Checklist of the Collembola: Heteromurinae

http://www.collembola.org/taxa/heteinae.htm - Last updated on 2024.08.26 by Frans Janssens

Checklist of the Collembola: Heteromurinae

Subfamilia Heteromurinae Absolon & Kseneman, 1942 sensu Zhang F et Deharveng L, 2015:307

Genera with number of species
Genus	Number of species
Alloscopus	13
Dicranocentrus	71
Falcomurus	5
GenusnovnotDicranocentrus	1
GenusnovnrDicranocentrus	1
Heteromurtrella	20
Heteromurus (Heteromurus)	18
Heteromurus (Verhoeffiella)	14
Heteromurus	32
Mastigoceras	1
Pseudodicranocentrus	3
Sinodicranocentrus	4
Heteromurinae	151

  Syn.: Entomobryidae Entomobryini(sic) Börner, C, 1901:15 partim ISS
  Syn.: Collembola Tomoceridae Islam, S, Alim, A et Hossain, M, 2014:107 lapsus
  Syn.: Collembola Orchesellini Winter, C, 1966:165
  Syn.: Entomobryidae Orchesellinae Zhang, F, Chen, Z, Dong, R-R, Deharveng, L, Stevens, MI, Huang, Y-H et Zhu, CD, 2014:234
  Syn.: Collembola Isotomidae Wynne, JJ, Boyero Gallardo, JR, Hershauer, S, Huerta, OT, Martín, RF, Cuenca Rodriguez, J, París, M et Pardo-Igúzquiza, E, 2018:27

Heteromurinae from Australia
2009.09.25 © Daley, A.

Syn.: Entomobryidae Heteromurininae(sic) Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:153 ISS

Remark by Bellini BC, & al. 2020:24: We consider the alternative hypothesis to Entomobryoidea topology as Orchesellidae + (Heteromurinae + Entomobryidae/Paronellidae), the presence of scales could be the synapomorphy of the more derived Entomobryoidea, including Heteromurinae, and it was secondarily lost possibly more than one time among the Entomobryinae.

Tribus Heteromurini Absolon & Kseneman, 1942 sensu Zhang F & Deharveng L, 2015:309

  Syn.: ... Verhoeffiella Bellini, 2014

Genus Sinodicranocentrus Zhang F in Zhang, F, Cipola, NG, Pan, Z-X et Ding, Y-H, 2020:4

  Syn.: Dicranocentrus gaoligongensis Zhang & al, 2018 gr
  Syn.: Dicranocentrus gaoligonensis(sic) Zhang, F, Cipola, NG, Pan, Z-X et Ding, Y-H, 2020:12,gr ISS

Species gaoligongensis (Zhang F in Zhang F & al, 2018 tt) Zhang, F, Cipola, NG, Pan, Z-X & Ding, Y-H, 2020:5

Species pallidus (Zhang F in Zhang F & al, 2018) Zhang, F, Cipola, NG, Pan, Z-X & Ding, Y-H, 2020:5

Species similis (Zhang F in Zhang F & al, 2018) Zhang, F, Cipola, NG, Pan, Z-X & Ding, Y-H, 2020:5

Species varicolor (Zhang F in Zhang F & al, 2018) Zhang, F, Cipola, NG, Pan, Z-X & Ding, Y-H, 2020:5

Genus Dicranocentrus Schött, H, 1893:21

  Syn.: Dicranocentrus Dicranocentrella Wray, DL, 1953:140
  Syn.: Heteromuricinae Heteromuricus Imms, 1912
  Syn.: Heteromurus Hetermuricus(sic) Mari Mutt, JA, 1979:544 ISS
  Syn.: Heteromurini Falcomurus Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:22 =?

Dicranocentrus from Peru
2009.01-03.dd © Damiano-Palomino., B

Dicranocentrus from Peru
2009.01-03.dd © Damiano-Palomino., B

Dicranocentrus from Peru
2009.01-03.dd © Damiano-Palomino., B

Syn.: Heteromurini Dicranoncetrus(sic) Bellini, BC, Greenslade, P, Baquero, E, Jordana, R, de Souza, PGC, 2022:254 ISS

Species sp.1 Janssens F & Cheng H-J, 2022

Dicranocentrus nr cercifer from Taiwan
2022.01.25 © Cheng, H.-J.

