http://www.collembola.org/taxa/bouridae.htm - Last updated on 2024.04.27 by Frans Janssens
Checklist of the Collembola: Bourletiellidae

Familia Bourletiellidae Börner, 1913:322, sensu Bretfeld, 1994

Genus Number
of species
A1
Acutoempodialis1
Adisianus3
Anarmatus5
Aneuempodialis2
Anjavidiella5
Arlesminthurus5
B1
Bourletides1
Bourletiella27
Bourletiellitas1
Bovicornia7
Cassagnaudiella8
Corynephoria7
Cyprania2
Deuterosminthurus24
Diksamella1
Ellisiella1
Fasciosminthurus30
Heterosminthurus25
Karakumiella1
Kaszabellina3
Koupelia1
Madecassiella1
Massoudia1
N.11
Nasosminthurus4
Navarrella1
Orenius2
Parabourletiella1
Paulianitas1
Prorastriopes41
Pseudobourletiella1
Rastriopes16
Sanaaiella5
Stenognathriopes (Stenognathriopes)6
Stenognathriopes (Tenentiella)2
Stenognathriopes9
Tritosminthurus1
Vatomadiella3
Bourletiellidae250
 Genera with number of species
Deuterosminthurus pallipes ♂ & var. bellingeri ♀ from France
Courtship ritual
2010.06.04 © Elpachato.
Bourletiella sp. from Australia
2012.07.03 © Hort, J.

  Syn.: ... Smynthuridae(sic) Lubbock, 1862, partim Lubbock, 1870
  Syn.: Sminthuridæ(sic) Smynthurus(sic) Packard, 1873 partim
  Syn.: Tridentata Bourletiellida(sic) Bretfeld, 1986
  Syn.: Tridentata Bourletiellida(sic) Bretfeld, G, 1999:213



  Syn.: Collembola Boerletiellidae(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:111 ISS
Bourletiella hortensis ♀ from Belgium
5 bothriotricha ABC+DE
2020.07.30 © Huskens, M.-L.





Genus A Bretfeld, G, 2005:50

Species spec Bretfeld, G, 2005:50

Gen. spec. A from Yemen
After Bretfeld G, 2005 Fig.101


Genus B Bretfeld, G, 2005:53

Species spec Bretfeld, G, 2005:53


Genus N.1 Ernesto, MV, Liberal, CN, Ferreira, AS, Alves, ACF, Zeppelini, D, Martins, CF, Pereira-Colavite, A, Creao-Duarte, AJ, Vasconcellos, A, 2018:5

Species sp. Ernesto, MV, Liberal, CN, Ferreira, AS, Alves, ACF, Zeppelini, D, Martins, CF, Pereira-Colavite, A, Creao-Duarte, AJ, Vasconcellos, A, 2018:5


Subfamilia Parabourletiellinae Betsch J-M, 1974:181

  Syn.: Bourletiellidae Parabourletiellidae(sic) Betsch, J-M, 1997:16 ISS


Genus Bourletiellitas Betsch, J-M, 1974:554

Species imerinensis Betsch, J-M, 1974:547, t.t.


Genus Anjavidiella Betsch, J-M, 1974:560

Species andoharianensis Betsch, J-M, 1980:182

Anjavidiella andoharianensis from Madagascar
After Cipola NG in Cipola NG & al, 2022 Fig.9.17.D

Species andringitrensis Betsch, J-M, 1980:182

Species anjavidilavensis Betsch, J-M, 1980:182

Species ankaratrensis Betsch, J-M, 1974:554, t.t.

Species vohidrayensis Betsch, J-M, 1980:184


Genus Vatomadiella Betsch, J-M, 1974:567

Species descarpentriesi Betsch, J-M, 1980:185

Species pauliani Betsch, J-M, 1974:560, t.t.

Species peyrierasi Betsch, J-M, 1980:184


Genus Parabourletiella Betsch, J-M, 1975:477

Species mahafalensis Betsch, J-M, 1975:478, t.t.


Subfamilia Bourletiellinae Börner, 1912, sensu Betsch J-M, 1974:181


Genus Acutoempodialis Bretfeld, 2000

Species lineatus Bretfeld, 2000, t.t.


Genus Adisianus Bretfeld, G, 2003:551

Species fuscus (Bretfeld, 2002) Bretfeld, G, 2003:551

Species maassius (Palacios-Vargas, JG & González, V, 1995:290) Bretfeld, G, 2003:551

Deuterosminthurus maassius from Cuba
After Palacios-Vargas JG & González V, 1995 Fig.17
Deuterosminthurus maassius from Cuba
After Palacios-Vargas JG & González V, 1995 Fig.18

Species maculatus (Bretfeld, 2002) Bretfeld, G, 2003:551, t.t.


Genus Anarmatus Bretfeld, 2000 key

Species diksamanus Bretfeld, G, 2005:2

Anarmatus diksamanus from Yemen
After Bretfeld G, 2005 Fig.1

Species goeehensis Bretfeld, G, 2005:4

Anarmatus goeehensis from Yemen
After Bretfeld G, 2005 Fig.5

Species irregularius Bretfeld, G, 2005:6

Anarmatus irregularius from Yemen
After Bretfeld G, 2005 Fig.9

Species longisetus Bretfeld, G, 2005:8

Species taizzus Bretfeld, 2000, t.t.


Genus Sanaaiella Bretfeld, 2000

Species circumscripta Bretfeld, 2000

Species dorsopallida Bretfeld, 2000

Species multisensillata Bretfeld, 2000

Species quadripunctata Bretfeld, 2000

Species septemlineata Bretfeld, G, 2005:19

Sanaaiella septemlineata from Yemen
After Bretfeld G, 2005 Fig.34


Genus Rastriopes Börner, C, 1906:182

Rastriopes sp. from Tasmania
2014.08.11 © Zosterops, K.
Rastriopes sp. from Tasmania
2014.08.11 © Zosterops, K.
Rastriopes sp. from Tasmania
2014.08.11 © Zosterops, K.
Rastriopes sp. from Tasmania
2016.04.14 © Zosterops, K.
Rastriopes sp. from Chile
2017.08.10 © Segura, Be.
Rastriopes sp. from Australia
2019.06.30 © Tony.
Rastriopes sp. from Australia
2019.06.30 © Tony.
Rastriopes sp. from South Africa
2020.01.12 © Manson, A.D.

Species sp.1 Zeppelini, D, Bellini, BC, Creao-Duarte, AJ et Hernández, MIM, 2009:1166

Species sp.2 Janssens F & Segura Be, 2017

Rastriopes sp. from Chile
2017.06.10 © Segura, Be.
Rastriopes sp. from Chile
2017.06.10 © Segura, Be.
Rastriopes sp. from Chile
2017.06.10 © Segura, Be.

Species aculeatus (Börner, C, 1906:182) Bonet, F, 1933:124, t.t.

             Syn.: Rastriopes aculeata(sic) Bonet, F, 1933:124 ISS

Species amphigyus Börner, 1906

Species deformis (Nicolet, H, 1847:359) Stach, J, 1956:238, i.s.

Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Species dromedarius (Womersley, H, 1932:28) Greenslade, P, 1974:259

Rastriopes nr dromedarius from Australia
2015.11.09 © Murray, A.

Species echidnus Yoshii, R et Suhardjono, YR, 1992:92

Species fulvipes (Nicolet, H, 1847:359) Stach, J, 1956:238, i.s.

Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Species fuscus Yosii, 1960

Species gisinianus Murphy, 1960

Species handschini (Womersley, H, 1932:28) Greenslade, P in Houston, WWK, 1994:129

            Syn.: Deuterosminthurus (Rastriopes) handschini var. atratus Womersley, H, 1932:28


Rastriopes handschini from Australia
2012.07.15 © Hort, J.
Rastriopes handschini from Australia
2016.11.20 © MacAulay, I.
Rastriopes handschini from Australia
2016.11.20 © MacAulay, I.

Species lineatus Womersley, 1931

            Syn.: Rastriopes lineata(sic) Womersley, 1931 IOS
            Syn.: Rastriopes schultzei Paclt, 1959


             Syn.: Rastriopes lineata(sic) Salmon, JT, 1964:610 ISS
             Syn.: Rastriopes lneata(sic) Thibaud, J-M, 2013:240 ISS
             Syn.: Rastriopes lineata(sic) Janion-Scheepers, C, Deharveng, L, Bedos, A et Chown, L, 2015:73 ISS



Species obscurus (Womersley, H, 1931:7) Greenslade, P, 1977:247

Deuterosminthurus obscurus from Tasmania
After Womersley, 1931 Pl.I Fig.4


Species schultzei Börner, C in Schultze, 1908:61

Species tricuspidatus (Börner, 1907) Mari Mutt, JA & Bellinger, PF, 1990:151

            Syn.: Sminthurus pallipes Parona, 1895
            Syn.: Sminthurus pallipes Börner, 1907


Species vexillarius (Schäffer, C, 1898:422) Stach, 1956


Genus Deuterosminthurus Börner, 1901 nc ICZN Opinion 1042, 1976 key

  Syn.: Sminthurides(sic) Sminthurus Bourlet, C, 1845:147
  Syn.: Bourletiellidae Adisianus Magaña-Martínez, CS, Palacios-Vargas, JG et Navarrete-Heredia, JL, 2019:62 lapsus


Remark by ICZN. 2007:241: Placed on the Official List of Names in Zoology. Opinion 1042.

Deuterosminthurus sp. from France
2010.05.24 © Elpachato.
Deuterosminthurus sp. from France
2010.05.24 © Elpachato.
Deuterosminthurus sp. juv. from Ibiza
2014.01.09 © Beer, R.
Deuterosminthurus sp. from Ibiza
2014.02.03 © Beer, R.
?Deuterosminthurus? sp. from Taiwan
2022.01.25 © Cheng, H.-J.

Species bicinctus (Koch in Herrich-Schäffer, 1840) Linnaniemi, 1912, t.t., B

            Syn.: ... pallipes Bourlet
            Syn.: Sminthurus pallipes Tullberg, 1871
            Syn.: Deuterosminthurus bicinctus var. pallipes Linnaniemi, 1912
            Syn.: Smynthurus(sic) luteus Lubbock, J, 1870:281 partim
            Syn.: Sminthurus luteus Tullberg, 1872
            Syn.: Sminthurus cinctus Tullberg, 1871
            Syn.: Sminthurus lupulinae Lubbock, J, 1873:114
            Syn.: Sminthurus speciosus Krausbauer, 1898
            Syn.: Sminthurus novemlineatus Börner, 1900, partim
            Syn.: Sminthurus (Deuterosminthurus) oblongus ssp. minor Börner, C, 1901:110
            Syn.: Sminthurus (Deuterosminthurus) oblongus ssp. minor var. puncteola(sic) Börner, C, 1901:112 IOS
Sminthurus oblongus var. puncteola(sic)
from Germany
After Börner C, 1901 Tafel I Fig.5

            Syn.: Bourletiella sulphurea ssp. bicincta var. puncteola Börner, C in Schultze, 1908:55
             Syn.: Deuterosminthurus bicinctus var. puncteola(sic) Stach, J, 1956:173 ISS
Deuterosminthurus bicinctus var. puncteola(sic)
from Poland
After Stach J, 1956 Plate XX Fig.3

             Syn.: Deuterosminthurus bicinctus var. puncteola(sic) Bretfeld, G, 1999:236 ISS
Deuterosminthurus bicinctus from Poland
Courtship with female var. puncteolus
2002 © Kozlowski, M.W. & Aoxiang, S.
Deuterosminthurus bicinctus var. puncteolus juv. from the UK
2012.05.14 © Nurcombe, E.
Deuterosminthurus bicinctus var. puncteolus ♀ from the UK
2012.06.20 © Nurcombe, E.
Deuterosminthurus bicinctus from the UK
Courtship with female var. puncteolus
2012.06.20 © Nurcombe, E.
Deuterosminthurus bicinctus var. puncteolus from France
2018.05.04 © Tiky, S.
Deuterosminthurus bicinctus var. puncteolus from France
2018.05.04 © Tiky, S.
Deuterosminthurus bicinctus var. puncteolus from France
2018.05.04 © Tiky, S.
Deuterosminthurus bicinctus var. puncteolus from Belgium
2018.05.05 © Huskens, M.-L.
Deuterosminthurus bicinctus var. puncteolus juv. from Belgium
2020.04.19 © Huskens, M.-L.
Deuterosminthurus bicinctus var. puncteolus ♀ & f. flavus ♂ from France
Courtship
2020.05.03 © Tiky, S.


            Syn.: Burletiella(sic) sulphurea Börner, 1908
            Syn.: ... sulphureus Collinge & Shoebotham, 1910
            Syn.: Deuterosminthurus bicinctus f. repandus Womersley, H, 1931:6
Deuterosminthurus bicinctus f. repandus from Tasmania
After Womersley, 1931 Pl.I Fig.2

             Syn.: Deuterosminthurus bicinctus f. repanda(sic) Womersley, H, 1931:6 ISS
Deuterosminthurus bicinctus
subsp. repandus from New Zealand
After Salmon JT, 1941 Pl.72 Fig.520

            Syn.: Bourletiella (Deuterosminthurus) flava Gisin, 1946
             Syn.: Deuterosminthurus bicinctus f. flava(sic) Bretfeld, 1970 ISS
             Syn.: Deuterosminthurus bicinctus f. flava(sic) Bretfeld, 1996 ISS
             Syn.: Deuterosminthurus bicinctus f. flava(sic) Bretfeld, G, 1999:236 ISS
Deuterosminthurus bicinctus from the USA
Deuterosminthurus bicinctus f. flavus ♀ & ♂
Deuterosminthurus bicinctus ♂
2007.06.04 © Reed, D.
Deuterosminthurus bicinctus ♂ & f. flavus ♀ from the UK
2010.05.22 © Michaelson, J.

             Syn.: Deuterosminthurus bicinctus var. flava(sic) Berg, M et den Ouden, A, 2013:5 ISS
Deuterosminthurus bicinctus f. flavus ♀ from France
2017.08.26 © Picard, J.
Deuterosminthurus bicinctus f. flavus ♀ from France
2017.08.26 © Picard, J.
Deuterosminthurus bicinctus f. flavus ♀ from France
2017.08.26 © Picard, J.
Deuterosminthurus bicinctus f. flavus ♀ from France
2017.08.26 © Picard, J.
Deuterosminthurus bicinctus f. flavus ♀ & ♂ from France
Courtship
2018.05.04 © Tiky, S.
Deuterosminthurus bicinctus f. flavus ♀ from Holland
Dark orange form
2018.06.14 © Kamsteeg, G.
Deuterosminthurus bicinctus ♂ & f. flavus ♀ from Holland
Courtship
2018.06.14 © Kamsteeg, G.
Deuterosminthurus bicinctus f. flavus ♂ & ♀ from Denmark
Spermatophore deposition in front of ♀
2018.07.12 © Schlütter, P.
Deuterosminthurus bicinctus f. flavus ♂ & ♀ from Denmark
♀ eats spermdrop; ♂ pulls ♀ over spermdrop
2018.07.12 © Schlütter, P.
Deuterosminthurus bicinctus f. flavus ♂♂ & ♀ from Denmark
Courtship
2018.08.26 © Schlütter, P.
Deuterosminthurus bicinctus f. flavus ♀ from Hungary
2018.09.28 © Ujvári, Z.
Deuterosminthurus bicinctus ♂ & f. flavus ♀ from Belgium
Courtship
2020.04.29 © Huskens, M.-L.
Deuterosminthurus bicinctus f. flavus from Belgium
2021.06.01 © Huskens, M.-L.




Deuterosminthurus bicinctus from Russia
After Potapov M. in Babenko A.B., 1988
Deuterosminthurus bicinctus from Poland
2002 © Kozlowski, M.W. & Aoxiang, S.
Deuterosminthurus bicinctus
2005 © Kozlowski, M.W. & Aoxiang, S.
Deuterosminthurus bicinctus
'Premating-dance'
2005 © Kozlowski MW & Aoxiang S.
Deuterosminthurus bicinctus
Sperm deposition and take-up
2005 © Kozlowski MW & Aoxiang S.
Deuterosminthurus bicinctus ♂ from the USA
2007.06.02 © Reed, D.
 
 
Deuterosminthurus bicinctus ♂ from the USA
2007.06.04 © Reed, D.
 
Deuterosminthurus bicinctus juv. from the UK
2012.04.15 © Nurcombe, E.
Deuterosminthurus bicinctus juv. from the UK
2012.04.15 © Nurcombe, E.
Deuterosminthurus bicinctus ♂ from the UK
2012.06.20 © Nurcombe, E.
 
Deuterosminthurus bicinctus juv. from the UK
2012.06.26 © Murray, A.
 
Deuterosminthurus bicinctus ♀ from the UK
2012.06.27 © Murray, A.
Deuterosminthurus bicinctus from the USA
2013.07.04 © Eiseman, C.
Deuterosminthurus bicinctus from the USA
Ant.4 with 7 subsegments
2013.07.04 © Eiseman, C.
Deuterosminthurus bicinctus ♂ & ♀ from Germany
Courtship
2014.06.20 © Beck, K.
Deuterosminthurus bicinctus from Holland
2017.07.15 © Kamsteeg, G.
Deuterosminthurus bicinctus from Holland
2017.07.15 © Kamsteeg, G.
Deuterosminthurus bicinctus from Holland
Dark form
2018.06.09 © Kamsteeg, G.
Deuterosminthurus bicinctus from Holland
Darker form
2018.06.09 © Kamsteeg, G.
Deuterosminthurus bicinctus from Belgium
2018.06.12 © Huskens, M.-L.
Deuterosminthurus bicinctus 2 ♂ & ♀ from Denmark
Courtship
2018.08.26 © Schlütter, P.
Sminthurinus bicinctus from Holland
Dark colour form
2019.06.20 © Kamsteeg, G.
Sminthurinus bicinctus from Holland
Dark colour form
2019.06.20 © Kamsteeg, G.
Deuterosminthurus bicinctus ♂ & ♀ from Belgium
Male deposited spermatophore in front of female
2020.04.19 © Huskens, M.-L.
Deuterosminthurus bicinctus ♂ & ♀ from Belgium
Male prevents female from eating the spermatophore
2020.04.19 © Huskens, M.-L.
Deuterosminthurus bicinctus ♀ from Belgium
Female connects gonopore to spermdrop of spermatophore
2020.04.19 © Huskens, M.-L.
Deuterosminthurus bicinctus ♀ from Belgium
Female takes up sperm from spermdrop
2020.04.19 © Huskens, M.-L.
Deuterosminthurus bicinctus ♀ from Belgium
Spermdrop partly emptied
2020.04.19 © Huskens, M.-L.
Deuterosminthurus bicinctus ♀ from Belgium
Spermdrop empty
2020.04.19 © Huskens, M.-L.
Deuterosminthurus bicinctus ♀ from Belgium
Female eats remainder of spermatophore
2020.04.19 © Huskens, M.-L.
Deuterosminthurus bicinctus ♂ & ♀ from Belgium
?Male offers drop of saliva to female?
2020.04.29 © Huskens, M.-L.
Deuterosminthurus bicinctus juv. from France
Feeding instar
2021.05.03 © Vol, F.
Deuterosminthurus bicinctus from the USA
2021.05.20 © BugIdentifier.
Sminthurinus bicinctus from Holland
Dark colour form
2021.05.20 © Kamsteeg, G.
Deuterosminthurus bicinctus ♂ & ♀ from Belgium
Courtship
2023.05.18 © Huskens, M.-L.



Species bisetosus Baquero, E, Moraza, ML et Jordana, R, 2003:9

Deuterosminthurus bisetosus from Spain
After Baquero E, Moraza ML & Jordana R, 2003 Fig.29
Deuterosminthurus bisetosus from Spain
Generalised colouration
After Baquero E, Moraza ML & Jordana R, 2003 Fig.37
Deuterosminthurus bisetosus from Spain
Pale specimen
After Baquero E, Moraza ML & Jordana R, 2003 Fig.36
Deuterosminthurus bisetosus from Spain
Pale specimen
After Baquero E, Moraza ML & Jordana R, 2003 Fig.39
Deuterosminthurus bisetosus from Spain
Pale specimen
After Baquero E, Moraza ML & Jordana R, 2003 Fig.42
Deuterosminthurus bisetosus from Spain
Orange-red specimen
After Baquero E, Moraza ML & Jordana R, 2003 Fig.38
Deuterosminthurus bisetosus from Spain
Red coloured specimen
After Baquero E, Moraza ML & Jordana R, 2003 Fig.40
Deuterosminthurus bisetosus from Spain
Dark pattern
After Baquero E, Moraza ML & Jordana R, 2003 Fig.41

Species caeruleacaudus (Scott, 1965)

            Syn.: Bourletiella (Deuterosminthurus) sp. Bellinger, PF, 1985:78

Species delatorrei Palacios-Vargas, JG et González, V, 1995:287

Deuterosminthurus delatorrei from Mexico
After Palacios-Vargas JG & González V, 1995 Fig.1
Deuterosminthurus delatorrei from Mexico
After Palacios-Vargas JG & González V, 1995 Fig.2
Deuterosminthurus delatorrei from Peru
Cephalic branching trachea
2016.06.09 © Palacios-Vargas, J.G.

