

Tomocerus plumbeus Willem, V, 1900:44
Tomocerus plumbeus Lameere, 1940:8
Tomocerus plumbeus Geyskens, 1945:16
Tomocerus tridentiferus Willem, V, 1900:6
Tomocerus tridentiferus Willem, V, 1902:281
Tomocerus minor Bonet, F, 1931:388
Tomocerus (Tomocerus) minor Leruth, R, 1939:200
Tomocerus minor Marlier, GJ, 1942:10
Tomocerus minor Delhez, F et Kersmaekers, M, 1973:8
Tomocerus minor Delhez, Gilson & Hubart, 1973
Tomocerus minor Heungens, A et Van Daele, E, 1973:138
Tomocerus minor Thibaud, J-M et Massoud, Z, 1977:6
Tomocerus minor Verlinden, 1977:114
Tomocerus minor Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Tomocerus minor Berbiers, P et Mertens J, 1989:236
Tomocerus minor Janssens, F, 1996:53
Tomocerus minor Jacobs, W, 1997:73
Tomocerus minor Dethier, M, 1998:57
Tomocerus minor Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Tomocerus minor Hubart, J-M et Dethier, M, 1999:168
Tomocerus minor Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Tomocerus minor Dethier, M et Hubart, J-M, 2000:19
Tomocerus minor Dethier, M et Hubart, J-M, 2000:29
Tomocerus minor Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Tomocerus minor De Boey, B, 2002:31

Species vulgaris (Tullberg, 1871) Tullberg, 1876

Tomocerus niger Lameere, A, 1895:591
Tomocerus vulgaris Lameere, A, 1895:591
Tomocerus vulgaris Willem, V, 1900:6
Tomocerus plumbeus Lameere, 1895:591, nec Lubbock, 1873 ^h
Tomocerus vulgaris Marlier, GJ, 1942:10
Tomocerus vulgaris Berbiers, P et Segers, H, 1991:83
Tomocerus vulgaris De Boey, B, 2002:31

Genus Pogonognathellus Paclt, 1944

Species flavescens (Tullberg, 1871) Yosii, 1967

Tomocerus flavescens Lameere, A, 1895:591
Tomocerus (Pogonognathus) flavescens Leruth, R, 1939:200
Tomocerus flavescens Marlier, GJ, 1942:10
Tomocerus flavescens Delhez, F et Kersmaekers, M, 1973:8
Tomocerus flavescens Berbiers, P et Segers, H, 1991:83
Pogonognathellus flavescens Janssens, F, 1996:53
Tomocerus flavescens Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Pogonognathellus flavescens Janssens, F in Bruers, J et Viskens, G, 2000:15
Pogonognathellus flavescens Dethier, M et Hubart, J-M, 2000:29 ^cf
Pogonognathellus flavescens Dethier, M et Hubart, J-M, 2000:29
Pogonognathellus flavescens Ponge, J-F, 2000:510
Pogonognathellus flavescens De Boey, B, 2002:32

Species longicornis (Müller OF, 1776) Börner, 1908

Tomocerus longicornis Lameere, 1895:591
... longicornis Willem
Tomocerus longicornis Marlier, GJ, 1942:10
Tomocerus longicornis Berbiers, P et Mertens, J, 1989:280
Tomocerus longicornis Berbiers, P et Mertens J, 1989:236
Pogonognathellus longicornis Janssens, F in Bruers, J et Viskens, G, 2000:15

Genus Plutomurus Yosii, 1956

Species unidentatus (Börner, 1901) Stomp, N & Weiner, WM, 1994:360

Tomocerus (Tomocerus) unidentatus Leruth, R, 1939:200
Tomocerus unidentatus Marlier, GJ, 1942:10
Tomocerus unidentatus Dethier, M, 1998:57
Tomocerus unidentatus Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Tomocerus unidentatus Hubart, J-M et Dethier, M, 1999:167