Dicranocentrus nr cercifer from Taiwan
Abd.6 with dorsal median cercus
2022.01.25 © Cheng, H.-J.

Dicranocentrus nr cercifer from Taiwan
Ant.3 distinctly longer and annulated
2022.01.25 © Cheng, H.-J.

Dicranocentrus nr cercifer from Taiwan
Abd.6 with dorsal median cercus
2022.01.29 © Cheng, H.-J.

Dicranocentrus nr cercifer from Taiwan
2022.02.01 © Cheng, H.-J.

Dicranocentrus nr cercifer from Taiwan
Antennae
2022.02.01 © Cheng, H.-J.

Dicranocentrus nr cercifer from Taiwan
Ant.1+2 subdivided
2022.02.01 © Cheng, H.-J.

Dicranocentrus nr cercifer from Taiwan
Abd.3 to 6
2022.02.01 © Cheng, H.-J.

Dicranocentrus nr cercifer from Taiwan
Abd.6 with elongated dorsal anal valve
2022.02.01 © Cheng, H.-J.

Dicranocentrus nr cercifer from Taiwan
Abd.4-5-6
2022.02.01 © Cheng, H.-J.

Dicranocentrus nr cercifer ♀ from Taiwan
Abd.6 dorsal anal valve not elongated
2022.02.03 © Cheng, H.-J.

Dicranocentrus nr cercifer ♀ from Taiwan
Abd.6 short
2022.02.03 © Cheng, H.-J.

Dicranocentrus nr cercifer ♀ from Taiwan
Female gonopore
2022.02.03 © Cheng, H.-J.

Dicranocentrus nr cercifer ♀ from Taiwan
Dental spines
2022.02.03 © Cheng, H.-J.

Dicranocentrus nr cercifer ♂? from Taiwan
Abd.6 dorsal anal valve elongated
2022.02.03 © Cheng, H.-J.

Dicranocentrus nr cercifer ♂? from Taiwan
Abd.6 dorsal anal valve elongated
2022.02.03 © Cheng, H.-J.

Dicranocentrus nr cercifer ♂? from Taiwan
Abd.6 dorsal anal valve elongated
2022.02.03 © Cheng, H.-J.

Dicranocentrus nr cercifer ♂? from Taiwan
Abd.6 dorsal anal valve elongated
2022.02.03 © Cheng, H.-J.

Dicranocentrus nr cercifer ♂ subadult? from Taiwan
Abd.6 dorsal anal valve slightly elongated
2022.02.05 © Cheng, H.-J.

Dicranocentrus nr cercifer ♀? from Taiwan
Abd.6 dorsal anal valve not elongated
2022.02.05 © Cheng, H.-J.

Species abestadorum Siqueira OJR, Bellini BC et Cipola NG in Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:4

Syn.: Dicranocentrus (marias-group) abestado(sic) Siqueira OJR, Bellini BC et Cipola NG in Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:4 IOS

Dicranocentrus abestado(sic) from Brazil
After Bellini BC & al, 2020 Fig.1

Remark by Janssens, F. 2020.10.26: The etymology (p.10) : "'abestado' or 'abestada' is a regional expression used in Northeastern Brazil which means 'fool'. The expression is commonly used between friends." does not explain the origin of the new name but simply the meaning of the vernacular word 'abestado'... So, the new name refers to 'Dicranocentrus, the fool'. Although it is not explicitly stated by the etymology, it is clear that it was the intention of the authors to reflect in the new name the friendship between the authors, as in the 'Dicranocentrus of the fools'. The mismatch between the name and the intention of the authors is clear evidence of an inadvertent error that must be corrected (Art.32.5.1). In addition, an unmodified vernacular word should not be used as a scientific name (Art.11.3 Rec.11A). Given appropriate latinization is the preferred means of formation of names from vernacular words (Rec.11A) : in analogy with 'muscorum' = 'of the mosses', we propose the following emendation of the name that matches the intention of the authors: 'abestadorum' = stem 'abestad' + latin plural genitive case suffix 'orum'.