Species doriae (Parona, 1884) Stach, J, 1956:238, i.s.

            Syn.: Smynthurus(sic) Doriæ(sic) Parona, 1884 IOS


             Syn.: Sminthurus Doriæ(sic) Schött, H, 1893:4 ISS



Species ezoensis Yosii, R, 1972:95

Species kugleri Bretfeld in Bretfeld, Poliakov & Broza, 2000

Species luridus (Snider, 1978) Palacios-Vargas, JG & González, V, 1995:293

             Syn.: Deuterosminthurus lurida(sic) Palacios-Vargas, JG et González, V, 1995:293 ISS
             Syn.: Bourletiella (Deuterosminthurus) luridus Christiansen, K et Bellinger, P, 1998:1302 ISS
             Syn.: Deuterosminthurus lurida(sic) Vázquez, M et Palacios-Vargas, JG, 2004:105 ISS
             Syn.: Deuterosminthurus lurida(sic) Kang, J-S, Lee, B-H et Park, K-H, 2005:238 ISS
             Syn.: Deuterosminthurus lurida(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:111 ISS
Deuterosminthurus luridus ♂ from the USA
2017.04.05 © Fritzl.
Deuterosminthurus luridus ♂ from the USA
Ant.4 with 10 subdivisions
2017.04.05 © Fritzl.
Deuterosminthurus luridus ♀ from the USA
2017.04.05 © Fritzl.
Deuterosminthurus luridus ♂ from the USA
2018.06.01 © Dobbs, Br.
Deuterosminthurus luridus from the USA
2018.06.01 © Dobbs, Br.



Species mediterraneus Ellis, 1974

Species nonfasciatus (Snider, 1978) Palacios-Vargas, JG & González, V, 1995:293

             Syn.: Deuterosminthurus nonfasciata(sic) Palacios-Vargas, JG, 2000:33,ca ISS
Deuterosminthurus nonfasciatus from Canada
2007.05.14 © Pilon, C.
Deuterosminthurus nonfasciatus from Canada
2007.05.14 © Pilon, C.
Deuterosminthurus nonfasciatus ♂ & ♀ from the USA
Courtship ritual
2007.05.25 © Reed, D.
Deuterosminthurus nonfasciatus ♂ & ♀ from the USA
2007.05.25 © Reed, D.
Deuterosminthurus nonfasciatus ♀ & ♂ from Canada
2007.05.27 © Marsman, I.
Deuterosminthurus nonfasciatus ♀ & ♂ from the USA
2007.06.02 © Reed, D.
Deuterosminthurus nonfasciatus from the USA
2012.04.07 © Kouri, J.
Deuterosminthurus nonfasciatus from the USA
2023.04.27 © Gruver, Be.
Deuterosminthurus nonfasciatus from the USA
2023.04.27 © Gruver, Be.
Deuterosminthurus nonfasciatus from the USA
Ant.4 with 15 subdivisions
2023.04.27 © Gruver, Be.
Deuterosminthurus nonfasciatus from the USA
Eye with 8 ocelli (A-H)
Ocelli BCH distinctly larger
2023.04.27 © Gruver, Be.



Species okinawanus Yosii, R, 1965:47

Deuterosminthurus okinawanus ♀ from Japan
After Yosii R, 1965 Fig.30A

Species pallipes (Bourlet, 1843) Handschin, 1924, B

            Syn.: Smynthurus(sic) echii Koch, 1840
            Syn.: Sminthurus ochraceus Koch, 1840
            Syn.: Smynthurus(sic) sulphureus Koch, 1840
            Syn.: Smynthurus(sic) lupulinae Bourlet, 1842
             Syn.: Smynthurus(sic) lupulinæ(sic) Lubbock, J, 1870:296 ISS
            Syn.: Sminthurus pallipes var. verticis maculis Bourlet, C, 1845:148
            Syn.: Sminthurus luteus Lubbock, 1867
            Syn.: Sminthurus atratus Schäffer, 1896
            Syn.: Sminthurus speciosus Krausbauer, 1898
            Syn.: Sminthurus repandus Ågren, 1903
             Syn.: Deuterosminthurus bicinctus f. repanda(sic) Handschin, E in Dahl, F, 1929:117 ISS
             Syn.: Deuterosminthurus repanda(sic) Nayrolles, P, 1996:549 ISS
             Syn.: Deuterosminthurus pallipes f. repanda(sic) Bretfeld, G, 1999:239 ISS
             Syn.: Deuterosminthurus pallipes f. repanda(sic) Bretfeld, G, 2001:89 ISS
             Syn.: Deuterosminthurus pallipes f. repanda(sic) Bretfeld, G, 2001:90 ISS
             Syn.: Deuterosminthurus pallipes f. repanda(sic) Bretfeld, G, 2001:99 ISS
             Syn.: Deuterosminthurus pallipes f. repanda(sic) Bretfeld, G, 2001:129 ISS
             Syn.: Deuterosminthurus repanda(sic) Baquero, E, Moraza, ML et Jordana, R, 2003:15 ISS
             Syn.: Deuterosminthurus pallipes f. repanda(sic) Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:895 ISS
Deuterosminthurus pallipes f. repandus from the UK
2005 © Hopkin, S.P.
Deuterosminthurus pallipes f. repandus from Belgium
2007.05.03 © Gielen, K.
Deuterosminthurus pallipes f. repandus from the UK
2008.07.13 © Michaelson, J.

             Syn.: Deuterosminthurus pallipes f. repanda(sic) Bretfeld, G, 2010:303 ISS
             Syn.: Deuterosminthurus pallipes f. repanda(sic) Petersen, H, 2011:472 ISS
             Syn.: Deuterosminthurus pallipes f. repanda(sic) Petersen, H, 2011:476,cf ISS
Deuterosminthurus pallipes f. repandus from Ibiza
Grooming with everted vesicle of collophore
2011.02.11 © Beer, R.
Deuterosminthurus pallipes f. repandus from Ibiza
Courtship ritual
2011.12.14 © Beer, R.
Deuterosminthurus pallipes f. repandus from the UK
2012.07.01 © Nurcombe, E.
Deuterosminthurus pallipes f. repandus from the UK
2012.07.01 © Nurcombe, E.
Deuterosminthurus pallipes f. repandus juv. from the UK
2012.07.09 © Nurcombe, E.
Deuterosminthurus pallipes f. repandus juv. from the UK
Subdivided 4th antennal segment
2013.02.21 © Phillips, E.
Deuterosminthurus pallipes f. repandus juv. from the UK
'Pearly' composed eye
2013.02.21 © Phillips, E.
Deuterosminthurus pallipes f. repandus from Russia
Predated by pseudoscorpion
2013.10.05 © Haritonoff.
Deuterosminthurus pallipes f. repandus ♀ & ♂ from Holland
2014.05.01 © van Duinen, J.
Deuterosminthurus pallipes f. repandus from Belgium
Ant.4 subdivided
2016.07.09 © Huskens, M.-L.
Deuterosminthurus pallipes f. repandus from Belgium
Dorsal anal valve (dav) longer than ventral valves (vav); * = anus
2017.05.07 © Huskens, M.-L.
Deuterosminthurus pallipes f. repandus from Belgium
Ant.4 with 7 subdivisions
2017.05.18 © Huskens, M.-L.
Deuterosminthurus pallipes f. repandus from Belgium
♂ guides ♀ to spermatophore
2017.06.20 © Huskens, M.-L.
Deuterosminthurus pallipes f. repandus from France
With "S" pattern of spots
2019.05.29 © Picard, J.
Deuterosminthurus pallipes f. repandus from France
Abd.6 with 2 spots
2019.05.29 © Picard, J.
Deuterosminthurus pallipes f. repandus from France
Ocellus H distincly largest
Central ocellus D distinctly smallest
2019.05.29 © Picard, J.
Deuterosminthurus pallipes f. repandus from France
Bicoloured form
2019.05.31 © Picard, J.
Deuterosminthurus pallipes f. repandus from France
Bicoloured form
2019.05.31 © Picard, J.
Deuterosminthurus pallipes f. repandus from France
Bicoloured form
2019.05.31 © Picard, J.
Deuterosminthurus pallipes f. repandus ♀ from Belgium
Ocelli BCH distinctly larger than ocellus D
2020.04.23 © Huskens, M.-L.
Deuterosminthurus pallipes f. repandus ♀ from Belgium
Distance ocelli GH larger then FG
2020.04.26 © Huskens, M.-L.
Deuterosminthurus pallipes f. repandus ♀ from Belgium
Bothriotricha A-D
2020.04.26 © Huskens, M.-L.
Deuterosminthurus pallipes f. repandus ♀ from the UK
With lateral "S" pattern
2020.04.30 © Barton, T.
Deuterosminthurus pallipes f. repandus ♂ & ♀ from the UK
Male deposited spermatophore in front of female
2020.05.02 © Barton, T.
Deuterosminthurus pallipes f. repandus ♂ & ♀ from Belgium
Courtship
2021.05.10 © Huskens, M.-L.
Deuterosminthurus pallipes f. repandus ♂ & ♀ from Belgium
♂ dips antenna in spermdrop
and pulls thread that ♀ should follow
2021.05.10 © Huskens, M.-L.
Deuterosminthurus pallipes f. repandus ♀ from Norway
2021.08.06 © Joachim, J.
Deuterosminthurus pallipes f. repandus ♀ from the Canary Islands
2023.11.23 © Huskens, M.-L.
Deuterosminthurus pallipes f. repandus from Belgium
Distance between EF and GH distinctly larger then between other ocelli
2024.04.25 © Huskens, M.-L.
Deuterosminthurus pallipes f. repandus from Belgium
Footcomplex with clavate tenent macrosetae
2024.04.25 © Huskens, M.-L.

            Syn.: Deuterosminthurus Fényesi(sic) Stach, 1926 IOS
            Syn.: Deuterosminthurus Fényesi(sic) Stach, J, 1930:271 IOS
             Syn.: Deuterosminthurus Fenyesi(sic) Denis, J-R, 1933:300 ISS
Deuterosminthurus pallipes fenyesi from Rhodos
"Abnormal pigmentation"
After Ellis WN, 1974 Fig.6e

             Syn.: Deuterosminthurus feneysi(sic) Nayrolles, P, 1996:536 ISS
Deuterosminthurus pallipes f. fenyesi ♀ from France
2010.05.24 © Elpachato.
Deuterosminthurus pallipes f. fenyesi ♀ from France
2010.06.04 © Elpachato.
Caudal view
Deuterosminthurus pallipes ♂ & f. fenyesi ♀ from France
Courtship ritual
2010.06.04 © Elpachato.
Deuterosminthurus pallipes f. fenyesi ♀ from France
2010.06.16 © Elpachato.
Deuterosminthurus pallipes f. fenyesi ♀ from France
2018.03.26 © Tiky, S.
Deuterosminthurus pallipes f. fenyesi ♀ from France
Caudal aspect
2018.03.26 © Tiky, S.
Deuterosminthurus pallipes f. fenyesi from France
2018.05.25 © Picard, J.
Deuterosminthurus pallipes f. fenyesi from France
2018.06.05 © Picard, J.
Deuterosminthurus pallipes f. fenyesi from France
Dorso-caudal aspect
2018.06.05 © Picard, J.
Deuterosminthurus pallipes f. fenyesi from France
2019.05.30 © Picard, J.
Deuterosminthurus pallipes f. fenyesi from France
With "S" pattern of spots
2019.05.30 © Picard, J.
Deuterosminthurus pallipes f. fenyesi from France
2020.04.09 © Nogues, Be.

            Syn.: Bourletiella (Deuterosminthurus) mixta Gisin, 1946
            Syn.: Deuterosminthurus beckeri Coineau & Delamare Deboutteville, 1961
            Syn.: Deuterosminthurus pallipes ssp. fenyesi f. pallida(sic) Ellis, WN, 1974:142 IOS
            Syn.: Deuterosminthurus pallipes ssp. fenyesi f. pallida(sic) Ellis, WN, 1978:62 IOS
Deuterosminthurus fenyesi f. pallidus from Greece
2008.01.14 © Markopoulos, S.

            Syn.: Deuterosminthurus maculatus Nayrolles, P, 1996:543 nec Womersley, 1939
Deuterosminthurus maculatus from France
After Nayrolles P, 1996 Fig.21

            Syn.: Deuterosminthurus bellingeri Nayrolles, P, 1997:473
Deuterosminthurus pallipes var. bellingeri from the UK
2009.07.10 © Ardron, P.A.
Deuterosminthurus pallipes var. bellingeri ♂ from the UK
2009.07.10 © Ardron, P.A.

            Syn.: Deuterosminthurus bicinctus Sys, S, Weißbach, S, Jakob, L, Gerber, S, et Schneider, C, 2022:2738
Deuterosminthurus bicinctus from Germany
After Sys S & al, 2022 Fig.1c
= Deuterosminthurus pallipes f. repandus juv.




Smynthurus(sic) pallipes from the UK
After Lubbock J, 1870 Pl.XXI Fig.13
Smynthurus(sic) pallipes from the UK
Antenna
After Lubbock J, 1870 Pl.XXI Fig.14
Smynthurus(sic) pallipes from the UK
Saltatory appendage
After Lubbock J, 1870 Pl.XXI Fig.15
Smynthurus(sic) pallipes from the UK
After Lubbock J, 1873 Pl.4

             Syn.: Sminthurus palliceps(sic) Reuter, OM, 1892:232 lapsus ISS
Deuterosminthurus pallipes from the UK
2005 © Hopkin, S.P.
Deuterosminthurus pallipes from the UK
2006 © Barton, T.
Deuterosminthurus pallipes ♂ from the UK
2007.04.28 © Barton, T.
Deuterosminthurus pallipes from the UK
2007.06.03 © Saul, R.
Deuterosminthurus pallipes from the UK
2007.06.20 © Barton, T.
Deuterosminthurus pallipes from the UK
2008.06.02 © Farrell, S.
Deuterosminthurus pallipes juv. from the UK
2009.04.24 © Valentine., B
Deuterosminthurus pallipes from the UK
2009.05.03 © Barton, T.
Deuterosminthurus pallipes from the UK
2009.05.03 © Barton, T.
Deuterosminthurus pallipes ♂ from the UK
2009.07.16 © Dawes, D.
Deuterosminthurus pallipes from the UK
Male with female forma repandus
2010.05.17 © Barton, T.
Deuterosminthurus pallipes from the UK
2010.05.17 © Barton, T.
Deuterosminthurus pallipes ♂ & ♀ from the UK
2012.05.12 © Murray, A.
Deuterosminthurus pallipes from the UK
2012.05.13 © Murray, A.
 
Deuterosminthurus pallipes from the UK
Pale colour form
2012.05.13 © Murray, A.
Deuterosminthurus pallipes ♀ & ♂ from the UK
2012.05.29 © Nurcombe, E.
Deuterosminthurus pallipes ♂ & ♀ from the UK
2012.06.01 © Nurcombe, E.
Deuterosminthurus pallipes ♂ from the UK
Raised genital papilla
2012.06.01 © Nurcombe, E.
Deuterosminthurus pallipes ♂ f. repandus & ♀ from the UK
2012.06.15 © Nurcombe, E.
Deuterosminthurus pallipes from the UK
2013.07.11 © Nurcombe, E.
Deuterosminthurus pallipes ♀ from the UK
2014.04.26 © Henton-Jones, D.
Deuterosminthurus pallipes ♀ from the UK
Ant.4 with 8 subsegments
2014.04.26 © Henton-Jones, D.
Deuterosminthurus pallipes ♂ & ♀ from the UK
Courtship
2014.05.03 © Henton-Jones, D.
Deuterosminthurus pallipes ♀ from the UK
Eyepatch with 3 large inner ocelli
2014.05.03 © Henton-Jones, D.
Deuterosminthurus pallipes ♀ from the UK
Pinkish colour form
2017.04.18 © Henton-Jones, D.
Deuterosminthurus pallipes ♂ from the UK
Tentative courtship complex of 4 males
2017.04.18 © Henton-Jones, D.
Deuterosminthurus pallipes ♂ & f. repandus ♀ from the UK
Courtship between 2 colour forms
2017.04.18 © Henton-Jones, D.
Deuterosminthurus pallipes ♀ & f. repandus ♂ from the UK
Courtship between 2 colour forms
2017.05.02 © Henton-Jones, D.
Deuterosminthurus pallipes ♀ from Belgium
Fight? with D. pallipes f. repandus ♀
2017.07.11 © Huskens, M.-L.
Deuterosminthurus pallipes ♀ juv. from France
2017.11.01 © Tiky, S.
Deuterosminthurus pallipes from France
Reddish colour form
2018.06.03 © Picard, J.
Deuterosminthurus pallipes juv. from Belgium
2021.07.30 © Huskens, M.-L.
Deuterosminthurus pallipes subadult from Belgium
2021.07.31 © Huskens, M.-L.
Deuterosminthurus pallipes ♀ & ♂ from the USA
Courtship. ♀ = f. repandus
First record for the USA
2022.05.23 © Lopez, A.





Species pleuracanthus Nayrolles, P, 1996:537

Species polenus (Christiansen, K et Bellinger, P, 1992:360)

Species propallipes (Börner, C, 1901:107) Stach, 1956

Species prospathaceus (Rapoport in Delamare Deboutteville & Massoud, 1963) Mari Mutt & Bellinger, 1990 nn

Species quadrangulatus (Loksa, I & Bogojevic,J, 1970:140) Ellis, WN, 1974:146

Bourletiella (Deuterosminthurus) quadrangulata from Jugoslavia
After Loksa I & Bogojevic J, 1970 Abb.61
             Syn.: Deuterosminthurus quadrangulata(sic) Ellis, WN, 1974:146 ISS
             Syn.: Deuterosminthurus quadranculata(sic) Nayrolles, P, 1996:549 ISS


Species russatus (Maynard, EA, 1951:258) Stach, J, 1956:160

Bourletiella (Deuterosminthurus) russata from the USA
After Maynard, E.A, 1951 Plate XXIII Fig.406
             Syn.: Deuterosminthurus russata(sic) Snider, RJ, 1967:228 ISS


Species separatus Arlé, R, 1943:112

Species serratus Kang J-S et Lee B-H in Kang, J-S, Lee, B-H et Park, K-H, 2005:238

Deuterosminthurus serratus from Korea
After Kang J-S, Lee B-H & Park K-H, 2005 Fig.2A

Species sp. nov. González, V, Díaz, M et Palacios-Vargas, JG, 1997:48

Species sulphureus (Koch, 1840) Stach, 1956:159, B

            Syn.: Smynthurus(sic) lupulinae Bourlet, 1841
            Syn.: Smynthurus(sic) baulastinus Harvey, 1848
            Syn.: Smynthurus(sic) solani Curtis, 1860
            Syn.: Smynthurus(sic) luteus Lubbock, J, 1870:296
Smynthurus(sic) luteus from the UK
After Lubbock J, 1870 Pl.XXI Fig.4
Smynthurus(sic) luteus from the UK
Antenna
After Lubbock J, 1870 Pl.XXI Fig.5
Smynthurus(sic) luteus from the UK
Foot with 2 tenent hairs
After Lubbock J, 1870 Pl.XXI Fig.6
Smynthurus(sic) luteus from the UK
Saltatory appendage
After Lubbock J, 1870 Pl.XXI Fig.7
Smynthurus(sic) luteus from the UK
After Lubbock J, 1873 Pl.3

             Syn.: Sminthurus luteo(sic) Reuter, OM, 1876:84 ISS
             Syn.: Bourletiella (Bourletiella) luteus(sic) Kloet & Hincks, 1964 ISS
             Syn.: Bourletiella luteus(sic) Janssens, F, 1995:19,B ISS



            Syn.: Smynthurus(sic) vitis Lichtenstein, 1883
            Syn.: Sminthurus fulvus Lie-Pettersen, 1896
            Syn.: Sminthurus repandus Ågren, 1902
            Syn.: Corynephoria gibbera Salmon, JT, 1941:416
Corynephoria gibbera from New Zealand
After Salmon JT, 1941 Pl.72 Fig.521

             Syn.: Corynephoria gibbosa(sic) Stach, J, 1956:133 ISS
            Syn.: Deuterosminthurus Gisini(sic) Stach, 1956



Neozygites sp. from Denmark. Bar=10µm
Conidia infecting Deuterosminthurus sulphureus
After Dromph & al, 2001 Fig.5
Deuterosminthurus sulphureus from the UK
Annulated fourth antennal segment
2004 © Hopkin, S.P.