Ordo Symphypleona Börner, 1901, sensu Moen, P et Ellis, WN, 1984:203

Collembola Symphypleona Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20

Familia Sminthurididae Börner, 1906, sensu Betsch J-M, 1974:181

Genus Sminthurides Börner, 1900

Species aquaticus (Bourlet, 1842) Börner, C, 1901:96

Sminthurus aquaticus Lameere, A, 1895:589
Prosminthurus aquaticus Willem, V, 1900:6
Sminthurus aquaticus Willem, V, 1900:55
Prosminthurus aquaticus Lameere, A, 1940:9
Sminthurides aquaticus Marlier, GJ, 1942:10
Sminthurus aquaticus Geyskens, 1945:16
Sminthurides aquaticus Heungens, A et Van Daele, E, 1973:140
Sminthurus aquaticus Verlinden, A, 1977:114
Sminthurides aquaticus Blancquaert, J-P, 1981:2
Sminthurides aquaticus Blancquaert, J-P, Coessens, R et Mertens, J, 1981:116
Sminthurides aquaticus Berbiers, P et Mertens J, 1989:237
Sminthurides aquaticus Viskens, G et Bruers, J, 1994:29
Sminthurides aquaticus Janssens, F, 1995:19

Species malmgreni (Tullberg, 1876) Axelson, 1905

Sminthurides malmgreni Marlier, GJ, 1942:10
Sminthurides malmgreni Heungens, A et Van Daele, E, 1973:140
Sminthurides malmgreni Blancquaert, J-P, 1981:1
Sminthurides malmgreni Berbiers, P et Mertens J, 1989:237

Species parvulus (Krausbauer, 1898) Stach, 1956:19

Sminthurides parvulus Blancquaert, J-P, 1981:1
Sminthurides parvulus Berbiers, P et Mertens J, 1989:237

Species penicillifer (Schäffer, 1896) Börner, C, 1901:92

Sminthurides penicillifer Janssens, F, 1995:18
Sminthurides malmgreni Janssens, 1992 ^e

Species schoetti (Axelson, 1903) Handschin, 1929:110

Smithurides(sic) schoetti Heungens, A et Van Daele, E, 1973:136
Sminthurides schoetti Blancquaert, J-P, 1981:1
Sminthurides schoetti Berbiers, P et Mertens J, 1989:237

Genus Sphaeridia Linnaniemi, WM, 1912:248

Species pumilis (Krausbauer, 1898) Kloet, GS & Hincks, WD, 1945:9

Sminthurides pumilis Heungens, A et Van Daele, E, 1973:138
Sphaeridia pumilis Blancquaert, J-P et Mertens, J, 1977:343
Sphaeridia pumilis Blancquaert, J-P, Coessens, R et Mertens, J, 1981:116
Sphaeridia pumilis Blancquaert, J-P, Mertens, J et Coessens, R, 1982:605
Sphaeridia pumilis Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Sphaeridia pumilis Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Sphaeridia pumilis Pollet, M et Desender, K, 1985:583
Sphaeridia pumilis Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Sminthurides pumilis Berbiers, P et Mertens J, 1989:237
Sphaeridia pumilis Ponge, J-F, 2000:510

Familia Katiannidae Börner, 1913, sensu Betsch J-M, 1974:181

Genus Sminthurinus Börner, C, 1901:99

Species aureus (Lubbock, 1862) Börner, C, 1901:103

Sminthurinus aureus Blancquaert, J-P, Coessens, R et Mertens, J, 1981:116
Sminthurinus aureus Grégoire-Wibo, C, 1981:632
Sminthurinus aureus Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Sminthurinus aureus Berbiers, P et Mertens J, 1989:237
Sminthurinus aureus Berbiers, P et Segers, H, 1991:83
Sminthurinus aureus Jacobs, W, 1997:73
Sminthurinus aureus Ponge, J-F, 2000:510
Sminthurinus aureus f. alba Marlier, GJ, 1942:10
Sminthurinus aureus var. ochropus Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:355