            Syn.: Dicranocentrus (marias-group) abestado(sic) Siqueira OJR, Bellini BC et Cipola NG in Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:5 IOS
            Syn.: Dicranocentrus (marias-group) abestado(sic) Siqueira OJR, Bellini BC et Cipola NG in Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:6 IOS
            Syn.: Dicranocentrus (marias-group) abestado(sic) Siqueira OJR, Bellini BC et Cipola NG in Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:7 IOS
            Syn.: Dicranocentrus (marias-group) abestado(sic) Siqueira OJR, Bellini BC et Cipola NG in Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:8 IOS
            Syn.: Dicranocentrus (marias-group) abestado(sic) Siqueira OJR, Bellini BC et Cipola NG in Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:11 IOS
            Syn.: Dicranocentrus (marias-group) abestado(sic) Siqueira OJR, Bellini BC et Cipola NG in Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:12 IOS
            Syn.: Dicranocentrus (marias-group) abestado(sic) Siqueira OJR, Bellini BC et Cipola NG in Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:14 IOS
            Syn.: Dicranocentrus (marias-group) abestado(sic) Siqueira OJR, Bellini BC et Cipola NG in Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:15 IOS
            Syn.: Dicranocentrus (marias-group) abestado(sic) Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:16 IOS
            Syn.: Dicranocentrus (marias-group) abestado(sic) Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:20 IOS
            Syn.: Dicranocentrus (marias-group) abestado(sic) Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:21 IOS
            Syn.: Dicranocentrus (marias-group) abestado(sic) Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:22 IOS
            Syn.: Dicranocentrus (marias-group) abestado(sic) Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:24 IOS
            Syn.: Dicranocentrus abestado(sic) Cipola, NG, 2022:8 IOS

Species amazonicus Bellini, Morais & Oliveira, 2013

Species antillensis Mari Mutt, JA, 1979:57

Subspecies jamaicensis Mari Mutt, JA, 1981:99

Species bicolor Handschin, 1924

Syn.: Dicranocentrus bicolor Xisto, T et de Mendonça, MC, 2017:34 nd

Dicranocentrus cf. bicolor from Brazil
After Cipola NG & al, 2019 Fig.3

Species biseriatus Mari Mutt, JA, 1981:97

Syn.: Dicranocentrus millsi Mari Mutt, JA, 1979:61

Species celatus Mari Mutt, JA, 1985:105

            Syn.: Dicranocentrus sp. Mari Mutt, 1977
            Syn.: Dicranocentrus millsi Mari Mutt, 1981

             Syn.: Dicranocentrus celarus(sic) Mari Mutt, JA, 1985:109 ISS

Species cercifer (Imms, 1912) Mari Mutt, JA, 1979:543

            Syn.: Tomocerus minor Islam, S, Alim, A et Hossain, M, 2014:107 lapsus
            Syn.: Tomocerus minor Islam, S, Alim, A et Hossain, M, 2014:109 lapsus
            Syn.: Tomocerus minor Islam, S, Alim, A et Hossain, M, 2014:110 lapsus
            Syn.: Tomocerus minor Islam, S, Alim, A et Hossain, M, 2014:111 lapsus

Tomocerus minor (lapsus) from Bangladesh
With 'extended organ at end point of anus'
After Islam S, Alim A & Hossain M, 2014 Fig.3O

Tomocerus minor (lapsus) from Bangladesh
With 'extended organ at end point of anus'
After Islam S, Alim A & Hossain M, 2014 Fig.3P

Syn.: Heteromuricus crucifer(sic) Schött, 1925 ISS

Heteromuricus cercifer from South-India
After Handschin E, 1929 Fig.48

Heteromuricus cercifer from India
After Mandal GP & Hazra AK, 2009 Fig.78a

Dicranocentrus cercifer from India
Abd.6 with dorsal median cercus
After Mandal GP & Suman KK, 2015 Fig.p.524