             Syn.: Deuterosminthurus sulphures(sic) Fjellberg, A, 2007:216 ISS
             Syn.: Deuterosminthurus sulphereus(sic) Berg, M et den Ouden, A, 2013:5 ISS
Deuterosminthurus sulphureus ♂ from France
Ant.1+2 without spine-like setae
Face laterally without enlarged setae
2021.07.12 © Picard, J.
Deuterosminthurus sulphureus ♂ from France
Abd.6 with enlarged distal setae
2021.07.12 © Picard, J.
Deuterosminthurus sulphureus ♂ from France
Antenna distally fading more reddish
2021.07.12 © Picard, J.
Deuterosminthurus sulphureus ♂ from France
Ocelli C+D not smaller then others
2021.07.12 © Picard, J.

  Subspecies mediterraneus Ellis, 1974



Species tristani Denis, J-R, 1933:297

            Syn.: Deuterosminthurus Tristani(sic) Denis, J-R, 1933:297 IOS
Deuterosminthurus Tristani(sic) ♀ from Costa Rica
After Denis J-R, 1933 Fig.113
Deuterosminthurus Tristani(sic) ♀ from Costa Rica
After Denis J-R, 1933 Fig.114
Deuterosminthurus Tristani(sic) ♂ from Costa Rica
After Denis J-R, 1933 Fig.115

            Syn.: Deuterosminthurus Tristani(sic) Denis, J-R, 1933:298 IOS
            Syn.: Deuterosminthurus Tristani(sic) Denis, J-R, 1933:299 IOS
            Syn.: Deuterosminthurus Tristani(sic) Denis, J-R, 1933:300 IOS
            Syn.: Deuterosminthurus Tristani(sic) Denis, J-R, 1933:315 IOS
            Syn.: Deuterosminthurus Tristani(sic) Denis, J-R, 1933:320 IOS


Species yumanensis Wray, 1967


Genus Karakumiella Bretfeld, G, 2010:310

Species kaplini (Martynova in Martynova & al, 1979 tt) Bretfeld, G, 2010:310


Genus Arlesminthurus Bretfeld, G, 1999:219 key

Species aueti (Arlé, R, 1961:37) Bretfeld, G, 1999:219

Species caatinguensis Cipola NG, Lycariao TA et Medeiros GDS in Cipola, NG, Medeiros, GDS, Oliveira, VAMD, Barbosa, LGDM, Lycariao, TA et Bellini, BC, 2021:3

Arlesminthurus caatinguensis ♀ from Brazil
After Cipola NG & al, 2021 Fig.4A
Arlesminthurus caatinguensis ♀ from Brazil
After Cipola NG & al, 2021 Fig.4B
Arlesminthurus caatinguensis ♀ from Brazil
After Cipola NG & al, 2021 Fig.4C

Species franzkafkai Palacios-Vargas, JG et Cabrera, M, 2015:63

Arlesminthurus franzkafkai ♀ from Nicaragua
After Palacios-Vargas JG & Cabrera M, 2015 Fig.1
Arlesminthurus franzkafkai ♂ from Nicaragua
After Palacios-Vargas JG & Cabrera M, 2015 Fig.2

Species richardsi (Arlé, 1971) Bretfeld, G, 1999:219, t.t.

Arlesminthurus nr richardsi ♂ from Costa Rica
2013.07.21 © Kouri, J.D.

Species salinensis (Arlé, 1971) Bretfeld, G, 1999:219


Genus Ellisiella Bretfeld, 1999

Species pandayi (Ellis, WN, 1974:143) Bretfeld, 1999, t.t.


Genus Corynephoria Absolon, 1907

Species absoloni Womersley, 1939

Corynephoria absoloni ♀ from Australia
With dorsal digitation
2015.11.09 © Murray, A.
Corynephoria absoloni ♂ from Australia
With dorsal digitation
2015.11.09 © Murray, A.
Corynephoria absoloni from Australia
2015.11.09 © Murray, A.

Species albidorsalis Womersley, H, 1932:36

Corynephoria albidorsalis from Australia
After Womersley H, 1932 Fig.14a
             Syn.: Corynephoria albodorsalis(sic) Stach, J, 1956:133 ISS


Species cassida Womersley, H, 1932:34

Corynephoria cassida from Australia
After Womersley H, 1932 Fig.12a

Species dubia Womersley, H, 1932:35

Species jacobsoni Absolon, 1907, t.t.

Species quadrimaculata Womersley, H, 1942:31

Corynephoria quadrimaculata from Australia
After Womersley H, 1942 Fig.8A

Species reticulata Salmon, 1963


Genus Bourletiella Banks, 1899, n.c. ICZN Opinion 291 key

  Syn.: Poduridæ(sic) Symnthurus(sic) Fitch, A, 1863:668 ISS
  Syn.: Poduridæ(sic) Symnthurus(sic) Fitch, A, 1863:669 ISS
  Syn.: Poduridæ(sic) Symnthurus(sic) Fitch, A, 1863:671 ISS
  Syn.: Poduridæ(sic) Symnthurus(sic) Fitch, A, 1863:672 ISS
  Syn.: Poduridæ(sic) Symnthurus(sic) Fitch, A, 1863:673 ISS
  Syn.: ... Symnthurus(sic) Fitch in Van Benthuysen, 1867 ISS
  Syn.: Sminthuridæ(sic) Sminthurus Schött, H, 1891:7 partim
  Syn.: ... Symnthurus(sic) Porter, 1894 ISS
  Syn.: Thysanura Symnthurus(sic) NewYorkStateMuseum, 1895 ISS
  Syn.: ... Symnthurus(sic) Porter, 1896 ISS
  Syn.: ... Symnthurus(sic) NewYorkStateMuseum, 1902 ISS
  Syn.: Sminthuridae Symnthurus(sic) Barnes, JK, 1988:89 ISS
  Syn.: Sminthuridae Symnthurus(sic) Barnes, JK, 1988:90 ISS
  Syn.: Sminthurus Pilosi Schött, H, 1891:11 partim
  Syn.: Collembola Sminthurinus(sic) Schött, H, 1917:59 lapsus


Remark by ICZN. 2007:112: Placed on the Official List of Names in Zoology. Opinion 291.

  Syn.: Bourletiellidae Borletiella(sic) Nickerl, J, Helbig, R, Schulz, H-J, Werner, C et Neinhuis, C, 2012:7 ISS
Bourletiella sp. from Australia
2015.11.22 © Steve & Alison.
Bourletiella sp. from Australia
2015.11.22 © Steve & Alison.
Bourletiella hortensis f. albamaculata from Belgium
Bothriotricha A-B-C in straight line
2019.09.03 © Huskens, M.-L.
Bourletiella hortensis f. albamaculata from Belgium
Footcomplex with knobbed sticky setae (*)
2019.09.03 © Huskens, M.-L.
Bourletiella sp. from Taiwan
2022.01.12 © Cheng, H.-J.
Bourletiella sp. from Taiwan
2022.01.13 © Cheng, H.-J.
Bourletiella sp. from Taiwan
2022.01.13 © Cheng, H.-J.
Bourletiella sp. from Taiwan
Tripod grooming
2022.01.13 © Cheng, H.-J.
Bourletiella sp. juv. from Taiwan
2022.01.14 © Cheng, H.-J.
Bourletiella sp. juv. from Taiwan
Bothriotricha ABC in straight line
2022.01.14 © Cheng, H.-J.
Bourletiella sp. from Taiwan
2022.01.14 © Cheng, H.-J.
Bourletiella sp. from Taiwan
2022.01.14 © Cheng, H.-J.
Bourletiella sp. from Taiwan
Eye with 8 equally sized ocelli
2022.01.14 © Cheng, H.-J.
Bourletiella sp. from Taiwan
Ant.4 with about 8 subdivisions
2022.01.14 © Cheng, H.-J.



Species sp.1 Janssens F sensu Brand & Dunn, 1998

            Syn.: Bourletiella sp.2 Brand, RH et Dunn, CP, 1998:238


Species sp.2 Janssens F sensu Christiansen & Bellinger, 1998

            Syn.: Bourletiella (Bourletiella) sp.A Christiansen & Bellinger, 1998
            Syn.: Bourletiella (Bourletiella) sp.new Rebek, EJ, Young, DK et Hogg, DB, 1999:61


Species sp.3 Janssens F senu Culik, de Souza & Ventura, 2002

            Syn.: Bourletiella sp.1 Culik, de Souza & Ventura, 2002


Species arvalis (Fitch, A, 1863:673) Tömösvary, 1883

            Syn.: Symnthurus(sic) arvalis Fitch, A, 1863:673
            Syn.: Smynthurus(sic) luteus Lubbock, 1867
            Syn.: Smynthurus(sic) roseus Packard, AS, 1873:43
            Syn.: Bourletiella hortensis f. hispanica Denis, 1930
            Syn.: Deuterosminthurus neopandus Wray, 1950
            Syn.: Bourletiella (Bourletiella) savona Maynard, EA, 1951:249 partim
             Syn.: Bourletiella repandus(sic) Scott, HG, 1963:107 ISS


Bourletiella arvalis from Tasmania
After Womersley, 1931 Pl.I Fig.5
Bourletiella arvalis from India
After Denis JR, 1936 Fig.49
Bourletiella arvalis from the USA
After Maynard, E.A, 1951 Plate XXIII Fig.407
Remark by Stach, J. 1956:147-148: In fig.567 given by Maynard, in, 1951, is the crest [of the male clasping organ] entirely wanting.

Bourletiella arvalis from the USA
2006 © McClarin, J.
Bourletiella arvalis from France
2007.04.16 © Lebeaux, P.
Bourletiella arvalis from Canada
2007.05.19 © Pilon, C.
Bourletiella arvalis from France
2007.06.18 © Lebeaux, P.
Bourletiella arvalis from Germany
2007.08.15 © Maier, T.
Bourletiella arvalis from the UK
2007.10.18 © Campbell, A.
Bourletiella arvalis from the USA
2008.05.11 © Thombs, D.
Bourletiella arvalis from the USA
2008.05.16 © Thombs, D.
Bourletiella arvalis from the USA
2008.05.18 © Thombs, D.
Bourletiella arvalis from the USA
2008.05.18 © Thombs, D.
Bourletiella arvalis from the USA
2008.06.02 © Thombs, D.
Bourletiella arvalis from the USA
Abdomen, lateral aspect
2008.06.16 © Thombs, D.
Bourletiella arvalis from the USA
2008.06.18 © Cowen, R.
 
Bourletiella arvalis from France
2008.10.06 © Lebeaux, P.
Bourletiella arvalis from France
2008.10.06 © Lebeaux, P.
Bourletiella arvalis from France
2008.10.19 © Lebeaux, P.
Bourletiella arvalis from the USA
2009.04.12 © Justis, S.
Bourletiella arvalis from the USA
2009.04.12 © Justis, S.
Bourletiella arvalis from the USA
2009.04.12 © Justis, S.
Bourletiella arvalis from the USA
2009.04.12 © Justis, S.
Bourletiella arvalis from the USA
2009.06.15 © Thombs, D.
Bourletiella arvalis from Tasmania
2009.08.16 © Daley, A.
Bourletiella arvalis from the USA
2011.08.23 © Carr, J.
Bourletiella arvalis from the UK
2012.08.06 © Nurcombe, E.
Bourletiella arvalis from Holland
2012.09.04 © van Duinen, J.
Bourletiella arvalis from New Zealand
2013.12.28 © Sullivan, J.
Bourletiella arvalis from Germany
2014.05.05 © Beck, K.
Bourletiella arvalis from Germany
Ant.4 with 7 annulations
2014.05.06 © Beck, K.
Bourletiella arvalis from the UK
2014.08.22 © Nurcombe, E.
Bourletiella arvalis ♂ from France
2019.06.06 © Garcelon, P.
Bourletiella arvalis ♀ from Belgium
2019.06.18 © Huskens, M.-L.
Bourletiella arvalis ♀ from Belgium
Female genital papilla (*)
2019.06.18 © Huskens, M.-L.
Bourletiella arvalis ♂ from the USA
2020.05.10 © Solomon.

Species captis Baijal, 1969

Species christianseni Snider, 1978

Bourletiella (Bourletiella) christianseni from the USA
After Christiansen K & Bellinger P, 1981 Fig.963A
             Syn.: Bourletiella Christianseni(sic) Fjellberg, A, 1984:207 ISS


Species drachi Delamare Deboutteville & Massoud, 1963:263

            Syn.: Bourletiella Drachi(sic) Delamare Deboutteville & Massoud, 1963 IOS

Species friedlaenderi (Börner, 1903) Stach, 1956:141

Species gibbonsi Snider, RJ, 1981:93

Bourletiella (Bourletiella) gibbonsi from the USA
After Snider RJ, 1981 Fig.1
Bourletiella (Bourletiella) gibbonsi from the USA
After Snider RJ, 1981 Fig.2

             Syn.: Bourletiella (Bourletiella) gibbons(sic) Christiansen, K et Bellinger, P, 1998:1304 ISS
Bourletiella gibbonsi from the USA
2017.04.05 © Fritzl.
Bourletiella gibbonsi from the USA
2017.04.05 © Fritzl.



Species guevarai Rapoport, 1970

Species hortensis (Fitch, A, 1863:668) Brown, 1918, t.t., B

Symnthurus(sic) hortensis from the USA
After Fitch A, 1863 p.670 left
Symnthurus(sic) hortensis from the USA
Antenna
Ant.4 with 7 subdivisions
Modified after Fitch A, 1863 p.670 right

            Syn.: Podura signata Fabricius, 1783
Smynthurus(sic) signatus from Switzerland
After Nicolet H, 1842 Pl.9 Fig.7

            Syn.: Smynthurus(sic) sp. Harris, 1844
            Syn.: Achetidæ(sic) Myrmecophila Harris, 1847 =?
            Syn.: Achetidæ(sic) Myrmecophila Fitch, A, 1863:668
             Syn.: Bourletiella spinata(sic) Lucic, 2006 lapsus ISS
             Syn.: Bourletiella spinata(sic) Grujic, NZ, Winkler, D et Daghighi, E, 2023:751 ISS
            Syn.: Smynthurus(sic) fuliginosus Nicolet, 1847
             Syn.: Smynthurella fuliginosus(sic) Houlbert, C, 1924:159 ISS
            Syn.: Smynthurus(sic) solani Curtis, 1860
            Syn.: Symnthurus(sic) hortensis var. apicalis Fitch, A, 1863:671
            Syn.: Symnthurus(sic) hortensis ssp. apicalis Barnes, JK, 1988:89
            Syn.: Smynthurus(sic) apicalis Poole, RW in Poole, RW et Gentili, P, 1997:54
            Syn.: Symnthurus(sic) hortensis var. basalis Fitch, A, 1863:671
            Syn.: Symnthurus(sic) hortensis ssp. basalis Barnes, JK, 1988:89
            Syn.: Smynthurus(sic) basalis Poole, RW in Poole, RW et Gentili, P, 1997:54
            Syn.: Symnthurus(sic) hortensis var. dorsalis Fitch, A, 1863:671
            Syn.: Symnthurus(sic) hortensis ssp. dorsalis Barnes, JK, 1988:89
            Syn.: Smynthurus(sic) dorsalis Poole, RW in Poole, RW et Gentili, P, 1997:54
            Syn.: Symnthurus(sic) hortensis var. juvenilis Fitch, A, 1863:671
            Syn.: Symnthurus(sic) hortensis ssp. juvenilis Barnes, JK, 1988:90
            Syn.: Smynthurus(sic) juvenilis Poole, RW in Poole, RW et Gentili, P, 1997:54
            Syn.: Symnthurus(sic) hortensis var. maturus Fitch, A, 1863:671
            Syn.: Symnthurus(sic) hortensis ssp. maturus Barnes, JK, 1988:90
            Syn.: Smynthurus(sic) maturus Poole, RW in Poole, RW et Gentili, P, 1997:54
            Syn.: Smynthurus(sic) cucumeris Harris, 1869
            Syn.: Sminthurus pruinosus Tullberg, 1871
Sminthurus pruinosus from Sweden
After Schött H, 1893 Taf.II Fig.13

            Syn.: Bourletiella pruinosa Folsom, 1899
Sminthurus pruinosus from Europe
After Folsom in Escherich K, 1923 Abb.1b

            Syn.: Sminthurus pruinosus Matsumura, S, 1931:1496
            Syn.: Bourletiella pruinosa Kinoshita, S, 1932:2116
Bourletiella pruinosa from Japan
After Kinoshita S, 1932 X23

            Syn.: Bourletiella pruinosa Uchida, 1940
            Syn.: Bourletiella pruinosa Kinoshita & Uchida, 1950
            Syn.: Sminthurus pruinosus Yosii, R, 1977:164
            Syn.: Bourletiella pruinosa Yosii, R, 1977:164
            Syn.: Smynthurus(sic) luteus Lubbock, 1873
            Syn.: Smynthurus(sic) 4-signatus(sic) Packard, AS, 1873:44 IOS
            Syn.: Sminthurus lineatus Reuter, OM, 1876:84
            Syn.: Smynthurus frontalis Uzel, 1890
            Syn.: Smynthurus(sic) signifer MacGillivray, 1891
            Syn.: Smynthurus(sic) albamaculata(sic) Harvey, FL, 1897:124 IOS
Smynthurus(sic) albamaculata(sic) from the USA
After Emerton JH in Harvey FL, 1897 Fig.1
Smynthurus(sic) albamaculata(sic) from the USA
Ventral aspect with everted collophore vesicles
After Emerton JH in Harvey FL, 1897 Fig.2
Smynthurus(sic) albamaculata(sic) from the USA
Dorsal aspect of pattern of abdominal patches
After Harvey FL, 1897 Fig.3
Smynthurus(sic) albamaculata(sic) from the USA
Lateral aspect of pattern of abdominal patches
After Harvey FL, 1897 Fig.4

             Syn.: Smynthurus(sic) albamaculata(sic) Folsom, JW, 1899:269 ISS
             Syn.: Sminthurus albamacula(sic) Reuter, 1900 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Patch, 1905 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Patch, 1906 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Patch, 1911 ISS
             Syn.: Sminthurus albamaculata(sic) Maynard, EA, 1951:254 ISS
             Syn.: Sminthurus albomaculatus(sic) Obenberger, 1955 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Paclt, 1956 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Stach, J, 1956:149 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Stach, J, 1956:237 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Salmon, JT, 1964:613 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Salmon, JT, 1964:614 ISS
             Syn.: Sminthurus albamaculata(sic) Salmon, JT, 1964:613 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Mari Mutt, 1978 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Christiansen, K et Bellinger, P, 1981:1147 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Christiansen, K et Bellinger, P, 1981:1151 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Christiansen, K et Bellinger, P, 1981:1152 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Christiansen, K et Bellinger, P, 1981:1153 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Poole, RW in Poole, RW et Gentili, P, 1997:54 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Christiansen, K et Bellinger, P, 1998:1307 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Christiansen, K et Bellinger, P, 1998:1311 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Christiansen, K et Bellinger, P, 1998:1312 ISS
             Syn.: Smynthurus(sic) albamaculata(sic) Christiansen, K et Bellinger, P, 1998:1313 ISS
Bourletiella hortensis f. albamaculata from the UK
2012.06.21 © Murray, A.
Bourletiella hortensis f. albamaculata ♂ subadult from Belgium
2017.08.20 © Huskens, M.-L.
Bourletiella hortensis f. albamaculata from Belgium
2017.08.25 © Huskens, M.-L.
Bourletiella hortensis f. albamaculata juv. from Belgium
Two different instars
2017.09.02 © Huskens, M.-L.
Bourletiella hortensis f. albamaculata ♂ from Belgium
Male genital papilla (gp)
and median crest and modified setae (*)
2018.05.28 © Huskens, M.-L.
Bourletiella hortensis f. albamaculata juv. from Belgium
Bicoloured: pale head, dark abdomen
2018.05.28 © Huskens, M.-L.
Bourletiella hortensis f. albamaculata ♂ & ♀ from Belgium
Courtship: ♀ follows ♂
2018.06.21 © Huskens, M.-L.
Bourletiella hortensis f. albamaculata ♂ & ♀ from Belgium
Courtship: ♀ stimulates abd.6 modified setae of ♂
2018.06.25 © Huskens, M.-L.
Bourletiella hortensis f. albamaculata from Belgium
2018.07.26 © Huskens, M.-L.
Bourletiella hortensis f. albamaculata from Belgium
2018.07.26 © Huskens, M.-L.
Bourletiella hortensis f. albamaculata from Belgium
2019.09.03 © Huskens, M.-L.