Species elegans (Fitch, 1862) Folsom, 1928

Sminthurinus elegans Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Sminthurinus elegans Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Sminthurinus elegans Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Sminthurinus elegans Berbiers, P et Mertens J, 1989:237
Sminthurinus elegans Janssens, F, 1996:54

Species niger (Lubbock, 1867) Börner, C, 1901:101

Sminthurus niger Lameere, A, 1895:589
Sminthurinus niger Marlier, GJ, 1942:10
Sminthurinus niger Heungens, A et Van Daele, E, 1973:140
Sminthurinus niger Ponge, J-F, 2000:510

Species trinotatus Axelson, 1905

Sminthurinus trinotatus Heungens, A et Van Daele, E, 1973:137

Familia Arrhopalitidae Richards, 1968, sensu Betsch J-M, 1974:181

Genus Arrhopalites Börner, 1906

Arrhopalitidae Arrhopalites Dethier, M et Hubart, J-M, 2000:30 ^sp

Species bifidus Stach, J, 1945:34

Arrhopalites bifidus Dethier, M, 1998:57
Arrhopalites bifidus Delhez, F(), Dethier, M et Hubart, J-M, 1999:35
Arrhopalites bifidus Hubart, J-M et Dethier, M, 1999:168
Arrhopalites bifidus Delhez, Dethier & Hubart, 2000 in prep

Species caecus Tullberg, 1871:146

Arrhopalites caecus Heungens, A et Van Daele, E, 1973:140

Species microphthalmus Cassagnau & Delamare Deboutteville, 1953

Arrhopalites microphtalmus(sic) Jacobs, W, 1997:71 ^iss
Arrhopalites microphthalmus Jacobs, W, De Bruyn, L et Lust, N, 1999:57
Arrhopalites microphtalmus(sic) Jacobs, W, De Bruyn, L et Lust, N, 1999:58 ^iss

Species principalis Stach, J, 1945:37

Arrhopalites principalis Berbiers, P et Mertens J, 1989:237

Species pygmaeus (Wankel, 1860:256) Stach, 1918

Sminthurinus binoculatus Willem, V, 1902:283
Arrhopalites binoculatus Denis, JR, 1937:3
Arrhopalites pygmaeus Bonet, F, 1931:391
Arrhopalites pygmaeus Leruth, R, 1939:201
Arrhopalites pygmaeus Marlier, GJ, 1942:10
Sminthurinus affinis Willem, V, 1902:283
Arrhopalites pygmaeus Godissart, J et Delhez, F, 1969:17
Arrhopalites pygmaeus Godissart, J et Delhez, F, 1969:38
Arrhopalites pygmaeus Delhez, F et Kersmaekers, M, 1973:8
Arrhopalites pygmaeus Delhez, Gilson & Hubart, 1973
Arrhopalites pygmaeus Thibaud, J-M et Massoud, Z, 1977:6
Arrhopalites pygmaeus Hubart, JM, 1982:343
Arrhopalites pygmaeus Dethier, M, 1998:57
Arrhopalites pygmaeus Delhez, F(), Dethier, M et Hubart, J-M, 1999:35
Arrhopalites pygmaeus Hubart, J-M et Dethier, M, 1999:168
Arrhopalites pygmaeus Delhez, Dethier & Hubart, 2000 in prep
Arrhopalites pygmaeus Dethier, M et Hubart, J-M, 2000:19
Arrhopalites pygmaeus Dethier, M et Hubart, J-M, 2000:30 ^cf
Arrhopalites pygmaeus Dethier, M et Hubart, J-M, 2000:30

Species sericus Gisin, 1947

Arrhopalites sericus Blancquaert, J-P, Coessens, R et Mertens, J, 1981:116
Arrhopalites sericus Berbiers, P et Mertens J, 1989:237