Dicranocentrus cercifer from India
After Mandal GP & al, 2019 Plate VI left

Species chenae Ma, Y-T, Chen, J-X et Soto-Adames, FN, [2007]:485

Species chimborazoensis Najt, J, Thibaud, J-M et Mari Mutt, JA, 1988:553

Dicranocentrus chimborazoensis
After Thibaud J-M & D'Haese C, 2010 Fig.9

Species dolosus Bellini BC, Greenslade P et Souza PGC in Bellini, BC, Greenslade, P, Baquero, E, Jordana, R, de Souza, PGC, 2022:224

Dicranocentrus dolosus from Cocos & Keeling Islands
After Bellini BC & al, 2022 Fig.2

Syn.: Dicranocentrus sp. Bellini, BC, de Souza, PGC et Greenslade, P, 2021:17
Syn.: Dicranocentrus sp. Bellini, BC, Greenslade, P, Baquero, E, Jordana, R, de Souza, PGC, 2022:222

Species eburneus Delamare Deboutteville, 1951 nn

Species fasciatus Yosii, 1961

Species gemellus Mari Mutt, JA, 1985:321

Species hainanicus Ren Y-H et Zang F in Ren, Y-H, Li, Z-H et Zhang, F, 2018:61

Dicranocentrus hainanicus from China
After Ren Y-H, Li Z-H & Zhang F, 2018 Fig.1

Dicranocentrus hainanicus from China
Eyepatch with 9 ocelli in lapsus
Modified after Ren Y-H, Li Z-H & Zhang F, 2018 Fig.17

Species icelosmarias Soto-Adames, FN et Anderson, EW, 2017:33

Syn.: Dicranocentrus marias Mari Mutt, 1979 from Panama =?

Species indicus Bonet, F, 1930:255

Dicranocentrus indicus from India
After Bonet F, 1930 Fig.3A

Dicranocentrus indicus from India
After Mandal GP & Hazra AK, 2009 Fig.36a

Species inermodentes (Uchida, 1944) Mari Mutt, JA, 1979:67

Species janetscheki Yosii, 1971

Species javanus Yoshii & Yayuk, 1989:62

Subspecies salatiganus Yoshii, R et Suhardjono, YR, 1989:62

Species lawrencei Mari Mutt, JA, 1979:66

Species linnaniemii Börner, 1912

             Syn.: Dicranocentrus Linnaniemi(sic) Denis, J-R, 1933:260 ISS
             Syn.: Dicranocentrus Linnaniemii(sic) Denis, J-R, 1933:260 ISS
             Syn.: Dicranocentrus Linnaniemii(sic) Delamare Deboutteville, C, 1945:33 ISS
             Syn.: Dicranocentrus linnaniemi(sic) Thibaud, J-M, 2013:239 ISS

Species liuae Xu, G-L et Zhang, F, 2014:3

Dicranocentrus liuae from China
After Xu G-L & Zhang F, 2014 Fig.2

Species marias Wray, DL, 1953:141

            Syn.: Dicranocentrus silvestrii Denis, 1931
             Syn.: Dicranocentrus Silvestrii(sic) Denis, J-R, 1933:255 ISS
             Syn.: Dicranocentrus Silvestrii(sic) Denis, J-R, 1933:261 ISS
             Syn.: Dicranocentrus Silvestrii(sic) Denis, J-R, 1933:319 ISS

             Syn.: Dicranocentrus maria(sic) Smith, CL, 2015:57 u ISS

Species millsi Mari Mutt, JA, 1979:61

Syn.: Dicranocentrus millsi ssp. cubensis Mari Mutt, JA, 1979:62
Syn.: Dicranocentrus sp. Mari Mutt, 1978