            Syn.: Sminthurus atratus Schäffer, 1896
            Syn.: Sminthurus pruinosus var. sabulosa Skorikov, 1899
            Syn.: Sminthurus pruinosus var. stscherbakowi Skorikov, 1899
            Syn.: Sminthurus pruinosus var. viridula Skorikov, 1899
            Syn.: Bourletiella hortensis f. Linnaniemi(sic) Denis, 1930
            Syn.: Bourletiella hortensis f. Folsomi(sic) Denis, 1930
            Syn.: Bourletiella insula Folsom, JW, 1932:73
             Syn.: Podura fusco-nigra(sic) Salmon, JT, 1964:612 ISS
            Syn.: Bourletiella viridis Greenslade, 2002



Sminthurus hortensis from Japan
After Folsom JW, 1899 Pl.3 Fig.37
Smynthurus(sic) hortensis from Japan
After Matsumura S, 1907 Pl.1 Fig.10
Bourletiella hortensis from Tasmania
After Womersley, 1931 Pl.I Fig.6
Bourletiella hortensis from Australia
After Greenslade P & Ireson JE, 1986 Fig.9
Bourletiella hortensis from Sweden
2005 © Hall, K.
Bourletiella hortensis from the USA
2006 © McClarin, J.
Bourletiella hortensis from Canada
2007.05.03 © Pilon, C.
Bourletiella hortensis from Canada
2007.05.04 © Pilon, C.
Bourletiella hortensis from the USA
2007.05.06 © Schimming, L.
Bourletiella hortensis from the USA
2007.05.10 © Murray, T.
Bourletiella hortensis from the USA
2007.05.10 © Murray, T.
Bourletiella hortensis from the USA
2007.05.10 © Murray, T.
Bourletiella hortensis from France
2007.05.14 © Lebeaux, P.
Bourletiella hortensis from the USA
2008.04.23 © Bradford, A.
Bourletiella hortensis from Hungary
2008.06.21 © Keresztes, G.
Bourletiella hortensis from Antarctica
2009.03.20 © Key, R.S.
Det. Greenslade P.
Bourletiella hortensis from Antarctica
2009.03.20 © Key, R.S.
Det. Greenslade P.
Bourletiella hortensis from the USA
2009.05.06 © Murray, T.
Bourletiella hortensis from the USA
2009.05.18 © Murray, T.
Bourletiella hortensis from the USA
2009.05.18 © Murray, T.
Bourletiella hortensis from Belgium
2009.06.18 © Hermans, R.
Bourletiella hortensis from the UK
2010.06.26 © Michaelson, J.
Bourletiella hortensis from the UK
Subdivided 4th antennal segment
2010.07.26 © Robertson, A.
Bourletiella hortensis from the UK
Footcomplex with clavate tenent hairs
2010.07.26 © Robertson, A.
Bourletiella hortensis from France
2011.08.29 © Lebeaux, P.
 
Borletiella(sic) hortensis (bar=2µm)
Rhombic epicuticular ultrastructure with secondary granules
After Nickerl J & al, 2012 Fig.6E
Bourletiella hortensis from the UK
2012.05.28 © Jones, A.
Bourletiella hortensis juv. from the UK
2012.06.24 © Nurcombe, E.
 
Bourletiella hortensis from the UK
With parasitic larva of Acari:Erythraeoidea
2012.06.27 © Nurcombe, E.
Bourletiella hortensis from the UK
With parasitic larva of Acari:Erythraeoidea
2012.06.27 © Nurcombe, E.
Bourletiella hortensis from Belgium
Mucro spoonshaped
2012.10.29 © Vannieuwenhuyse., B
Bourletiella hortensis from the UK
Grooming using drop of saliva on front claw and everted tube-like vesicle of collophore
2013.05.25 © Nurcombe, E.
Bourletiella hortensis from the UK
2014.05.04 © Evans, M.
Bourletiella hortensis from Belgium
2017.08.27 © Huskens, M.-L.
Bourletiella hortensis ♂ & ♀ from Belgium
♀ stimulates ♂ by nibbling on its abd.6 setae
2018.09.01 © Huskens, M.-L.
Bourletiella hortensis from Belgium
Moulting
2018.09.12 © Huskens, M.-L.
Bourletiella ?hortensis? from Holland
2018.11.04 © van Duinen, J.
Bourletiella hortensis from Belgium
With parasitic Erythraeoidea nymph
2019.06.18 © Huskens, M.-L.
Bourletiella hortensis from France
Hatchling
2019.10.31 © Garcelon, P.
Bourletiella hortensis juv. from Belgium
Ocelli C+D distinctly smaller than other ocelli
2020.04.23 © Huskens, M.-L.
Bourletiella hortensis ♂ from Belgium
2020.06.21 © Huskens, M.-L.
Bourletiella hortensis ♂ from Belgium
Genital papilla (*)
2020.06.21 © Huskens, M.-L.
Bourletiella hortensis from Belgium
Feeding
2020.07.28 © Huskens, M.-L.
Bourletiella hortensis ♀ from Belgium
Bothriotricha ABC+DE
2020.07.30 © Huskens, M.-L.
Bourletiella hortensis from Belgium
Pale colour form
2021.07.07 © Huskens, M.-L.

  Subspecies melitensis Janssens F & Ujvári Z, 2018

Bourletiella hortensis ssp. melitensis from Malta
2018.11.02 © Ujvári, Z.
Bourletiella hortensis ssp. melitensis ♀ from Malta
2018.11.02 © Ujvári, Z.
Bourletiella hortensis ssp. melitensis ♀ from Malta
2018.11.02 © Ujvári, Z.
Bourletiella hortensis ssp. melitensis ♀ from Malta
2018.11.02 © Ujvári, Z.
Bourletiella hortensis ssp. melitensis ♀ from Malta
Eye with 8 ocelli; C+D small
2018.11.02 © Ujvári, Z.
Bourletiella hortensis ssp. melitensis from Malta
Ant.4 with long basal and 7 short apical subdivisions
2018.11.02 © Ujvári, Z.
Bourletiella hortensis ssp. melitensis ♂ from Malta
2018.11.02 © Ujvári, Z.
Bourletiella hortensis ssp. melitensis ♂ from Malta
Male genital papilla (*)
2018.11.02 © Ujvári, Z.
Bourletiella hortensis ssp. melitensis ♂ from Malta
Median crest and modified setae (*)
2018.11.02 © Ujvári, Z.



Species insula Folsom, JW, 1932:73

Species juanitae Maynard, EA, 1951:249

Bourletiella juanitae from the USA
After Maynard, E.A., 1951 Pl.XXIII Fig.387
Bourletiella juanitae from the USA
After Maynard, E.A., 1951 Pl.XXIII Fig.388

            Syn.: Bourletiella (Deuterosminthurus) fallonae Maynard, EA, 1951:255
Bourletiella fallonae from the USA
After Maynard, E.A., 1951 Pl.XXIII Fig.404
Bourletiella fallonae from the USA
After Maynard, E.A., 1951 Pl.XXIII Fig.405




             Syn.: Bourletiella jaunitae(sic) Richards, WR, 1968:33 ISS
Bourletiella juanitae ♀ from the USA
2007.08.30 © Murray T.
Bourletiella juanitae ♀ from the USA
2007.08.30 © Murray T.
Bourletiella juanitae ♂ from the USA
2007.09.05 © Murray T.
Bourletiella juanitae ♂ from the USA
2007.09.05 © Murray T.
Bourletiella juanitae ♂ from the USA
2007.09.05 © Murray T.
Bourletiella juanitae ♀ from the USA
2019.03.28 © Jimenez, A.
Bourletiella juanitae ♀ from the USA
2019.03.28 © Jimenez, A.
Bourletiella juanitae ♀ from the USA
2019.03.28 © Jimenez, A.
Bourletiella juanitae ♀ from the USA
2019.06.23 © Duh, E.C. in Casey, R.
Bourletiella juanitae ♀ from the USA
2019.06.23 © Duh, E.C. in Casey, R.
Bourletiella juanitae ♀ from the USA
2019.07.18 © Solomon.
Bourletiella juanitae ♀ from the USA
2021.11.29 © Wildwings.
Bourletiella juanitae ♀ from the USA
2021.11.29 © Wildwings.



Species khenifrana Bretfeld, G, 2001:93

Species meghalayensis Mandal, GP, 2018:114

Bourletiella meghalayensis from India
After Mandal GP, 2018 Plate.3 Fig.1

Species millsi Pedigo, 1968

Species pisifera Fjellberg, A, 1984:207

Species pistilla Gisin, 1946

            Syn.: Bourletiella pistillum(sic) Gisin, 1946 IOS
            Syn.: Bourletiella pruinosa Ågren, 1903
            Syn.: Bourletiella agreni Stach, J, 1956:140
            Syn.: Bourletiella (Deuterosminthurus) cruciata Haybach, 1972
            Syn.: Bourletiella mongolica Betsch, J-M, 1977:77
            Syn.: Bourletiella pistillum(sic) ssp. incisa Ellis, WN, 1975:77


             Syn.: Bourletiella pistillum(sic) Stach, J, 1956:140 ISS
             Syn.: Bourletiella (Bourletiella) pistillum(sic) Gisin, H, 1960:286 ISS
             Syn.: Bourletiella pistillum(sic) Ellis, WN, 1975:76 ISS
             Syn.: Bourletiella (Cassagnaudiella) pistillum(sic) Ellis, WN, 1975:78 ISS
             Syn.: Bourletiella (Cassagnaudiella) pistillum(sic) ssp. pistillum(sic) Ellis, WN, 1975:79 ISS
             Syn.: Bourletiella pistillum(sic) ssp. pistillum(sic) Ellis, WN, 1975:81 ISS
             Syn.: Bourletiella (Cassagnaudiella) pistillum(sic) Klaver, E, 1975:185 ISS
             Syn.: Bourletiella pistillum(sic) Rusek, J in Dlabola, J, 1977:16 ISS
             Syn.: Bourletiella pistillum(sic) Fjellberg, A, 1984:87 ISS
             Syn.: Bourletiella pistillum(sic) Nayrolles, P, 1993:661 ISS
             Syn.: Bourletiella pistillum(sic) Dallai, R, Malatesta, E et Ramellini, P in Minelli, A, Ruffo, S et La Posta, S, 1995:14 ISS
             Syn.: Bourletiella pistillum(sic) Nayrolles, P, 1995:47 ISS
             Syn.: Bourletiella (Bourletiella) pistillum(sic) ssp. pistillum(sic) Bretfeld, G, 1999:226 ISS
             Syn.: Bourletiella pistillum(sic) Auclerc, A, Ponge, JF, Barot, S et Dubs, F, 2009:1602 ISS
             Syn.: Bourletiella pistillum(sic) Dunger, W et Voigtländer, K, 2009:18 ISS
             Syn.: Bourletiella (Bourletiella) pistillum(sic) Schulz, H-J, 2015:17 ISS
             Syn.: Bourletiella pistillum(sic) Burkhardt, U, Schulz, H-J et Stark, A, 2016:13 ISS



Species radula Gisin, 1946

            Syn.: Bourletiella rustica Cassagnau, 1968 lapsus


Species rapoporti Delamare Deboutteville & Massoud, 1963:259

            Syn.: Bourletiella Rapoporti(sic) Delamare Deboutteville & Massoud, 1963 IOS

Species rustica Maynard, EA, 1951:248

Bourletiella rustica from the USA
After Maynard EA PlateXXII Fig.373
Bourletiella rustica from the USA
After Maynard EA PlateXXX Fig.568
Bourletiella rustica from the USA
After Maynard EA PlateXXX Fig.569
Bourletiella rustica from the USA
2015.11.24 © Snyder, S.L.
Bourletiella rustica from the USA
2015.11.24 © Snyder, S.L.
Bourletiella rustica from the USA
Feeding instar
2016.12.13 © Snyder, S.L.
Bourletiella rustica? from the USA
2019.07.19 © Doby, J.

Species savona Maynard, EA, 1951:249

Bourletiella savona from the USA
After Maynard EA, 1951 Pl.XXIII Fig.384

Species sicki Bretfeld, 1994

Species spectabilis Handschin, 1930

Species tullbergi (Reuter, OM, 1876:85) Stach, J, 1956:241, i.s.

            Syn.: Sminthurus Tullbergi(sic) Reuter, OM, 1876:85 IOS
             Syn.: Sminthurus Tullbergii(sic) Reuter, OM, 1895:5 ISS
            Syn.: Sminthurus Tullbergi(sic) Reuter, OM, 1895:6 IOS



Species venusta Bretfeld, G, 2001:94

Species viridescens Stach, 1920

            Syn.: Bourletiella arvalis var. fusca Womersley, H, 1932:27
            Syn.: Bourletiella arvalis ssp. dorsobscura Salmon, JT, 1941:415
Bourletiella arvalis
subsp. dorsobscura from New Zealand
After Salmon JT, 1941 Pl.71 Fig.516
Remark by Stach, J. 1956:148: Fig.515 of Bourletiella arvalis dorsobscura Salmon, 1941, from New Zealand, without crest [of male clasping organ].

Bourletiella viridescens f. dorsobscura from New Zealand
2007.12.29 © van de Vyver, L.

            Syn.: Bourletiella lutea Gisin, 1946, nec Lubbock, 1867
            Syn.: Bourletiella hortensis Stach, 1956, nec Fitch, 1863
            Syn.: Sminthurus fulvus Fjellberg, A, 2007:217



Bourletiella viridescens from Australia
After Greenslade P & Ireson JE, 1986 Fig.7
Bourletiella viridescens from Tasmania
After Ireson JE, 1990 Fig.42
Bourletiella viridescens from France
2007.05.14 © Lebeaux, P.
Bourletiella viridescens from France
2007.05.14 © Lebeaux, P.
Bourletiella viridescens from New Zealand
2007.12.21 © van de Vyver, L.
Bourletiella viridescens from New Zealand
2007.12.21 © van de Vyver, L.
Bourletiella viridescens from the UK
2009.08.20 © Ransom, T.
Bourletiella viridescens from the UK
2009.08.24 © Ransom, T.
Bourletiella viridescens from the UK
2013.08.01 © Garlick, T.
Bourletiella viridescens juv. from Holland
2015.08.21 © van Duinen, J.
Bourletiella viridescens from Holland
2015.08.28 © van Duinen, J.
Bourletiella viridescens from Holland
2015.08.28 © Wieringa, P.
Bourletiella viridescens from Holland
2015.08.28 © Wieringa, P.
Bourletiella viridescens from Holland
2015.09.18 © van Duinen, J.
Bourletiella viridescens from Holland
2016.09.03 © van Duinen, J.
Bourletiella viridescens from Greece
2017.03.04 © Dimitrakopoulos, C.


Genus Diksamella Bretfeld, G, 2005:13

Species pohli Bretfeld, G, 2005:13, t.t.

Diksamella pohli from Yemen
After Bretfeld G, 2005 Fig.22


Genus Nasosminthurus Stach, 1955

Species analis (Womersley, H, 1939:251) Greenslade P in Houston, 1994:127

Species dimorphus (Womersley, H, 1932:31) Stach, J, 1956:134, t.t.

Species nigridorsalis (Womersley, H, 1932:30) Greenslade, P in Houston, WWK, 1994:127

Species sigmoides (Womersley, 1933) Greenslade P in Houston, 1994:127


Genus Navarrella Bretfeld & Arbea, 2000

Species apicalis Bretfeld & Arbea, 2000, t.t.


Genus Orenius Bretfeld in Bretfeld, Poliakov & Broza, 2000

Species maculatus Bretfeld, G, 2005:16

Orenius maculatus from Yemen
After Bretfeld G, 2005 Fig.29

Species parvus Bretfeld in Bretfeld, Poliakov & Broza, 2000, t.t.


Genus Pseudobourletiella Stach, J, 1956:142

  Syn.: Thysanura Smynthurus(sic) MacGillivray, AD, 1893:127 partim ISS
  Syn.: Smynthuri(sic) Smynthurus(sic) Folsom, JW, 1896:449 partim ISS
  Syn.: ... Symnthurus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:54 lapsus ISS


Species spinata (MacGillivray, AD, 1893:127) Stach, J, 1956:142, t.t.

            Syn.: Smynthurus(sic) spinatus MacGillivray, AD, 1893:127,tt IOS
            Syn.: Pseudobourletiella chandleri Pedigo, 1968


             Syn.: Bourletiella spinatus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:54 ISS
Pseudobourletiella spinata from the USA
2006 © Web, S.
Pseudobourletiella spinata from Canada
2006.06.23 © Pilon, C.
Pseudobourletiella spinata ♂ from Canada
2006.07.23 © Pilon, C.
Pseudobourletiella spinata ♂ from Canada
2006.09.06 © Pilon, C.
Pseudobourletiella spinata from Canada, juvenile
2007.07.07 © Pilon, C.
Pseudobourletiella spinata from Canada
2007.07.18 © Pilon, C.
 
Pseudobourletiella spinata ♀ from the USA
2008.06.07 © Murray, T.
Pseudobourletiella spinata ♀ from the USA
2008.06.07 © Murray, T.
Pseudobourletiella spinata ♀ from the USA
2008.06.13 © Murray, T.
Pseudobourletiella spinata ♂ from the USA
2008.06.13 © Murray, T.
Pseudobourletiella spinata ♀ from the USA
2008.08.15 © Murray, T.
Pseudobourletiella spinata ♂ from the USA
2008.12.07 © Fleming, K.
Pseudobourletiella spinata ♀ from the USA
2009.02.01 © Fleming, K.
Pseudobourletiella spinata ♀ from the USA
2009.07.28 © Murray, T.
Pseudobourletiella spinata ♀ from the USA
2010.05.15 © Justis, S.
Pseudobourletiella spinata juv. from the USA
2010.05.15 © Justis, S.
Pseudobourletiella spinata juv. from the USA
2010.05.15 © Justis, S.
Pseudobourletiella spinata juv. from the USA
2010.05.15 © Justis, S.
Pseudobourletiella spinata juv. from the USA
2010.05.20 © Justis, S.
Pseudobourletiella spinata juv. from the USA
2010.05.20 © Justis, S.
Pseudobourletiella spinata juv. from the USA
2010.05.24 © Justis, S.
Pseudobourletiella spinata ♀ from the USA
2011.01.21 © Montgomery, G.
Pseudobourletiella spinata ♀ from the USA
2011.05.27 © Carr, J.
Pseudobourletiella spinata ♀ from the USA
2011.05.29 © Murray, T.
Pseudobourletiella spinata ♂ from the USA
2011.08.04 © Eiseman, C.
Pseudobourletiella spinata ♀ from the USA
2011.08.04 © Eiseman, C.
Pseudobourletiella spinata ♀ from the USA
2012.01.01 © Justis, S.
Pseudobourletiella spinata ♀ from the USA
2012.01.01 © Justis, S.
Pseudobourletiella spinata ♂ & ♀ from Canada
Courtship
2015.06.05 © Murray, A.
Pseudobourletiella spinata juv. ♂ from Canada
2015.06.07 © Murray, A.
Pseudobourletiella spinata ♂ from Canada
2015.06.07 © Murray, A.
Pseudobourletiella spinata ♀ from Canada
2015.06.07 © Murray, A.
Pseudobourletiella spinata ♀ from Canada
With dorsal thoracic bulge (*)
2015.06.07 © Murray, A.
Pseudobourletiella spinata ♀ from Canada
2015.06.07 © Murray, A.
Pseudobourletiella spinata ♀ juv. from the USA
2017.06.25 © Dobbs, Br.
Pseudobourletiella spinata ♀ & ♂ from Canada
Courtship
2018.09.23 © Hogue, A.
Pseudobourletiella spinata juv. from the USA
2019.08.12 © Solomon.
Pseudobourletiella spinata ♀ from the USA
2019.08.12 © Solomon.
Pseudobourletiella spinata juv. from the USA
2019.08.12 © Solomon.
Pseudobourletiella spinata ♀ from Canada
Thoracic pigmentation
2020.07.01 © Barbin, T.
Pseudobourletiella spinata ♀ from Canada
Predated by Dictyna minuta
2020.09.12 © Johnson, D.
Pseudobourletiella spinata ♂ & ♀ from Canada
Courtship
♀ stimulates ♂
2020.10.04 © Hogue, A.
Pseudobourletiella spinata ♂ from Canada
2020.10.04 © Hogue, A.
Pseudobourletiella spinata ♂♂ & ♀ from the USA
courtship
2023.04.15 © Shahan, T.