Familia Dicyrtomidae Börner, 1906, sensu Betsch J-M, 1974:181

Genus Ptenothrix Börner, 1906

Species atra (Linnæus, C, 1758:608) Börner, 1906

Ptenothrix atra Ball, A, 1926:261

Species setosa (Krausbauer, 1898:24) Börner, 1909

Ptenothrix setosa Leruth, R, 1939:201
Ptenothrix setosa Marlier, GJ, 1942:11

Genus Dicyrtoma Bourlet, 1841

Species fusca (Lubbock, 1873:120) Axelson, 1903

Papirius fuscus Lameere, A, 1895:590
Papirius fuscus Willem, V, 1900:6
Papirius fuscus Marlier, 1942:11
Dicyrtoma fusca Berbiers, P et Mertens J, 1989:237

Genus Dicyrtomina Börner, 1903

Species minuta (Fabricius O, 1783) Börner, 1906

Papirius minutus Lameere, A, 1895:590
Papirius minutus Willem, V, 1900:63
Dicyrtoma minuta Marlier, GJ, 1942:11
Dicyrtoma minuta Heungens, A et Van Daele, E, 1973:140
Dicyrtomina minuta Thibaud, J-M et Massoud, Z, 1977:6
Dicyrtoma minuta Berbiers, P et Mertens J, 1989:237
Dicyrtomina minuta Janssens, F, 1996:54
Dicyrtomina minuta Ponge, J-F, 2000:510
Papirius flavo-signatus(sic) Willem, V, 1900:63 ^iss
Dicyrtomina ornata Thibaud, J-M et Massoud, Z, 1977:6
Dicyrtoma ornata Berbiers, P et Mertens J, 1989:237
Dicyrtoma ornata Berbiers, P et Segers, H, 1991:83

Species saundersi (Lubbock, 1862) Kloet, GS & Hincks, WD, 1945:9

Dicyrtoma saundersi Berbiers, P et Mertens J, 1989:237

Familia Bourletiellidae Börner, 1912, sensu Betsch J-M, 1974:181

Genus Deuterosminthurus Börner, C, 1901:104 nc , 1976

Species bicinctus (Koch in Herrich-Schäffer, 1840) Linnaniemi, 1912

Deuterosminthurus bicinctus Marlier, GJ, 1942:10
Bourletiella bicincta Heungens, A et Van Daele, E, 1973:138
Deuterosminthurus bicinctus Verlinden, 1977:114
Deuterosminthurus bicinctus Berbiers, P et Mertens J, 1989:237

Species pallipes (Bourlet, 1843) Handschin, 1924

Sminthurus pallipes Lameere, 1895:589
Bourletiella pallipes Heungens, A et Van Daele, E, 1973:138
Deuterosminthurus pallipes Berbiers, P et Mertens J, 1989:237

Species repandus (Ågren, 1903) Linnaniemi, 1912

Deuterosminthurus repandus Berbiers, P et Mertens J, 1989:237

Species sulphureus (Koch, 1840) Bretfeld, G, 1971:147

Bourletiella sulphurea Berbiers, P et Mertens J, 1989:237

Genus Bourletiella Banks, 1899

Sminthuridae Bourletiella Marlier, GJ, 1942:5 ^sp

Species arvalis (Fitch, 1862) Folsom, 1928

Sminthurus luteus Lameere, A, 1895:589
Bourletiella lutea Marlier, GJ, 1942:10
Bourletiella luteus(sic) Janssens, F, 1995:19 ^iss

Species hortensis (Fitch, 1862) Brown, 1918

Bourletiella hortensis Grégoire-Wibo, C, 1983:41
Bourletiella hortensis Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Bourletiella hortensis Berbiers, P et Mertens J, 1989:237
Bourletiella hortensis Viskens, G et Bruers, J, 1994:29