Species orientalis Mari Mutt, JA, 1979:67

Species peckorum Najt, J, Thibaud, J-M et Mari Mutt, JA, 1988:556

Species pikachu Xisto, T et de Mendonça, MC, 2017:16

Dicranocentrus pikachu from Brazil
After Xisto T & de Mendonça MC, 2017 Fig.6

Species problematicus Denis, 1948

Species reunionensis Mari Mutt, JA, 1985:299

Species silvestrii Absolon, K, 1903:105

            Syn.: Dicranocentrus Silvestrii(sic) Absolon, K, 1903:105 IOS
             Syn.: Tomocerus plumbaeus(sic) Parona, 1895 ISS
             Syn.: Tomocerus plumbaeus(sic) Bonet, F, 1933:123 ISS
             Syn.: Tomocerus plumbaeus(sic) Bonet, F, 1933:125 =? ISS
            Syn.: Dicranocentrus silvestrii var. nigrescens Absolon, 1903 =?
            Syn.: Dicranocentrus silvestrii var. annulata Börner, C, 1906:176
            Syn.: Dicranocentrus bicolor Cassagnau, P in Delamare Deboutteville, C et Rapoport, E, 1963:128 =?
            Syn.: Dicranocentrus bicolor Arlé & Mendonça, 1982 =?
            Syn.: Dicranocentrus termitophilus Arlé & Mendonça, 1982 =?

             Syn.: Dicranocentrus Silvestrii(sic) Cassagnau, P in Delamare Deboutteville, C et Rapoport, E, 1963:127 ISS
             Syn.: Dicranocentrus silvestri(sic) Abrantes, EA, Bellini, BC, Bernardo, AN, Fernandes, LH, Mendonça, MC, Oliveira, EP, Queiroz, GC, Sautter, KD, Silveira, TC et Zeppelini, D, 2010:2 ISS
            Syn.: Dicranocentrus silvestrii Xisto, T et de Mendonça, MC, 2017:34 nd

Species simplex Yosii, R, 1959:41

Species solomonensis Mari Mutt, JA, 1979:68

Species spinosus Prabhoo, 1971

Species stachi (Denis, 1925) Handschin, 1929

            Syn.: Heteromurus (Heteromuricus) Stachi(sic) Denis, 1925 IOS
            Syn.: Heteromurus major ssp. Stachi(sic) Stach, J, 1930:294 =? IOS
            Syn.: Heteromuricus Stachi(sic) Denis, J-R, 1933:259 IOS
            Syn.: Heteromurus Stachi(sic) Denis, J-R, 1933:259 IOS
            Syn.: Heteromurus Stachi(sic) Denis, J-R, 1933:320 IOS
            Syn.: Dicranocentrus? Stachi(sic) Delamare Deboutteville, C, 1945:34 IOS

Species sundanensis Schött, 1925

Syn.: Dicranocentrus sudanensis(sic) Jacquemart, S, 1980:5 ISS

Species termitophilus Handschin, 1924

Syn.: Dicranocentrus termitophilus Xisto, T et de Mendonça, MC, 2017:34 nd
Syn.: Dicranocentrus termithophilus(sic) Xisto, T et de Mendonça, MC, 2017:35 ISS

Species wangi Ma, YT et Chen, J-X, 2007:?

Species yoshiius Mari Mutt, JA, 1988:179

Species zebratus Mari Mutt, JA, 1985:106

Syn.: Dicranocentrus sp. Mari Mutt, 1977
Syn.: Dicranocentrus millsi Mari Mutt, 1981

Species-group gracilis

Species albicephalus Xisto, T et de Mendonça, MC, 2017:23

Dicranocentrus albicephalus from Brazil
After Xisto T & de Mendonça MC, 2017 Fig.10

Species assimilis Schött, 1927

Species bidentatus Mari Mutt, JA, 1983:154

Species capitaneus Mari Mutt, JA, 1985:297

Species colombiensis Mari Mutt, JA, 1979:23

Syn.: Dicranocentrus gracilis Winter, C, 1963:507 =?

Dicranocentrus colombiensis from Colombia
After Cipola NG, 2023 Fig.2C

Species cuprum Xisto & de Mendonça, 2016

Species deharvengi Mari Mutt, JA, 1981:493

Species delamarei Mari Mutt, JA, 1981:160

Species edicitae Jacquemart, S, 1980:2

Species fraternus Mari Mutt, JA et Bhattacharjee, RK, 1980:162

Species gapudi Mari Mutt, JA, 1985:303

Species gracilis Schött, H, 1893:23, t.t.