Genus Heterosminthurus Stach, 1955

  Syn.: Collembola Heteromurus(sic) Hopkin, SP, 1997:82 lapsus
Heterosminthurus insignis from the UK
Unguiculus of metaleg with distinct lamella
2008.06.30 © Ardron, P.A. (Det.)
Heterosminthurus insignis ♀ from Sweden
Unguiculus with distinctly large basal lamella (*)
2022.09.26 © Kamsteeg, G.



Species aquaticus (Maynard, EA, 1951:259)

Bourletiella (Deuterosminthurus) aquatica ♀
from the USA
After Maynard, E.A, 1951 Plate XXIII Fig.408
Bourletiella (Deuterosminthurus) aquatica ♂
from the USA
After Maynard, E.A, 1951 Plate XXIII Fig.409

            Syn.: Bourletiella (Heterosminthurus) insignis American authors
            Syn.: Deuterosminthurus insignis Mills, 1939
            Syn.: Deuterosminthurus insignis Hammer, M, 1953:59
            Syn.: Deuterosminthurus insignis Snider, RJ, 1969:358



Heterosminthurus aquaticus ♀ from the USA
2017.06.10 © Dobbs, Br.
Heterosminthurus aquaticus ♀ from the USA
Ant.4 with 7 subdivisions
2017.06.10 © Dobbs, Br.
Heterosminthurus aquaticus ♀ from the USA
2020.07.09 © Cam, C.
Heterosminthurus aquaticus ♀ from the USA
2020.07.09 © Cam, C.
Heterosminthurus aquaticus ♀ from the USA
2020.07.09 © Cam, C.
Heterosminthurus aquaticus ♀ from the USA
2020.07.09 © Cam, C.
Heterosminthurus aquaticus ♂ from the USA
2020.07.09 © Cam, C.
Heterosminthurus aquaticus ♀ from the USA
2020.07.13 © Cam, C.
Heterosminthurus aquaticus ♀ from the USA
2020.07.16 © Cam, C.
Heterosminthurus aquaticus ♀ from the USA
Bothriotricha ABC form large obtuse angle
2020.07.16 © Cam, C.
Heterosminthurus aquaticus ♀ from the USA
2020.07.16 © Cam, C.
Heterosminthurus aquaticus ♀ from the USA
Ant.4 with 7 subdivisions
2020.07.16 © Cam, C.

Species bilineatus (Bourlet, 1842) Stach, J, 1956:182, B

            Syn.: Smynthurus(sic) oblongus Nicolet, H, 1842:81
Smynthurus(sic) oblongus from Switzerland
After Nicolet H, 1842 Pl.9 Fig.8

            Syn.: Smynthurus(sic) Bourletii(sic) Gervais in Walkenaër, 1844 IOS
             Syn.: Smynthurus(sic) Bourletii(sic) Nicolet, H, 1847:359 ISS
             Syn.: Smynthurus(sic) Bourletii(sic) Lubbock, J, 1870:297 ISS
Smynthurus(sic) Bourletii(sic) from the UK
After Lubbock J, 1870 Pl.XXI Fig.8
Smynthurus(sic) Bourletii(sic) from the UK
Antenna
After Lubbock J, 1870 Pl.XXI Fig.9
Smynthurus(sic) Bourletii(sic) from the UK
Saltatory appendage
After Lubbock J, 1870 Pl.XXI Fig.10

             Syn.: Smynthurus(sic) Bourletii(sic) Lubbock, J, 1873:110 ISS
Smynthurus(sic) Bourletii(sic) from the UK
After Lubbock, J, 1873 Pl.5.


            Syn.: Deuterosminthurus coronatus Bagnall, 1921
            Syn.: Heterosminthurus cornutus Stach, J, 1956:190
Heterosminthurus cornutus from Poland
After Stach, J, 1956 Pl.XXII Fig.7




Heterosminthurus bilineatus from Poland
After Stach, J, 1956 Pl.XXII Fig.2
Heterosminthurus bilineatus from Poland
After Stach, J, 1956 Pl.XXII Fig.1
Heterosminturus bilineatus from Germany
Courtship ritual
After Bretfeld G, 1973 Fig.1B
             Syn.: Heterosminthurus bilinetus(sic) Bretfeld, G, 1999:262 ISS
Heterosminthurus bilineatus from the UK
Footcomplex, empodium with lamella
2004 © Hopkin, S.P.
Heterosminthurus bilineatus from the UK
2008.07.13 © Ardron, P.A. (Det.)
Heterosminthurus bilineatus from the UK
2008.07.13 © Ardron, P.A. (Det.)
Heterosminthurus bilineatus from the UK
2008.07.13 © Ardron, P.A. (Det.)
Heterosminthurus bilineatus from the UK, ♀
2008.07.13 © Ardron, P.A. (Det.)
Heterosminthurus bilineatus from the UK, ♂
Male interocular sinusoid setae
2008.07.13 © Ardron, P.A. (Det.)
Heterosminthurus bilineatus from France
2009.09.dd © Lebeaux, P.
Heterosminthurus bilineatus from Holland
2015.05.04 © van Duinen, J.
Heterosminthurus bilineatus ♀ from Belgium
Cleaning antenna
2017.08.28 © Huskens, M.-L.
Heterosminthurus bilineatus ♀ from Belgium
2017.08.31 © Huskens, M.-L.
Heterosminthurus bilineatus from France
2017.09.03 © Tiky, S.
Heterosminthurus bilineatus from France
Caudal aspect
2017.09.03 © Tiky, S.
Heterosminthurus bilineatus from France
2017.09.03 © Tiky, S.
Heterosminthurus bilineatus ♀ from France
2017.09.16 © Picard, J.
Heterosminthurus bilineatus ♀ from France
Back of head
2017.09.16 © Picard, J.
Heterosminthurus bilineatus ♀ from Belgium
4 flank setae (*)
2018.05.19 © Huskens, M.-L.
Heterosminthurus bilineatus ♂ & ♂ from Belgium
Courtship between 2 males...
2018.05.19 © Huskens, M.-L.
Heterosminthurus bilineatus from the UK
Caudal aspect
2018.05.20 © Cann, A.J.
Heterosminthurus bilineatus from the UK
Ventral aspect with exposed long tube-like collophore vesicles
2018.05.20 © Cann, A.J.
Heterosminthurus bilineatus ♀ & ♂ from Belgium
Courtship
2018.05.21 © Huskens, M.-L.
Heterosminthurus bilineatus from Holland
2018.08.23 © Kamsteeg, G.
Heterosminthurus bilineatus from Holland
2018.08.23 © Kamsteeg, G.
Heterosminthurus bilineatus from Holland
2018.08.23 © Kamsteeg, G.
Heterosminthurus bilineatus ♀ from France
Bothriotrichum C and 4 flank setae
2019.08.03 © Picard, J.
Heterosminthurus bilineatus ♀ & ♂ from Belgium
Courtship
Male stimulates female
2023.05.25 © Huskens, M.-L.

  Subspecies caucasicus Stach, 1956





Species borealis Bretfeld, G et Zöllner, E, 2000:136

Species chaetocephalus Hüther, 1971

Heterosminthurus chaetocephalus from Germany
Courtship ritual
After Bretfeld G, 1976

Species claviger (Gisin, 1958) Bretfeld, G, 1971:150, B

            Syn.: Bourletiella bilineata Linnaniemi, 1911 :
            Syn.: Deuterosminthurus bilineatus Tuxen, 1944 :
            Syn.: Deuterosminthurus bilineatus Bödvarsson, 1957 :
            Syn.: Bourletiella bilineata Haybach, 1971 :
            Syn.: Bourletiella bilineata Haybach, 1972 :
            Syn.: Bourletiella craggi Murphy, 1960


Bourletiella (Heterosminthurus) clavigera
from Süd-Bayern
After Gisin H, 1960 Abb.521
Heterosminthurus claviger from the UK
2008.07.10 © Ardron, P.A. (Det.)
Heterosminthurus claviger ♀ from the UK
2008.07.10 © Ardron, P.A. (Det.)
Heterosminthurus claviger ♀ from the UK
2008.07.10 © Ardron, P.A. (Det.)
Heterosminthurus claviger ♀ from the UK
2008.07.10 © Ardron, P.A. (Det.)
Heterosminthurus claviger ♂ from the UK
2008.07.10 © Ardron, P.A. (Det.)
Heterosminthurus claviger ♂ from the UK
Interocular paddle-like setae
2008.07.10 © Ardron, P.A. (Det.)
Heterosminthurus claviger ♂ from the UK
Interocular paddle-like setae
2008.07.10 © Ardron, P.A. (Det.)
Heterosminthurus claviger from Holland
2013.08.09 © den Ouden, A.
Heterosminthurus claviger from Holland
2014.10.06 © van Duinen, J.
Heterosminthurus claviger from Holland
2014.10.06 © van Duinen, J.
Heterosminthurus claviger from Holland
2014.10.06 © van Duinen, J.
Heterosminthurus claviger from Holland
2014.10.06 © van Duinen, J.
Heterosminthurus claviger ♀ from Holland
2015.06.01 © van Duinen, J.
Heterosminthurus claviger ♂ from Holland
Interocular paddle-like setae (caudal aspect)
2015.06.01 © van Duinen, J.
Heterosminthurus claviger from Holland
2015.07.26 © van Duinen, J.
Heterosminthurus claviger from the UK
2015.09.10 © Boardman, P.
Heterosminthurus claviger from Holland
2016.09.03 © van Duinen, J.
Heterosminthurus claviger ♀ from Belgium
Relative short antennae
2017.07.18 © Huskens, M.-L.
Heterosminthurus claviger ♀ from Belgium
2017.07.18 © Huskens, M.-L.
Heterosminthurus claviger ♀ from Belgium
2017.07.18 © Huskens, M.-L.
Heterosminthurus claviger ♂ from Belgium
Relative long antennae
2017.08.07 © Huskens, M.-L.
Heterosminthurus claviger ♂ from Belgium
Interocular paddle-like setae
2017.08.07 © Huskens, M.-L.
Heterosminthurus claviger ♂ from Belgium
Interocular paddle-like setae
2017.08.07 © Huskens, M.-L.
Heterosminthurus claviger ♀ & ♂ from Holland
Courtship ritual
2017.09.10 © van Duinen, J.
Heterosminthurus claviger from France
2018.05.10 © Picard, J.
Heterosminthurus claviger from the UK
2018.05.20 © Cann, A.J.
Heterosminthurus claviger from the UK
Antenna
2018.05.20 © Cann, A.J.
Heterosminthurus claviger from Belgium
2018.06.14 © Huskens, M.-L.
Heterosminthurus claviger ♂? from France
Non-reproductive instar
2018.11.22 © Dejoux, P.
Heterosminthurus claviger ♂ from Belgium
With interocular paddele-shaped macrosetae
2019.08.20 © Huskens, M.-L.
Heterosminthurus claviger ♂ from France
2020.09.05 © Picard, J.
Heterosminthurus claviger ♂? from France
Non-reproductive instar
2020.10.13 © Dejoux, P.
Heterosminthurus claviger ♂? from France
Non-reproductive instar
2020.10.13 © Dejoux, P.
Heterosminthurus claviger ♀ from Belgium
2020.10.14 © Huskens, M.-L.
Heterosminthurus claviger ♀ from Belgium
With 5 flank setae f1-f5 + seta i
2020.10.14 © Huskens, M.-L.
Heterosminthurus claviger ♀ from Belgium
2020.10.14 © Huskens, M.-L.
Heterosminthurus claviger ♀ from Belgium
Small abdomen dorsal pattern
2020.10.14 © Huskens, M.-L.
Heterosminthurus claviger ♂ from Belgium
Interocular paddle shaped macrosetae
2020.10.14 © Huskens, M.-L.
Heterosminthurus claviger from Belgium
With 2 parasitic protonymph Acari
2021.06.24 © Huskens, M.-L.
Heterosminthurus claviger ♂ from France
Interocular paddle-like setae
2021.08.02 © Picard, J.
Heterosminthurus claviger ♂ from France
Distinctly long antenna
2021.08.02 © Picard, J.
Heterosminthurus claviger ♂ from France
Interocular paddle-like setae sticking together
2021.08.02 © Picard, J.

Species diffusus (Gisin, H, 1962:23) Bretfeld, 1999

            Syn.: Bourletiella nonlineata Haybach, 1972


Species dreisbachi (Snider, RJ, 1969:360) Christiansen & Bellinger, 1981

Deuterosminthurus dreisbachi from the USA
After Snider RJ, 1969 Fig.1
            Syn.: ... bilineatus Hammer, 1953 =?
            Syn.: Heterosminthurus bilineatus Christiansen & Bellinger, 1981 =?
             Syn.: Bourletiella (Bourletiella) bilineatus(sic) Skidmore, RE, 1995:62 =? ISS
            Syn.: Heterosminthurus bilineatus Christiansen & Bellinger, 1998 =?
            Syn.: Heterosminthurus bilineatus Bretfeld, G, 1999:262



Species ihu Christiansen, K et Bellinger, P, 1992:365

Species insignis (Reuter, OM, 1876:83) Stach, J, 1956:181, t.t., B

            Syn.: Sminthurus novemlineatus var. palustris Becker, 1902
            Syn.: Sminthurus repandus Ågren, 1902
            Syn.: Bourletiella (Deuterosminthurus) aquatica Maynard, EA, 1951:259 =?
            Syn.: Heterosminthurus insignis f. cyaneus Bretfeld, G, 1988:2
             Syn.: Deuterosminthurus novemlineatus var. pilosicauda(sic) Linnaniemi, WM, 1912:307 partim ISS
            Syn.: Heterosminthurus novemlineatus Cassagnau, 1961 :
            Syn.: Heterosminthurus novemlineatus Cassagnau, 1964 :
            Syn.: Heterosminthurus insignis f. maculatus Bretfeld, G, 1988:2
Heterosminthurus insignis f. maculatus ♀ from France
2017.07.01 © Tiky, S.
Heterosminthurus insignis f. maculatus ♀ & ♂ from France
Male stimulates genital region of female
2017.07.01 © Tiky, S.
Heterosminthurus insignis f. maculatus ♂ from France
Caudal aspect
2017.07.01 © Tiky, S.
Heterosminthurus insignis f. maculatus ♀ from France
2017.07.01 © Tiky, S.
Heterosminthurus insignis f. maculatus ♀ from France
2017.07.01 © Tiky, S.
Heterosminthurus insignis f. maculatus from France
Foraging
2017.07.07 © Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
2017.07.07 © Picard, J.
Heterosminthurus insignis f. maculatus ♂ from France
2017.07.07 © Picard, J.
Heterosminthurus insignis f. maculatus ♂ from France
2017.07.07 © Picard, J.
Heterosminthurus insignis f. maculatus ♂ from France
2017.08.15 © Picard, J.
Heterosminthurus insignis f. maculatus ♂ from France
2017.08.15 © Picard, J.
Heterosminthurus insignis f. maculatus ♂ from France
Long spiky interocular macrosetae (*)
2017.08.15 © Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
2017.08.15 © Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
2017.08.15 © Picard, J.
Heterosminthurus insignis f. maculatus ♂ from France
Spiky interocular macrosetae (*)
2017.08.15 © Picard, J.
Heterosminthurus insignis f. maculatus ♂ from France
With long antennae
2017.08.15 © Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
Ventral aspect
2017.08.15 © Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
Caudal aspect
2017.08.16 © Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
Lateral aspect
2017.09.09 © Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
Dorsal aspect
2017.09.09 © Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
Caudal aspect
2017.09.09 © Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
2017.09.10 © Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
2017.09.10 © Picard, J.
Heterosminthurus insignis f. maculatus ♂ from France
Spiky interocular macrosetae (*)
2017.09.10 © Picard, J.
Heterosminthurus insignis f. maculatus ♂ from France
2017.09.16 © Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
2017.09.16 © Picard, J.
Heterosminthurus insignis f. maculatus ♂ & ♀ from France
Dark male & pale female
2017.09.21 © Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
Lateral anterior pattern on great abdomen
2017.09.21 © Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
Lateral anterior pattern on great abdomen
2017.09.21 © Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
Facial chaetotaxy
2017.09.30 © Picard, J.
Heterosminthurus insignis f. maculatus ♂ from France
Subadult with undeveloped genital papilla
2017.09.30 © Picard, J.
Heterosminthurus insignis f. maculatus ♂ from France
2017.10.22 © Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
Dental macrosetae as long as mucro
2017.11.22 © Janssens F. Magn. 100x
Leg.: Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
Sub-anal appendage curved at basis
2017.11.22 © Janssens F. Magn. 600x
Leg.: Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
Unguiculus of foot3 with large lamella (*)
2017.11.23 © Janssens F. Magn. 600x
Leg.: Picard, J.
Heterosminthurus insignis f. maculatus ♀ from France
Mucro spoonshaped with transparant flexible smooth edges
2017.11.24 © Janssens F. Magn. 600x
Leg.: Picard, J.
Heterosminthurus insignis f. maculatus ♂ from France
Subadult
2017.11.25 © Picard, J.
Heterosminthurus insignis f. maculatus ♂ from France
Subadult
2017.11.25 © Picard, J.
Heterosminthurus insignis f. maculatus ♂ from France
2018.04.07 © Picard, J.
Heterosminthurus insignis f. maculatus ♀ & ♂ from France
Courtship
2018.10.02 © Picard, J.
Heterosminthurus insignis f. maculatus ♂ from France
With 4 long interocular macrosetae (*)
2018.12.15 © Picard, J.

            Syn.: Heterosminthurus insignis f. principalis Bretfeld, G, 1988:1
            Syn.: Heterosminthurus insignis f. marmoratus Janssens F, Picard J & Tiky S, 2018
            Syn.: Heterosminthurus sp.1 f. marmoratus Janssens F, Picard J & Tiky S, 2017
Heterosminthurus sp.1 f. marmoratus ♀ from France
Marmbled colour form
2017.08.15 © Picard, J.
Heterosminthurus sp.1 f. marmoratus ♂ from France
2017.09.10 © Picard, J.
Heterosminthurus sp.1 f. marmoratus from France
Caudal aspect
2017.09.10 © Picard, J.
Heterosminthurus sp.1 f. marmoratus ♂ from France
2017.09.13 © Picard, J.
Heterosminthurus sp.1 f. marmoratus from France
2017.09.23 © Picard, J.
Heterosminthurus sp.1 f. marmoratus from France
2018.04.07 © Picard, J.
Heterosminthurus insignis f. marmoratus ♂ from France
Lateral aspect
2018.07.07 © Picard, J.
Heterosminthurus insignis f. marmoratus ♂ from France
Long interocular macrosetae (*)
2018.07.07 © Picard, J.
Heterosminthurus insignis f. marmoratus ♂ from France
Facial aspect
2018.07.07 © Picard, J.
Heterosminthurus insignis f. marmoratus ♂ from France
Caudal aspect
2018.07.07 © Picard, J.
Heterosminthurus insignis f. marmoratus ♂ from France
Four long interocular macrosetae
2018.07.17 © Picard, J.
Heterosminthurus insignis f. marmoratus from France
Pale form
2018.11.17 © Picard, J.
Heterosminthurus insignis f. marmoratus from France
Pale form
2018.11.17 © Picard, J.