Genus Heterosminthurus Stach, 1955

Species bilineatus (Bourlet, 1842:146) Stach, 1956:181

Sminthurus bilineatus Lameere, 1895:589
Heterosminthurus bilineatus Berbiers, P et Mertens J, 1989:237

Species claviger Gisin, 1958

Heterosminthurus claviger Berbiers, P et Mertens J, 1989:237

Species insignis (Reuter, 1876) Stach, 1956:181

Deuterosminthurus insignis Marlier, GJ, 1942:10

Genus Fasciosminthurus Gisin, 1960

Species quinquefasciatus (Krausbauer, 1898) Salmon, 1964

Bourtellia(sic) quinquefasciata Jacquemart, S, 1967:5
Bourletellia(sic) quinquefasciata Jacquemart, S, 1967:7

Familia Sminthuridae Lubbock, 1862, sensu Betsch J-M, 1974:181

Genus Sphyrotheca Börner, 1906

Sminthuridae Sphyroteca(sic) Jacobs, W, 1997:73 ^sp

Genus Lipothrix Börner, 1906

Species lubbocki (Tullberg, 1872) Ellis & Bellinger, 1973:29

Sminthurus lubbocki Jacquemart, S, 1967:4
Sphyrotheca lubbocki Berbiers, P et Mertens J, 1989:237
Lipothrix lubbocki Ponge, J-F, 2000:510

Genus Allacma Börner, 1906

Species fusca (Linnæus, C, 1758:608:608) Börner, 1906

Sminthurus fuscus Lameere, A, 1895:589
Sminthurus fuscus Willem, V, 1900:56
... fuscus Willem, 1901
Allacma fusca Marlier, GJ, 1942:11
Sminthurus fuscus Geyskens, 1945:16
Allacma fusca Berbiers, P et Mertens, J, 1989:280
Allacma fusca Berbiers, P et Mertens J, 1989:237
Allacma fusca Dall'Asta, 1990:135

Genus Sminthurus Latreille in Sonnini, [1802], n.c.

Species multipunctatus Schäffer, 1896

Sminthurus multipunctatus Marlier, GJ, 1942:11
Sminthurus multipunctatus Dethier, M et Hubart, J-M, 2000:30 ^cf

Species viridis (Linnæus, C, 1758:608) Tullberg, 1871

Sminthurus viridis Lameere, A, 1895:589
... viridis Willem
Sminthurus viridis Lameere, A, 1940:9
Sminthurus viridis Marlier, GJ, 1942:11
Sminthurus viridis Heungens, A et Van Daele, E, 1973:136
Sminthurus viridis Verlinden, A, 1977:114
Sminthurus viridis Grégoire-Wibo, C, 1981:632
Sminthurus viridis Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Sminthurus viridis Grégoire-Wibo, C, 1983:41
Sminthurus viridis Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Sminthurus viridis Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Sminthurus viridis Berbiers, P et Mertens J, 1989:237

Genus Caprainea Dallai, 1970

Species marginata (Schött, H, 1893:25) Dallai, 1972

Caprainea marginata Jacobs, W, 1997:71
Caprainea marginata Jacobs, W, De Bruyn, L et Lust, N, 1999:59

Discussion

Chen & Christiansen (1997) consider European records of Sinella caeca as records of the widespread Sinella tenebricosa.

Acknowledgements

We thank Wim Jacobs for his contribution based on his unpublished thesis. We thank Peter Bellinger for reviewing the species list. Especially for pointing out the problematic issue of the European records of Sinella caeca versus Sinella tenebricosa. We thank Jean-François Ponge for providing a species list from beech forests of the Belgian Ardennes. We thank Michel Dethier for the provisional catalogue (including addenda & corrigenda) of the Delhez Collection, for providing additional locality data and for providing additional papers on Belgian cave Collembola. We thank Koen Lock and Jos Bruers for their contribution in the form of an exhaustive bibliographic research.

Appendix

Loci Collembola Belgica.