Heterosminthurus insignis from Poland, ♂
Male with 4 distincly longer interocular setae
After Stach, J, 1956 Pl.XXV Fig.1
Heterosminthurus insignis from Germany
Courtship ritual
After Bretfeld G, 1976
Heterosminthurus insignis from the UK, ♂
Some interocular setae are longer than the others
2005 © Hopkin, S.P.
Heterosminthurus insignis from France
2007.06.18 © Lebeaux, P.
Heterosminthurus insignis from France
2007.06.18 © Lebeaux, P.
Heterosminthurus insignis from the UK, ♀
Det. Ardon, P.A.
2008.06.21 © Ardron, P.A.
Heterosminthurus insignis from the UK
Det. Ardon, P.A.
2008.06.21 © Ardron, P.A.
Heterosminthurus insignis from the UK
Det. Ardon, P.A.
2008.06.21 © Ardron, P.A.
Heterosminthurus insignis from the UK
Male chasing female
2008.06.30 © Ardron, P.A. (Det.)
Heterosminthurus insignis from the UK
Male rejected by female
2008.06.30 © Ardron, P.A. (Det.)
Heterosminthurus insignis ♀ from the UK
2008.06.30 © Ardron, P.A. (Det.)
Heterosminthurus insignis from the UK
2008.06.30 © Ardron, P.A. (Det.)
Heterosminthurus insignis from the UK
2008.08.06 © Robertson, A.
Heterosminthurus insignis from France
2009.09.dd © Lebeaux, P.
Heterosminthurus insignis from France
2009.09.dd © Lebeaux, P.

             Syn.: Heterosminthurus insigris(sic) Nickerl, J, Helbig, R, Schulz, H-J, Werner, C et Neinhuis, C, 2012:7 ISS
Heterosminthurus insignis from the UK
Glossy cephalic cuticula
Dark ant.3+4
2012.05.09 © Murray, A.
Heterosminthurus insignis from France
2012.07.26 © Lebeaux, P.
Heterosminthurus insignis juv. from France
2012.07.31 © Lebeaux, P.
Heterosminthurus insignis ♀ & ♂ from France
Courtship ritual
2012.07.31 © Lebeaux, P.
Heterosminthurus insignis from Holland
Glossy cephalic cuticula
2013.08.22 © van Duinen, J.
Heterosminthurus insignis from the UK
2014.06.13 © Michaelson, J.
Heterosminthurus insignis ♀ from Holland
2016.08.28 © van Duinen, J.
Heterosminthurus insignis ♂ from Holland
2016.08.28 © van Duinen, J.
Heterosminthurus insignis ♀ from Sweden
Glossy cephalic cuticula
Dark ant.3+4
2022.09.26 © Kamsteeg, G.
Heterosminthurus insignis ♀ from Holland
Eyebrow distinctly indented
2023.06.30 © Kamsteeg, G.
Heterosminthurus insignis ♂ from Holland
Distinct interocular macrosetae
2023.07.14 © Kamsteeg, G.
Heterosminthurus insignis ♂ from Holland
Glossy cephalic cuticula
2023.07.14 © Kamsteeg, G.
Heterosminthurus insignis from Holland
2 ocular setae
2023.07.14 © Kamsteeg, G.
Heterosminthurus insignis from Holland
Whitish midstripe
2023.07.14 © Kamsteeg, G.
Heterosminthurus insignis from Holland
Abdominal dorsal reticulation
Enhanced contrast
2023.07.14 © Kamsteeg, G.
Heterosminthurus insignis from Holland
Ant.4 articulations
2023.07.14 © Kamsteeg, G.



Species kiianus Yoshii & Sawada, 1997

Species koontzi Snider, RJ et Calandrino, FJ, 1987:10

Bourletiella (Heterosminthurus) koontzi from the USA
After Snider RJ & Calandrino FJ, 1987 Fig.14
Bourletiella (Heterosminthurus) koontzi from the USA
After Snider RJ & Calandrino FJ, 1987 Fig.15

Species linnaniemii (Stach, 1920) Stach, 1955

            Syn.: Sminthurus nigriceps Schille, 1908
            Syn.: Deuterosminthurus linnaniemii var. decolorata Stach, 1919
Heterosminthurus linnanieimii var. decolorata from Poland
After Stach J, 1956 PlateXXIII Fig.4

            Syn.: Heterosminthurus linnaniemii f. principalis Stach, J, 1956:189
            Syn.: Heterosminthurus linnaniemii f. rubra Bretfeld, 2000



Heterosminthurus linnanieimii ♀ from Poland
After Stach J, 1956 PlateXXIII Fig.1
Heterosminthurus linnanieimii ♂ from Poland
After Stach J, 1956 PlateXXIII Fig.3




             Syn.: Deuterosminthurus linnaniemi(sic) Szeptycki & Weiner in Razowski, 1990 ISS

Species multiornatus Bretfeld, G et Zöllner, E, 2000:138

Heterosminthurus multiornatus ♂ from Russia
After Bretfeld G & Zöllner E, 2000 Fig.3
Heterosminthurus multiornatus ♂ from Russia
Head frontodorsal
After Bretfeld G & Zöllner E, 2000 Fig.5

Species nonlineatus (Gisin, 1946)

            Syn.: Heterosminthurus cornutus ssp. caucasicus Stach, J, 1956:192


Species novemlineatus (Tullberg, 1871) Stach, J, 1956:195

            Syn.: Sminthurus novem-lineatus(sic) Tullberg, 1871 IOS
            Syn.: Sminthurus pilosicauda(sic) Reuter, OM, 1876:84 IOS
            Syn.: Sminthurus pilosicauda(sic) Reuter, OM, 1892:232 IOS
             Syn.: Sminthurus (Deuterosminthurus) oblongus ssp. major var. pilosicauda(sic) Börner, C, 1901:107 ISS
             Syn.: Deuterosminthurus novemlineatus var. pilosicauda(sic) Linnaniemi, WM, 1912:307 partim ISS
             Syn.: Deuterosminthurus novemlineatus f. pilosicauda(sic) Handschin, E in Dahl, F, 1929:117 ISS
             Syn.: Deuterosminthurus novemlineatus var. pilosicauda(sic) Kloet, GS et Hincks, WD, 1945:9 ISS
             Syn.: Heterosminthurus novemlineatus var. pilosicauda(sic) Stach, J, 1956:195 ISS
Heterosminthurus novemlineatus from Poland
Varietas pilosicauda(sic)
After Stach, J, 1956 Pl.XXIV Fig.5

             Syn.: Heterosminthurus novemlineatus f. pilosicauda(sic) Stach, J, 1956:197 ISS
            Syn.: Sminthurus novemlineatus var. cyaneofuscus Becker, 1902
             Syn.: Heterosminthurus novemlineatus var cyanofusca(sic) Bretfeld, G, 1999:270 ISS
            Syn.: Heterosminthurus novemlineatus f. bifasciata Stach, J, 1956:196
Heterosminthurus novemlineatus from Poland
Forma bifasciata
After Stach, J, 1956 Pl.XXIV Fig.2

            Syn.: Heterosminthurus novemlineatus f. malopigmentata Stach, J, 1956:196
Heterosminthurus novemlineatus from Poland
Forma malopigmentata
After Stach, J, 1956 Pl.XXIV Fig.1

            Syn.: Heterosminthurus novemlineatus f. unilineata Stach, J, 1956:195
Heterosminthurus novemlineatus f. unilineata from Poland
After Stach J, 1956 Pl.XXIV Fig.4




             Syn.: Sminthurus (Setosi) novem-lineatus(sic) Tullberg, T, 1872:30 ISS
             Syn.: Sminthurus novemlineato(sic) Reuter, OM, 1876:84 ISS
             Syn.: Sminthurus 9-lineatus(sic) f. principalis Börner, 1900 ISS
Heterosminthurus novemlineatus from Poland
After Stach, J, 1956 Pl.XXIV Fig.6
Heterosminthurus novemlineatus from Poland
After Stach, J, 1956 Pl.XXIV Fig.3
Heterosminthurus novemlineatus from Poland
2015.05.13 © Beck, K.
Heterosminthurus novemlineatus from Poland
2015.05.13 © Beck, K.
Heterosminthurus novemlineatus from Poland
2015.05.13 © Beck, K.
Heterosminthurus novemlineatus from Poland
Caudal aspect
With faecal pellet
2015.05.13 © Beck, K.
Heterosminthurus novemlineatus from Holland
2017.05.17 © van Duinen, J.
Heterosminthurus novemlineatus from Holland
2017.05.17 © van Duinen, J.
Heterosminthurus novemlineatus juv. from Holland
2017.05.17 © van Duinen, J.
Heterosminthurus novemlineatus from Belgium
2020.09.08 © Smets, W.
Heterosminthurus novemlineatus from Belgium
2020.09.08 © Smets, W.





Species nymphes Yosii, R, 1970:23

Species pirika Yosii, R, 1972:96

Species punctatus Bretfeld, 1996

Species putoranae Bretfeld, 2000

Species quadristrigatus Bretfeld, 2000

Species setiger Gisin, 1961

Species stebaevae Bretfeld, 1996

Species umbonicus Bretfeld, 2000

Species undulans Yosii, R et Lee, C-E, 1963:11

Heterosminthurus undulans from Korea
After Yosii R & Lee C-E, 1963 Fig.6A

            Syn.: Heterosminthurus undulatus Yosii, R et Lee, C-E, 1963:12 =?
            Syn.: Heterosminthurus insignis Yosii, 1954 =?
            Syn.: Heterosminthurus insignis Yosii, R, 1977:164 =?



Heterosminthurus undulans from Japan
2023.01.18 © Cheng, H.-J.
Leg.: Nakamori, T.
Heterosminthurus undulans from Japan
2023.01.18 © Cheng, H.-J.
Leg.: Nakamori, T.
Heterosminthurus undulans from Japan
2023.01.18 © Cheng, H.-J.
Leg.: Nakamori, T.
Heterosminthurus undulans from Japan
2023.01.18 © Cheng, H.-J.
Leg.: Nakamori, T.
Heterosminthurus undulans from Japan
Ant.4 subdivided
2023.01.18 © Cheng, H.-J.
Leg.: Nakamori, T.
Heterosminthurus undulans from Japan
Foot2complex with 2 clavate setae
Unguiculus with 2 lamellae
2023.01.18 © Cheng, H.-J. Leg.: Nakamori, T.
Heterosminthurus undulans ♀ from Japan
Sub-anal appendage
2023.02.07 © Cheng, H.-J. Leg.: Nakamori, T.
Heterosminthurus undulans from Japan
Foot1complex
Unguiculus filamenteous
2023.02.08 © Cheng, H.-J. Leg.: Nakamori, T.
Heterosminthurus undulans from Japan
Foot3complex
Unguiculus with distinctly large lamella
2023.02.08 © Cheng, H.-J. Leg.: Nakamori, T.
Heterosminthurus undulans from Japan
Mucro edge with undulations
2023.02.10 © Cheng, H.-J. Leg.: Nakamori, T.

Species undulatus (Hammer, 1953) Stach, 1956:180


Genus Aneuempodialis Stach, 1955

Aneuempodialis? sp. from Singapore
2022.11.14 © Bay, N.
Aneuempodialis? sp. from Singapore
2022.11.14 © Bay, N.
Aneuempodialis? sp. from Singapore
Eye with smaller ocelli C + D
2022.11.14 © Bay, N.
Aneuempodialis? sp. from Singapore
Dorsal macrosetae distinctly spiky
2022.11.14 © Bay, N.

Species cinereus (Womersley, H, 1932:31) Stach, 1955 tt

Deuterosminthurus cinereus from Australia
After Womersley H, 1932 Fig.10d
            Syn.: Deuterosminthurus (Deuterosminthurus) cinereus var. quadripunctatus Womersley, H, 1932:31
            Syn.: Deuterosminthurus (Deuterosminthurus) cinereus f. principalis Womersley, H, 1939:249



Aneuempodialis cinereus from Australia
2016.07.10 © Steve & Alison.
Aneuempodialis cinereus from Australia
2016.07.10 © Steve & Alison.
Aneuempodialis cinereus from Australia
2016.07.10 © Steve & Alison.
Aneuempodialis cinereus from Australia
2016.08.07 © Steve & Alison.

Species maculatus (Womersley, H, 1939:322) Greenslade, P, 1977:249


Genus Bovicornia Delamare Deboutteville, 1947

  Syn.: Bourletiellinae Bovicordia(sic) Massoud, Z et Betsch, J-M, 1972:90 ISS
Bovicornia sp. from Botswana
2015.01.06 © Dabek, L.



Species bidoma Mari Mutt, JA, 1978:166

Species coronata Delamare Deboutteville, 1947, t.t.

Species greensladei Massoud & Delamare Deboutteville, 1967

Species pauliani Massoud & Delamare Deboutteville, 1965

             Syn.: Bovicornia paulani(sic) Mari Mutt, JA, 1978:165 ISS

Species rhinoceros Massoud & Delamare Deboutteville, 1964

Species somala Dallai, R, 1981:242

Species uniformis Murphy, 1960


Genus Prorastriopes Delamare Deboutteville, 1947

  Syn.: Bourletiellidae Andiella Stach, 1955
  Syn.: Bourletiellidae Andiella Betsch, 1977
  Syn.: Bourletiellidae Andiella Bretfeld, 1992
  Syn.: Bourletiellidae Andiella Nayrolles, 1993
  Syn.: Bourletiellidae Andiella Nayrolles, P, 1997:372


Prorastriopes sp. from the USA
2021.03.27 © Ostrowski, M.

Species sp.1 Janssens F sensu Christiansen & Bellinger, 1998

            Syn.: Bourletiella (Prorastriopes) sp.F Christiansen, K et Bellinger, P, 1998:1298


Species sp.2 Janssens F sensu Christiansen & Bellinger, 1998

            Syn.: Bourletiella (Prorastriopes) sp.G Christiansen, K et Bellinger, P, 1998:1298


Species sp.3 Janssens F sensu Christiansen & Bellinger, 1998

            Syn.: Bourletiella (Prorastriopes) sp.H Christiansen, K et Bellinger, P, 1998:1298


Species sp.4 Janssens F sensu Christiansen & Bellinger, 1998

            Syn.: Bourletiella (Prorastriopes) sp.I Christiansen, K et Bellinger, P, 1998:1298


Species sp.5 Janssens F sensu Christiansen & Bellinger, 1998

            Syn.: Bourletiella (Prorastriopes) sp.J Christiansen, K et Bellinger, P, 1998:1298


Species sp.6 Janssens F sensu Christiansen & Bellinger, 1998

            Syn.: Bourletiella (Prorastriopes) sp.K Christiansen, K et Bellinger, P, 1998:1298


Species sp.7 Janssens F sensu Christiansen & Bellinger, 1998

            Syn.: Bourletiella (Prorastriopes) sp.L Christiansen, K et Bellinger, P, 1998:1298


Species sp.8 Janssens F sensu Christiansen & Bellinger, 1998

            Syn.: Bourletiella (Prorastriopes) sp.M Christiansen, K et Bellinger, P, 1998:1298


Species sp.9 Janssens F sensu Bellini & Zeppelini, 2004

            Syn.: Prorastriopes sp.undescribed Bellini, BC et Zeppelini, D, 2004:588


Species sp.10 Janssens F & Takahashi M, 2009

Prorastriopes sp. from the USA
2009.08.25 © Takahashi, M.
Prorastriopes sp. from the USA
Fourth antennal segment with 12 subsegments
2009.08.27 © Takahashi, M.
Prorastriopes sp. from the USA
2009.08.27 © Takahashi, M.
Prorastriopes sp. from the USA
2009.08.27 © Takahashi, M.

Species sp.11 Janssens F sensu Bellini & Zeppelini, 2004

            Syn.: Prorastriopes sp.n. Zeppelini, D, Bellini, BC, Creao-Duarte, AJ et Hernández, MIM, 2009:1166


Species sp.12 Janssens F & Takahashi M, 2009

Prorastriopes sp. from the USA
2009.08.25 © Takahashi, M.
Prorastriopes sp. from the USA
Fourth antennal segment with 12 subsegments
2009.08.27 © Takahashi, M.
Prorastriopes sp. from the USA
2009.08.27 © Takahashi, M.
Prorastriopes sp. from the USA
2009.08.27 © Takahashi, M.
Prorastriopes sp. from the USA
2020.07.10 © Cam, C.
Prorastriopes sp. from the USA
2020.07.10 © Cam, C.
Prorastriopes sp. from the USA
2020.07.10 © Cam, C.
Prorastriopes sp. from the USA
2020.07.11 © Cam, C.
Prorastriopes sp. from the USA
2020.07.11 © Cam, C.
Prorastriopes sp. from the USA
Ant.4 with 12 subdivisions
2020.07.11 © Cam, C.
Prorastriopes sp. from the USA
Ant.4 with 12 subdivisions
2020.07.11 © Cam, C.
Prorastriopes sp. from the USA
2020.08.12 © Blazeclaw.
Prorastriopes sp. from the USA
2020.08.12 © Blazeclaw.

Species sp.13 Janssens F, Miller T & Barbin T, 2022

Prorastriopes sp. from Canada
2021.05.21 © Miller, T.
Prorastriopes sp. from Canada
2021.05.21 © Miller, T.
Prorastriopes sp. from Canada
2022.05.23 © Barbin, T.
Prorastriopes sp. from Canada
2022.05.23 © Barbin, T.
Prorastriopes sp. from Canada
Ant.4 with 12 subdivisions
2022.05.23 © Barbin, T.
Prorastriopes sp. from Canada
Abdominal bothriotricha ABC in straight line
2022.05.23 © Barbin, T.
Prorastriopes sp. from Canada
Size of ocellus C = ocellus H
2022.05.23 © Barbin, T.

Species sp.14 Janssens F & Hays T, 2022

Prorastriopes sp. from the USA
2022.09.10 © Hays, T.
Prorastriopes sp. from the USA
2022.09.10 © Hays, T.
Prorastriopes sp. from the USA
2022.09.10 © Hays, T.
Prorastriopes sp. from the USA
2022.09.10 © Hays, T.

Species alatipes Murphy, 1960

Species azuladus Bretfeld, 1992

Species barnardi (Womersley, 1931)

Species carrerensis (Rapoport & Bianco, 1962) Betsch, 1980

Species cingulus (Bonet, F, 1933:188) Betsch, J-M, 1980:178 partim

            Syn.: Deuterosminthurus cingula(sic) Bonet, F, 1933:188 IOS
Deuterosminthurus cingula(sic) ♀ from Argentina
After Bonet F, 1933 Fig.XXIII.1
Deuterosminthurus cingula(sic) ♀ from Argentina
Abd.5+6
Backwards directed sub-anal appendages (<-)
After Bonet F, 1933 Fig.XXII.5
Deuterosminthurus cingula(sic) ♂ from Argentina
After Bonet F, 1933 Fig.XXIII.2
Deuterosminthurus cingula(sic) ♂ from Argentina
Abd.5+6
Genital papilla (*)

After Bonet F, 1933 Fig.XXII.4

            Syn.: ... procingulus Delamare Deboutteville & Massoud, 1963



             Syn.: Deuterosminthurus cingula(sic) Stach, 1955 ISS
             Syn.: Deuterosminthurus cingula(sic) Stach, J, 1956:163 ISS
             Syn.: Prorastriopes cingula(sic) Mari Mutt, JA et Bellinger, PF, 1990:151 ISS
Prorastriopes cingulus ♀ from Argentina
After Heckman CW, 2001 Fig.1.568 left
Prorastriopes cingulus ♂ from Argentina
Type 1
After Heckman CW, 2001 Fig.1.568 midleft
Prorastriopes cingulus ♂ from Argentina
Type 2
After Heckman CW, 2001 Fig.1.568 midright
Prorastriopes cingulus ♂ from Peru
Type 1
2009.03.23 © Contreras Valdez, A.
Prorastriopes cingulus ♀ from Argentina
2009.06.11 © Gonzalez-Carducci, E.
Prorastriopes cingulus ♂ from Argentina
Type 2
2012.02.20 © Gonzalez-Carducci, E.
Prorastriopes cingulus ♂ from Argentina
Type 2
2014.10.17 © Gonzalez-Carducci, E.
Prorastriopes cingulus ♂ from Argentina
Type 2
2014.10.17 © Gonzalez-Carducci, E.
Prorastriopes cingulus ♂ from Argentina
Type 2
2017.12.23 © Gonzalez-Carducci, E.





Species coalingaensis Snider, 1978

            Syn.: Deuterosminthurus cingulus Scott & Yosii, 1972
             Syn.: Deuterosminthurus cingula(sic) Christiansen, K et Bellinger, P, 1981:1147 ISS
             Syn.: Deuterosminthurus cingula(sic) Christiansen, K et Bellinger, P, 1998:1307 ISS


             Syn.: Prorastriopes coalingensis(sic) Snider, RJ et Loring, SJ, 1984:37 ISS
             Syn.: Bourletiella (Prorastriopes) coalingensis(sic) Christiansen, K et Bellinger, P, 1998:1323 ISS
Prorastriopes ?coalingaensis? ♀ from the USA
2006 © Gross, J.
Prorastriopes coalingaensis ♀ from the USA
2008.08.20 © Montgomery, G.
Prorastriopes coalingaensis ♂ from the USA
2009.06.25 © Melder, R.
Prorastriopes coalingaensis ♀ from the USA
2010.08.16 © Zungri, D.
Prorastriopes coalingaensis ♂ from the USA
2010.08.18 © Zungri, D.
Prorastriopes coalingaensis ♀ from the USA
2010.08.21 © Zungri, D.
Prorastriopes coalingaensis ♂ from the USA
2010.08.21 © Zungri, D.
Prorastriopes coalingaensis ♀ from the USA
2011.06.18 © Schneider, J.
Prorastriopes coalingaensis ♀ from the USA
Ant.4 with 10 subdivisions
2011.06.18 © Schneider, J.
Prorastriopes coalingaensis ♀ from the USA
2013.06.01 © Heo, C.-C.
Prorastriopes coalingaensis ♂ from the USA
2013.06.01 © Heo, C.-C.
Prorastriopes coalingaensis juv. from the USA
2014.05.29 © Dankowicz, E.
Prorastriopes coalingaensis ♂ from the USA
2014.08.24 © Dankowicz, E.
Prorastriopes coalingaensis ♂ from the USA
2015.08.21 © Rosenfeld, J.
Prorastriopes coalingaensis ♂ from the USA
2016.08.07 © Bailey, J.
Prorastriopes coalingaensis ♂ from the USA
2016.08.07 © Bailey, J.
Prorastriopes coalingaensis ♂ from the USA
2018.06.17 © Dobbs, Br.
Prorastriopes coalingaensis ♂ from the USA
2018.06.17 © Dobbs, Br.
Prorastriopes coalingaensis ♂ from the USA
2018.06.17 © Dobbs, Br.
Prorastriopes coalingaensis ♀ from the USA
2018.06.17 © Dobbs, Br.
Prorastriopes coalingaensis ♀ from the USA
2018.06.17 © Dobbs, Br.
Prorastriopes coalingaensis ♀ from the USA
2018.06.17 © Dobbs, Br.
Prorastriopes ?coalingaensis ♂? from the USA
2019.04.11 © Jimenez, A.
Prorastriopes ?coalingaensis ♂? from the USA
2019.04.11 © Jimenez, A.
Prorastriopes ?coalingaensis ♂? from the USA
2019.04.11 © Jimenez, A.
Prorastriopes coalingaensis ♂ from the USA
2019.07.18 © Solomon.
Prorastriopes coalingaensis ♀ from the USA
2020.07.19 © Fandre, T.
Prorastriopes coalingaensis ♀ from the USA
2020.07.19 © Fandre, T.
Prorastriopes coalingaensis ♀ from the USA
2023.04.23 © Gruver, Be.
Prorastriopes coalingaensis ♀ from the USA
2023.04.23 © Gruver, Be.
Prorastriopes coalingaensis ♀ from the USA
2023.04.23 © Gruver, Be.
Prorastriopes coalingaensis ♂ from the USA
Drinking : collophore vesicles pressed against moist substrate
2023.07.06 © Gruver, Be.
Prorastriopes coalingaensis ♀ from the USA
2023.07.06 © Gruver, Be.



Species domeykoi (Stach, 1955) Betsch, 1977

Species izarrae (Najt, 1967) Betsch, 1980

            Syn.: Andiella izarrai(sic) de Izarra, DC, 1965:275 nn IOS




Species lippsoni Snider, 1978

Species marginatus Bretfeld, 1992

Species marmoratus (Womersley, 1931)

Species olivieri Delamare Deboutteville & Massoud, 1963:267

            Syn.: Bourletiella Olivieri(sic) Delamare Deboutteville & Massoud, 1963 IOS

Species pampeanus (Rapoport & Bianco, 1962) Betsch, 1980

Species paryskii (Stach, 1955) Betsch, 1977

Species patagonicus (Delamare Deboutteville & Massoud, 1963) Betsch, 1980

Species pauliani Delamare Deboutteville, C et Massoud, Z, 1965:70

Prorastriopes pauliani from Angola
After Delamare Deboutteville C & Massoud Z, 1965 Fig.2E
            Syn.: Prorastriopes pauliani Delamare Deboutteville, C et Massoud, Z, 1964:70 u


Species pulcher Delamare Deboutteville, 1947, t.t.

            Syn.: Prorastriopes pulchra(sic) Delamare Deboutteville, 1947,tt IOS
             Syn.: Prorastriopes pulchrus(sic) Salmon, JT, 1964:610 ISS
            Syn.: Prorastriopes pulchra(sic) Thibaud, J-M, 2013:236 IOS

  Subspecies punctatus Delamare Deboutteville, 1947





Species risbeci Delamare Deboutteville, 1947

Species savannus Murphy, 1960

Species schultzei (Börner, C in Schultze, 1908:61)

Species spathaceus (Börner, 1907)

Species validentatus (Snider, 1978)

            Syn.: Bourletiella hoffi Scott, 1965 =?
            Syn.: Bourletiella multimaculata Scott, 1965 =?


Species venezuelae Bretfeld, 1992

Species violaceus Murphy, 1960

Species webbi Paclt, 1964

Species wexfordensis (Snider, RJ, 1969:362) Magaña-Martínez, CS, Palacios-Vargas, JG & Navarrete-Heredia, JL, 2019:62

Deuterosminthurus wexfordensis from the USA
After Snider RJ, 1969 Fig.23
Prorastriopes wexfordensis from the USA
2020.02.02 © Kneidel, K.
Prorastriopes wexfordensis from the USA
2020.02.02 © Kneidel, K.

Species xeromorphus Snider, 1978


Genus Bourletides Betsch, J-M et Massoud, Z, 1972:234

Species wallacei Betsch, J-M et Massoud, Z, 1972:225, t.t.


Genus Cassagnaudiella Ellis, WN, 1975:78 key

Species caeruleovernalis Nayrolles, P, 1995:48

            Syn.: Cassagnaudiella cæruleovernalis(sic) Nayrolles, P, 1995:48 IOS


             Syn.: Bourletiella (Cassagnaudiella) coeruleovernalis(sic) Schulz, H-J et van Harten, A, 2013:13 ISS
Bourletiella (Cassagnaudiella) coeruleovernalis(sic) from the UAE
After Schulz H-J & van Harten A, 2013 Pl.2





Species cadiensis Nayrolles, P, 1995:51

Cassagnaudiella cadiensis from Spain
After Nayrolles P, 1995 Fig.15

Species canigouensis Nayrolles, P, 1995:56

Bourletiella (Cassagnaudiella) cf. canigouensis from the UAE
After Schulz H-J & van Harten A, 2013 Pl.1
Cassagnaudiella canigouensis from France
2018.05.20 © Picard, J.
Cassagnaudiella canigouensis ♂ from France
Male with special butt
2018.05.20 © Picard, J.
Cassagnaudiella canigouensis from France
2018.05.20 © Picard, J.
Cassagnaudiella canigouensis from France
Ant.4 with constriction (*)
2018.05.20 © Picard, J.
Cassagnaudiella canigouensis ♀ from France
Caudal aspect
2018.05.20 © Picard, J.
Cassagnaudiella canigouensis ♂ from France
Caudal aspect
2018.05.20 © Picard, J.
Cassagnaudiella canigouensis ♂ from France
2018.05.20 © Picard, J.
Cassagnaudiella canigouensis ♂ from France
2019.06.21 © Picard, J.
Cassagnaudiella canigouensis ♂ from France
Distinctly large ocelli A & B
2019.06.21 © Picard, J.
Cassagnaudiella canigouensis ♂ from France
Distinctly large ocelli B
2019.06.21 © Picard, J.
Cassagnaudiella canigouensis ♀ from France
2019.06.21 © Picard, J.
Cassagnaudiella canigouensis ♀ from France
Abd.6 with 2 white spots
2019.06.21 © Picard, J.
Cassagnaudiella canigouensis ♀ from France
2019.06.21 © Picard, J.
Cassagnaudiella canigouensis ♀ from France
2019.06.21 © Picard, J.

Species cruciata (Haybach, 1972) Nayrolles, 1995

            Syn.: Cassagnaudiella trichroma Nayrolles, P, 1995:56
Cassagnaudiella trichroma from France
After Nayrolles P, 1995 Fig.27
Cassagnaudiella cruciata f. trichroma from France
2021.03.01 © Tiky, S.
Cassagnaudiella cruciata f. trichroma from France
2021.03.01 © Tiky, S.
Cassagnaudiella cruciata f. trichroma from France
2021.03.01 © Tiky, S.
Cassagnaudiella cruciata f. trichroma from France
2021.03.01 © Tiky, S.
Cassagnaudiella cruciata f. trichroma from France
2021.03.01 © Tiky, S.
Cassagnaudiella cruciata f. trichroma from Spain
2021.09.12 © Zeeders, M.
Cassagnaudiella cruciata f. trichroma from Spain
2021.09.12 © Zeeders, M.
Cassagnaudiella cruciata f. trichroma from Spain
2021.09.12 © Zeeders, M.
Cassagnaudiella cruciata f. trichroma ♂ & ♀ from Spain
Courtship
2021.09.15 © Zeeders, M.




Bourletiella (Cassagnaudiella) cruciata from the UAE
After Schulz H-J & van Harten A, 2013 Pl.3
Cassagnaudiella cruciata from France
2019.05.30 © Picard, J.
Cassagnaudiella cruciata from France
Caudal aspect
2019.05.30 © Picard, J.

Species gamae (Bretfeld, 1994)

Bourletiella gamae from Portugal
After Bretfeld, 1994 in Bretfeld, 1999 Fig.477
Cassagnaudiella gamae from France
2017.07.18 © Tiky, S.
Cassagnaudiella gamae from France
2017.07.18 © Tiky, S.
Cassagnaudiella gamae from France
2017.07.18 © Tiky, S.
Cassagnaudiella gamae from France
2017.07.18 © Tiky, S.
Cassagnaudiella gamae from France
Ant.4 with about 10 subdivisions
2017.07.18 © Tiky, S.
Cassagnaudiella gamae from France
2017.08.02 © Tiky, S.
Cassagnaudiella gamae from France
2018.05.10 © Tiky, S.
Cassagnaudiella gamae ♂ from France
2018.05.10 © Tiky, S.
Cassagnaudiella gamae from France
Small abdomen
2018.05.10 © Tiky, S.
Cassagnaudiella gamae from France
2018.06.04 © Picard, J.
Cassagnaudiella gamae from France
2019.06.05 © Tiky, S.

Species lanceolata Bretfeld, G, 2001:95

Species luteovernalis Nayrolles, P, 1995:51

Cassagnaudiella luteovernalis ♀ from France
2019.06.17 © Picard, J.
Cassagnaudiella luteovernalis ♀ from France
2019.06.17 © Picard, J.
Cassagnaudiella luteovernalis ♂ from France
2019.06.17 © Picard, J.
Cassagnaudiella luteovernalis ♂ from France
2019.06.17 © Picard, J.

Species pruinosa (Tullberg, 1871) Boguleanu, 1982

            Syn.: Sminthurus luteo(sic) Reuter, OM, 1876:84
            Syn.: Smynthurus(sic) frontalis Uzel, 1890
            Syn.: Smynthurus fulvus Lie-Petterson, 1896 =?
            Syn.: Bourletiella radula Cassagnau, 1964 partim
            Syn.: Bourletiella radula Raynal, 1973
            Syn.: Bourletiella radula Raynal, G, 1974:182
            Syn.: Bourletiella (Cassagnaudiella) radula Ellis, WN, 1975:78
            Syn.: Bourletiella (Cassagnaudiella) radula Klaver, E, 1975:185
            Syn.: Bourletiella radula Nayrolles, 1995



Cassagnaudiella pruinosa from France
After Nayrolles P, 1995 Fig.16
Lowland colour form
Cassagnaudiella pruinosa from France
After Nayrolles P, 1995 Fig.17
Medium mountain colour form
Cassagnaudiella pruinosa from France
After Nayrolles P, 1995 Fig.18
High mountain colour form
Cassagnaudiella pruinosa ♂ from Holland
2013.09.19 © van Duinen, J.
Cassagnaudiella pruinosa ♂ from Holland
2013.09.19 © van Duinen, J.
Cassagnaudiella pruinosa ♂ from Holland
Abd.5 with modified setae
2013.09.19 © van Duinen, J.
Cassagnaudiella pruinosa from Holland
2018.08.23 © van Leeuwen, J.
Cassagnaudiella ?pruinosa? ♂ from Holland
2019.11.05 © de Haas, M.
Cassagnaudiella ?pruinosa? ♂ from Holland
2019.11.05 © de Haas, M.
Cassagnaudiella ?pruinosa? from Portugal
2021.02.16 © Almeida, J.
Cassagnaudiella ?pruinosa? from Portugal
2021.02.16 © Almeida, J.


Genus Cyprania Bretfeld, 1992

Species gisae Bretfeld, 1992

Species inopinata Bretfeld, 1996


Genus Fasciosminthurus Gisin, H, 1960:281

  Syn.: Sminthuridae Burletiella(sic) Loksa, I, 1978:62 ISS


?Fasciosminthurus? from the USA
2017.08.14 © Hatfield, M.J.
?Fasciosminthurus? from the USA
2021.12.28 © Wildwings.

Species albanicus (Stach, J, 1956:164) Bretfeld, 1992

Deuterosminthurus albanicus from Poland
After Stach J, 1956 Plate XXI Fig.6
Deuterosminthurus albanicus from Poland
After Stach J, 1956 Plate XXI Fig.7

Species angulipunctatus (Loksa, I & Bogojevic,J, 1970:138) Bretfeld, 1992

Bourletiella (Fasciosminthurus) albanica ssp. angulipunctata from Jugoslavia
After Loksa I & Bogojevic J, 1970 Abb.51
Fasciosminthurus ?angulipunctatus? from Hungary
Dark form
2008.06.07 © Keresztes, G.

Species badikahanus Bretfeld, G, 2001:101

Fasciosminthurus badikahanus from the Ivory Coast
Small abdomen
After Bretfeld, 2001 Fig.14

Species bedosae Nayrolles, P, 1994:326

Fasciosminthurus bedosae from Spain
After Nayrolles P, 1994 Fig.21

Species canariensis (Paclt, 1964) Bretfeld, 1992

            Syn.: Fasciosminthurus melanocephalus Bretfeld, 1992 =?


Fasciosminthurus canariensis from the Canary Islands
After Bretfeld G, 1999 Fig.523
Fasciosminthurus canariensis from the Canary Islands
2018.12.06 © Huskens, M.-L.
Fasciosminthurus canariensis from the Canary Islands
2018.12.06 © Huskens, M.-L.
Fasciosminthurus canariensis from the Canary Islands
Caudal aspect
2018.12.06 © Huskens, M.-L.
Fasciosminthurus canariensis from the Canary Islands
Ant.1:2:3:4 = 1:2.1:3.3:7.7
2018.12.06 © Huskens, M.-L.
Fasciosminthurus canariensis from the Canary Islands
2018.12.08 © Huskens, M.-L.
Fasciosminthurus canariensis from the Canary Islands
2018.12.08 © Huskens, M.-L.
Fasciosminthurus canariensis from the Canary Islands
2018.12.08 © Huskens, M.-L.
Fasciosminthurus canariensis from the Canary Islands
2018.12.08 © Huskens, M.-L.
Fasciosminthurus canariensis ♂ from the Canary Islands
2018.12.08 © Huskens, M.-L.
Fasciosminthurus canariensis ♂ from the Canary Islands
Deposition of spermatophore
2018.12.08 © Huskens, M.-L.
Fasciosminthurus canariensis from the Canary Islands
2024.02.09 © Picard, J.
Fasciosminthurus canariensis from the Canary Islands
2024.02.09 © Picard, J.
Fasciosminthurus canariensis from the Canary Islands
2024.02.09 © Picard, J.

Species cassagnaui Nayrolles, P, 1994:330

Fasciosminthurus cassagnaui from Spain
After Nayrolles P, 1994 Fig.30
Fasciosminthurus cassagnaui from Ibiza
2015.09.15 © Beer, R.
Fasciosminthurus cassagnaui from France
2018.05.24 © Tiky, S.
Fasciosminthurus cassagnaui from France
Pollen feeding
2018.05.24 © Tiky, S.
Fasciosminthurus cassagnaui from France
2019.06.06 © Picard, J.
Fasciosminthurus cassagnaui from France
2019.06.06 © Picard, J.
Fasciosminthurus cassagnaui from France
2019.06.06 © Picard, J.
Fasciosminthurus cassagnaui from France
Central ocellus D small
2019.06.06 © Picard, J.

Species circumfasciatus (Stach, J, 1956:176) Martynova, 1964

Deuterosminthurus circumfasciatus from Poland
After Stach J, 1956 Plate XXI Fig.1
Deuterosminthurus circumfasciatus from Poland
After Stach J, 1956 Plate XXI Fig.2

             Syn.: Burletiella(sic) (Fasciosminthurus) cirkumfasciata(sic) Loksa, I, 1978:62 ISS
  Subspecies maculatus Betsch, J-M, 1977:85

Prorastriopes maculatus from Mongolia
After Betsch J-M, 1977 Fig.2D
Fasciosminthurus circumfasciatus ssp. maculatus ♀? from France
2018.05.04 © Picard, J.
Fasciosminthurus circumfasciatus ssp. maculatus ♀? from France
2018.05.04 © Picard, J.
Fasciosminthurus circumfasciatus ssp. maculatus ♀? from France
Dark head
2018.05.04 © Picard, J.
Fasciosminthurus circumfasciatus ssp. maculatus ♀? from France
Caudal aspect
2018.05.04 © Picard, J.
Fasciosminthurus circumfasciatus ssp. maculatus ♀? from France
Distinctly large ocellus B
2018.05.04 © Picard, J.
Fasciosminthurus circumfasciatus ssp. maculatus ♂? from France
Dark broad lateral band
2018.05.04 © Picard, J.
Fasciosminthurus circumfasciatus ssp. maculatus ♂? from France
Ant.1-3 apically dark
2018.05.04 © Picard, J.
Fasciosminthurus circumfasciatus ssp. maculatus ♂? from France
2018.05.04 © Picard, J.
Fasciosminthurus circumfasciatus ssp. maculatus ♀? from France
2018.05.04 © Picard, J.



Species coronatus (Betsch, J-M, 1977:85)

Species cugnyi (Nayrolles, 1987) Nayrolles, P, 1993:671

Species dictyostigmatus Nayrolles, P, 1993:669

Fasciosminthurus dictyostigmatus from France
After Nayrolles P, 1993 Fig.10
Fasciosminthurus dictyostigmatus from France
After Nayrolles P, 1993 Fig.9
Fasciosminthurus dictyostigmatus from Morocco
After Bretfeld, 2001 Fig.19
Fasciosminthurus dictyostigmatus from Spain
2012.04.09 © Joaquim, F.P.
Fasciosminthurus dictyostigmatus from France
2014.05.06 © Lebeaux, P.
Fasciosminthurus dictyostigmatus from France
2014.05.06 © Lebeaux, P.
Fasciosminthurus dictyostigmatus from France
2014.05.06 © Lebeaux, P.
Fasciosminthurus dictyostigmatus from France
2014.05.06 © Lebeaux, P.

Species lacazei (Denis, J-R, 1924:199) Delamare Deboutteville, C & Massoud, Z, 1963:256

            Syn.: Deuterosminthurus Lacazei(sic) Denis, J-R, 1924:199 IOS
             Syn.: Fasciosminthurus Lacazei(sic) Delamare Deboutteville, C et Massoud, Z, 1963:256 ISS
Fasciosminthurus lacazei from Spain
After Nayrolles P, 1993 Fig.1
Fasciosminthurus lacazei from Morocco
After Bretfeld, 2001 Fig.21 left
Fasciosminthurus lacazei from Morocco
After Bretfeld, 2001 Fig.21 right
Fasciosminthurus lacazei from the Canary Islands
Pale form
2019.12.14 © Huskens, M.-L.
Fasciosminthurus lacazei from the Canary Islands
Pale form
2019.12.14 © Huskens, M.-L.
Fasciosminthurus lacazei from the Canary Islands
Facial pattern
2019.12.14 © Huskens, M.-L.
Fasciosminthurus lacazei from the Canary Islands
2019.12.14 © Huskens, M.-L.
Fasciosminthurus lacazei from the Canary Islands
Caudal aspect
2019.12.14 © Huskens, M.-L.
Fasciosminthurus lacazei from the Canary Islands
2019.12.14 © Huskens, M.-L.
Fasciosminthurus lacazei from the Canary Islands
Small abdomen
2019.12.14 © Huskens, M.-L.
Fasciosminthurus lacazei from the Canary Islands
Ocellus D distinctly low in eyepatch
2019.12.14 © Huskens, M.-L.
Fasciosminthurus lacazei from the Canary Islands
Ant.4 with 8 subdivisions
2019.12.14 © Huskens, M.-L.
Fasciosminthurus lacazei ♂ from the Canary Islands
Facial pattern
2020.03.18 © Huskens, M.-L.
Fasciosminthurus lacazei ♂ from the Canary Islands
2020.03.18 © Huskens, M.-L.
Fasciosminthurus lacazei ♂ from the Canary Islands
Ant.4 with 9 subdivisions
2020.03.18 © Huskens, M.-L.
Fasciosminthurus lacazei ♂ from the Canary Islands
Facial pattern
2020.03.18 © Huskens, M.-L.
Fasciosminthurus lacazei ♂ from the Canary Islands
2020.03.18 © Huskens, M.-L.
Fasciosminthurus lacazei ♂ from the Canary Islands
2020.03.18 © Huskens, M.-L.
Fasciosminthurus lacazei from the Canary Islands
2023.03.15 © Huskens, M.-L.
Fasciosminthurus lacazei from the Canary Islands
2023.03.15 © Huskens, M.-L.
Fasciosminthurus lacazei from the Canary Islands
2023.03.15 © Huskens, M.-L.
Fasciosminthurus lacazei from the Canary Islands
Caudal aspect
2023.03.15 © Huskens, M.-L.
Fasciosminthurus lacazei from the Canary Islands
2023.03.15 © Huskens, M.-L.
Fasciosminthurus lacazei from the Canary Islands
Caudal aspect
2023.03.15 © Huskens, M.-L.





Species lacunifer Bretfeld, G, 2001:104

Fasciosminthurus lacunifer from Morocco
After Bretfeld, 2001 Fig.25

Species longisetus Nayrolles, P, 1994:322

Fasciosminthurus longisetus ♀ from Spain
After Nayrolles P, 1994 Fig.7
Fasciosminthurus longisetus ♀ from Spain
After Nayrolles P, 1994 Fig.8

Species mascaraeus Bretfeld, G, 2001:106

Species melanocephalus (Dallai, 1966)

Fasciosminthurus melanocephalus from Italy
After Dallai, 1966 in Bretfeld G, 1999 Fig.523

Species mertensi Bretfeld, G, 2001:106

Fasciosminthurus mertensi from Algeria
After Bretfeld, 2001 Fig.28

Species nairicus Bretfeld, G, 2001:108

Fasciosminthurus nairicus from Algeria
After Bretfeld, 2001 Fig.33 right
Fasciosminthurus nairicus from Algeria
After Bretfeld, 2001 Fig.33 left

Species niger Bretfeld, G, 2001:110

Species obtectus Bretfeld, 1992

Species perforatus Bretfeld, G, 2001:110

Species pontignanoi Bretfeld, 1992

Species pseudovirgulatus Nayrolles, P, 1997:368

Fasciosminthurus pseudovirgulatus from France
After Nayrolles P, 1997 Fig.8
Fasciosminthurus pseudovirgulatus from France
After Nayrolles P, 1997 Fig.9
Fasciosminthurus pseudovirgulatus from France
2012.07.26 © Lebeaux, P.
Fasciosminthurus pseudovirgulatus from France
2017.07.03 © Picard, J.
Fasciosminthurus pseudovirgulatus from France
2017.07.03 © Picard, J.
Fasciosminthurus pseudovirgulatus from France
2017.07.03 © Picard, J.
Fasciosminthurus pseudovirgulatus juv. from France
2017.07.22 © Picard, J.
Fasciosminthurus pseudovirgulatus juv. from France
2017.07.23 © Picard, J.
Fasciosminthurus pseudovirgulatus from France
2017.07.23 © Picard, J.
Fasciosminthurus pseudovirgulatus from France
Feeding instar
2017.07.23 © Picard, J.
Fasciosminthurus pseudovirgulatus from France
2017.07.24 © Picard, J.
Fasciosminthurus pseudovirgulatus from France
2017.08.04 © Picard, J.
Fasciosminthurus pseudovirgulatus from France
2017.08.23 © Tiky, S.
Fasciosminthurus pseudovirgulatus from France
2017.08.23 © Tiky, S.
Fasciosminthurus pseudovirgulatus from France
2017.09.03 © Picard, J.
Fasciosminthurus pseudovirgulatus from France
2018.05.02 © Picard, J.

Species quinquefasciatus (Krausbauer, 1898) Bretfeld, 1970, t.t., B

            Syn.: Sminthurus zebrinus Börner, 1900
            Syn.: Bourletiella zebrina Denis, 1922


             Syn.: Deuterosminthurus cinquefasciatus(sic) Womersley, H, 1931:7 ISS
Deuterosminthurus cinquefasciatus(sic) from Tasmania
After Womersley, 1931 Pl.I Fig.3

             Syn.: Deuterosminthurus 5-fasciatus(sic) Womersley, H, 1931:7 ISS
             Syn.: Deuterosminthurus cinquefasciatus(sic) Womersley, 1932 ISS
             Syn.: Deuterosminthurus cinquefasciatus(sic) Womersley, 1939 ISS
             Syn.: Prorastriopes cinquefasciatus(sic) Greenslade, P et Ireson, JE, 1986:282 ISS
Prorastriopes cinquefasciatus(sic) from Australia
After Greenslade P & Ireson JE, 1986 Fig.1

             Syn.: Prorastriopes cinquefasciatus(sic) Ireson, 1993 ISS
Fasciosminthurus quinquefasciatus from Portugal
2012.05.15 © Jacinto, V.
Fasciosminthurus quinquefasciatus from France
2012.09.16 © Lebeaux, P.
Fasciosminthurus quinquefasciatus from France
2012.09.16 © Lebeaux, P.
Fasciosminthurus quinquefasciatus from France
2012.09.16 © Lebeaux, P.
Fasciosminthurus quinquefasciatus from Holland
2013.08.11 © den Ouden, A.
Fasciosminthurus quinquefasciatus juv. from Holland
After Berg M, 2013 Fig.2 lower
Fasciosminthurus quinquefasciatus juv. from Holland
After Berg M, 2013 Fig.2 upper
Fasciosminthurus quinquefasciatus from Holland
2014.06.24 © van Duinen, J.
Fasciosminthurus quinquefasciatus from Holland
2014.06.24 © van Duinen, J.
Fasciosminthurus quinquefasciatus from Holland
2014.06.24 © van Duinen, J.
Fasciosminthurus quinquefasciatus from Holland
2014.08.14 © Wieringa, P.
Fasciosminthurus quinquefasciatus from Holland
2014.08.14 © van Duinen, J.
Fasciosminthurus quinquefasciatus ♀ from Holland
Sub-anal appendage (s.a.a.)
2014.08.14 © van Duinen, J.
Fasciosminthurus quinquefasciatus from Canada
2015.06.18 © Murray, A.
Fasciosminthurus quinquefasciatus from Canada
2015.06.18 © Murray, A.
Fasciosminthurus quinquefasciatus from Canada
Walking
2015.06.18 © Murray, A.
Fasciosminthurus quinquefasciatus ♀ from the USA
2016.07.27 © Rockenbeck, Bi.
Fasciosminthurus quinquefasciatus ♂ from France
Depositing a spermatophore
2017.05.03 © Tiky, S.
Fasciosminthurus quinquefasciatus from France
2017.06.24 © Tiky, S.
Fasciosminthurus quinquefasciatus from France
Pale colour form
2017.07.14 © Picard, J.
Fasciosminthurus quinquefasciatus ♂ from France
Spermatophore freshly deposited
2017.07.21 © Tiky, S.
Fasciosminthurus quinquefasciatus from France
2017.07.22 © Picard, J.
Fasciosminthurus quinquefasciatus from France
2017.07.22 © Picard, J.
Fasciosminthurus quinquefasciatus ♂ & ♀ from the USA
2017.07.31 © Dobbs, Br.
Fasciosminthurus quinquefasciatus ♂ from the USA
Depositing spermatophore
2017.07.31 © Dobbs, Br.
Fasciosminthurus quinquefasciatus ♂ from the USA
Freshly deposited spermatophore
2017.07.31 © Dobbs, Br.
Fasciosminthurus quinquefasciatus juv. from the USA
2017.08.05 © Dobbs, Br.
Fasciosminthurus quinquefasciatus ♂ from the USA
2017.08.05 © Dobbs, Br.
Fasciosminthurus quinquefasciatus from the USA
Facial markings
2017.08.21 © Dobbs, Br.
Fasciosminthurus quinquefasciatus ♂ & ♀ from the USA
Male meets female
2017.08.21 © Dobbs, Br.
Fasciosminthurus quinquefasciatus ♂ from the USA
2017.08.21 © Dobbs, Br.
Fasciosminthurus quinquefasciatus ♀ from the USA
2017.08.21 © Dobbs, Br.
Fasciosminthurus quinquefasciatus from the USA
Caudal aspect
2017.08.24 © Dobbs, Br.
Fasciosminthurus quinquefasciatus ♀ from the USA
Genital papilla (gp) oval + sub anal appendages (saa)
2017.08.31 © Dobbs, Br.
Fasciosminthurus quinquefasciatus ♂ from the USA
Genital papilla (gp) round
2017.09.03 © Dobbs, Br.
Fasciosminthurus quinquefasciatus ♂ from France
Deposition of spermatophore
2020.04.24 © Reinach, Br.
Fasciosminthurus quinquefasciatus from France
2020.09.22 © Dejoux, P.
Fasciosminthurus quinquefasciatus from the UK
Typical form
After Hutchinson FN & McCulloch JI, 2024 Fig.1a
Fasciosminthurus quinquefasciatus from the UK
Intermediate form
After Hutchinson FN & McCulloch JI, 2024 Fig.1b

  Subspecies canariensis Janssens F & Huskens M-L, 2018

Fasciosminthurus quinquefasciatus ssp. canariensis from the Canary Islands
2018.12.06 © Huskens, M.-L.
Fasciosminthurus quinquefasciatus ssp. canariensis from the Canary Islands
2018.12.06 © Huskens, M.-L.
Fasciosminthurus quinquefasciatus ssp. canariensis from the Canary Islands
2018.12.06 © Huskens, M.-L.
Fasciosminthurus quinquefasciatus ssp. canariensis from the Canary Islands
Caudal aspect
2018.12.06 © Huskens, M.-L.
Fasciosminthurus quinquefasciatus ssp. canariensis from the Canary Islands
2018.12.06 © Huskens, M.-L.
Fasciosminthurus quinquefasciatus ssp. canariensis from the Canary Islands
2018.12.06 © Huskens, M.-L.
Fasciosminthurus quinquefasciatus ssp. canariensis from the Canary Islands
2018.12.08 © Huskens, M.-L.
Fasciosminthurus quinquefasciatus ssp. canariensis from the Canary Islands
2018.12.08 © Huskens, M.-L.
Fasciosminthurus quinquefasciatus ssp. canariensis from the Canary Islands
2024.02.09 © Picard, J.
Fasciosminthurus quinquefasciatus ssp. canariensis from the Canary Islands
2024.02.09 © Picard, J.
Fasciosminthurus quinquefasciatus ssp. canariensis from the Canary Islands
2024.02.09 © Picard, J.
Fasciosminthurus quinquefasciatus ssp. canariensis from the Canary Islands
2024.02.09 © Picard, J.



Species raynalae (Nayrolles, 1987) Nayrolles, P, 1993:659

Species saportae Nayrolles, P, 1997:366

Fasciosminthurus saportae from Spain
After Nayrolles P, 1997 Fig.1
Fasciosminthurus saportae from France
2017.04.22 © Tiky, S.
Fasciosminthurus saportae from France
2017.04.22 © Tiky, S.
Fasciosminthurus saportae from France
2017.04.22 © Tiky, S.
Fasciosminthurus saportae from France
2017.06.28 © Picard, J.
Fasciosminthurus saportae from France
Dorsal anal valve
2017.06.28 © Picard, J.
Fasciosminthurus saportae from France
Antenna
2017.06.28 © Picard, J.
Fasciosminthurus saportae from France
With albino form
2018.05.18 © Tiky, S.
Fasciosminthurus saportae ♂ from France
Albino form
2018.05.18 © Tiky, S.
Fasciosminthurus saportae ♂ from France
Deposition of spermatophore in front of female
2018.05.21 © Picard, J.
Fasciosminthurus saportae ♂ from France
With dark genital papilla (gp)
and pale middorsal Abd.5 (*)
2018.05.21 © Picard, J.
Fasciosminthurus saportae ♀? from France
With pale genital papilla (gp)
and dark middorsal Abd.5 (*)
2018.05.21 © Picard, J.
Fasciosminthurus saportae ♀? from France
Eats spermatophore
2018.05.21 © Picard, J.
Fasciosminthurus saportae from France
2018.05.22 © Picard, J.
Fasciosminthurus saportae from France
2018.05.27 © Picard, J.
Fasciosminthurus saportae ♂ from France
2019.05.25 © Picard, J.

Species sauteri (Nayrolles & Lienhard, 1990) Bretfeld, 1992

Fasciosminthurus sauteri from Switzerland
After Nayrolles P & Lienhard C, 1990 in Bretfeld, 1999 Fig.529

Species strasseni (Paclt, 1964) Bretfeld, G, 1999:247

Species strigatus (Stach, 1922) Rusek, J in Dlabola, J, 1977:16

            Syn.: Fasciosminthurus circumfaciatus Dunger, 1975 cf
            Syn.: Prorastriopes circumfaciatus Christian & Kampichler, 1984
            Syn.: Prorastriopes circumfaciatus Thibaud & Christian, 1986 cf
            Syn.: Bourletiella carpatica Nosek, 1978
            Syn.: Bourletiella carpatica Bretfeld, 1992


  Subspecies sajanensis Bretfeld, 1996

             Syn.: Fasciosminthurus strigatus ssp. sajanensi(sic) Bretfeld, G, 1999:256 ISS

Species tarianus Bretfeld, G, 2001:111

Fasciosminthurus tarianus from Algeria
After Bretfeld, 2001 Fig.35 right
Fasciosminthurus tarianus from Algeria
After Bretfeld, 2001 Fig.35 left

Species virgulatus (Skorikov, 1899) Delamare Deboutteville, C & Massoud, Z, 1963:256

            Syn.: Deuterosminthurus bilineatus var. plusilineata Stach, 1922
            Syn.: Bourletiella pluslineata Gisin, H, 1960:289,sd
            Syn.: Deuterosminthurus bilineatus ab. prodromata Stach, 1922


Bourletiella (Fasciosminthurus) virgulata from Jugoslavia
After Loksa I & Bogojevic J, 1970 Abb.39
Fasciosminthurus virgulatus from France
After Nayrolles P, 1997 Fig.11
Fasciosminthurus virgulatus from France
After Nayrolles P, 1997 Fig.12
Fasciosminthurus virgulatus from Ibiza
2010.08.27 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
2010.08.27 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
2010.08.27 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
2010.09.14 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
2014.10.14 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
2015.09.10 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
2015.09.10 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
2015.09.11 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
2015.09.14 © Beer, R.
Fasciosminthurus virgulatus from France
2017.04.22 © Tiky, S.
Fasciosminthurus virgulatus from France
2017.04.22 © Tiky, S.
Fasciosminthurus virgulatus from France
2017.05.29 © Tiky, S.
Fasciosminthurus virgulatus from France
2017.05.22 © Tiky, S.
Fasciosminthurus virgulatus from France
2017.07.20 © Tiky, S.
Fasciosminthurus virgulatus from Ibiza
2017.09.02 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
Pollen feeding
2017.09.03 © Beer, R.
Fasciosminthurus virgulatus from France
2018.05.06 © Tiky, S.
Fasciosminthurus virgulatus from France
2018.05.06 © Tiky, S.
Fasciosminthurus virgulatus from France
2018.05.06 © Tiky, S.
Fasciosminthurus virgulatus from France
Spermatophore
2018.05.10 © Tiky, S.
Fasciosminthurus virgulatus from France
2018.06.23 © Picard, J.
Fasciosminthurus virgulatus from France
2018.06.24 © Picard, J.
Fasciosminthurus virgulatus from France
Caudal aspect
2018.06.24 © Picard, J.
Fasciosminthurus virgulatus from France
Caudal aspect
2018.06.24 © Picard, J.


Genus Kaszabellina Betsch, J-M, 1977:79

Species major Betsch, J-M, 1977:83

Species minima Betsch, J-M, 1977:84

Species variabilis Betsch, J-M, 1977:79, t.t.


Genus Koupelia Bretfeld, G, 2001:111

Species foliata Bretfeld, G, 2001:112, t.t.


Genus Madecassiella Betsch & Waller, 1996

Species ibityensis Betsch & Waller, 1996, t.t.


Genus Massoudia Betsch, 1975

Species griveaudi Betsch, 1975


Genus Paulianitas Betsch, 1978

Species vietti Betsch, 1978


Genus Stenognathriopes Betsch, J-M et Lasebikan, BA, 1979:166

  Syn.: Sminthuridae(lapsus) Senognathriopes(sic) Vázquez-G, M et Tun Osorio, R, 2004:283 ISS

Species sp.1 Janssens F sensu Samalot-Roque, 2006

            Syn.: Stenognathriopes sp.nov. Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:24

Subgenus Stenognathriopes Betsch, J-M et Lasebikan, BA, 1979:166 s.s.

Species sp. Bretfeld, G, 2005:45

Species huetheri Betsch, J-M et Lasebikan, BA, 1979:166

            Syn.: Stenognathriopes hütheri(sic) Betsch, J-M et Lasebikan, BA, 1979:166 IOS
             Syn.: Stenognathriopes hutheri(sic) Zeppelini, D et da Silva, DD, 2012:52 ISS
            Syn.: Stenognathriopes hütheri(sic) Thibaud, J-M, 2013:236 IOS



Species interpositus (Hüther, 1967) Betsch, J-M & Lasebikan, BA, 1979:166

Species rastrifer (Denis, 1948) Betsch, J-M & Lasebikan, BA, 1979:166

Species vilhenai (Delamare Deboutteville, C & Massoud, Z, 1965:73) Betsch, J-M & Lasebikan, BA, 1979:166

Rastriopes vilhenai from Angola
After Delamare Deboutteville C & Massoud Z, 1965 Fig.4E
            Syn.: Rastriopes vilhenai Delamare Deboutteville, C et Massoud, Z, 1964:73 u


Species yemenensis Bretfeld, G, 2005:42

Subgenus Tenentiella Palacios-Vargas, JG et Vázquez, M, 1997:285
Species janssensi Zeppelini, D et da Silva, DD, 2012:52

Stenognathriopes (Tenentiella) janssensi ♀ from Brazil
After Zeppelini D & da Silva DD, 2012 Fig.1A
Stenognathriopes (Tenentiella) janssensi ♀ from Brazil
After Zeppelini D & da Silva DD, 2012 Fig.1B
Stenognathriopes (Tenentiella) janssensi from Brazil
After Zeppelini D & da Silva DD, 2012 Fig.2

Species siankaana Palacios-Vargas, JG et Vázquez, M, 1997:286, t.t.


Genus Tritosminthurus Snider, RJ, 1988:260

Species schuhi Snider, RJ, 1988:260, t.t.


References