Species heloisae Arlé & Mendonça, 1982

Dicranocentrus heloisae from Brazil
After Xisto T & de Mendonça MC, 2014 Fig.1

Dicranocentrus heloisae from Brazil
After Xisto T & de Mendonça MC, 2017 Fig.18a

Dicranocentrus heloisae from Brazil
After Xisto T & de Mendonça MC, 2017 Fig.18b

Species indecisus Mari Mutt, JA, 1985:301

Species inermis Schött, 1927

Syn.: Dicranocentrus insignis(sic) Yosii, 1961 lapsus

Species longicornis (Carpenter, GH, 1916:36) Handschin, 1929

Species luzonensis Mari Mutt, JA, 1985:317

Species magnus Xisto, T et de Mendonça, MC, 2017:11

Dicranocentrus magnus from Brazil
After Xisto T & de Mendonça MC, 2017 Fig.2

Species marimutti Xisto, T et de Mendonça, MC, 2017:28

Dicranocentrus marimutti from Brazil
After Xisto T & de Mendonça MC, 2017 Fig.14

Species melinus Xisto & de Mendonça, 2016

Species meruensis Wahlgren, 1908

             Syn.: Heteromurus (Heteromuricus) Stachi(sic) Denis, 1925 =? ISS
            Syn.: Dicranocentrus aethiopicus Handschin, 1929
             Syn.: Dicranocentrus æthiopicus(sic) Denis, J-R, 1933:260 ISS
            Syn.: Dicranocentrus aethiopicus Mari Mutt, JA, 1979:55
            Syn.: Dicranocentrus Jeanneli(sic) Delamare Deboutteville, C, 1945:31 IOS
            Syn.: Dicranocentrus Jeanneli(sic) Jeannel, R, 1945:18 IOS

Species nepalensis Mari Mutt JA in Mari Mutt, JA et Bhattacharjee, RK, 1980:165

Dicranocentrus nepalensis from India
After Mandal P & al, 2024 Fig.2A

Species nigritus Mari Mutt, JA, 1979:23

Species orellanae Jacquemart, S, 1980:6

Syn.: Dicranocentrus orellanae ssp. cuzcoensis Jacquemart, S, 1980:8

Species pampaensis Silva CDD et Bellini BC in da Silva, CDD, Nunes, RC, Winck, BR et Bellini, BC, 2024:538

Dicranocentrus pampaensis from Brazil
After da Silva CDD & al, 2024 Fig.1

Syn.: Dicranocentrus sp.1 da Silva & al, 2022
Syn.: Dicranocentrus sp.1 da Silva, CDD, Nunes, RC, Winck, BR et Bellini, BC, 2024:551

Species paramoensis Mari Mutt, JA, 1983:149

Syn.: Dicranocentrus paramoense(sic) Mari Mutt, JA, 1983:149 IOS
Syn.: Dicranocentrus paramoense(sic) Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:158 ISS

Species platensis (Izarra, 1972) Mari Mutt, JA, 1977:248

Syn.: Tomocerus plumbeus Parona, 1895 =?
Syn.: Dicranocentrus silvestrii Bonet, 1934

Dicranocentrus platensis from Argentina
2016.03.27 © Chiavone, J.

Species schoetti Mari Mutt, JA, 1979:56

Syn.: Dicranocentrus maculatus Winter, C, 1963:509 nn

Remark by Winter, C. 1963:395: 37 von mir neu aufgestellte Arten bzw. Formen sind in der vorliegende Arbeit mit n.sp. bzw. n.f. (ohne Autorennamen) bezeichnet und müssen bis zu ihrer Beschreibung (voraussichtlich im kommenden Jahr) als Nomina nuda betrachtet werden.

Syn.: Dicranocentrus maculatus Mari Mutt, JA, 1979:57 =?
Syn.: Dicranocentrus muyensis Winter, C, 1963:508 nn

Syn.: Dicranocentrus muyensis Mari Mutt, JA, 1979:57 =?
Syn.: Dicranocentrus silvestrii f. circulata(sic) Winter, C, 1963:508 nn =? ISS

Syn.: Dicranocentrus silvestrii var. circulata(sic) Mari Mutt, JA, 1979:57 =? ISS

Species singularis Mari Mutt, JA et Bhattacharjee, RK, 1980:168

Species thaicus Yosii, 1961

Species violaceus Mari Mutt, JA, 1981:496

Genus GenusnovnotDicranocentrus Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:154

Species pilosus (Mari Mutt JA in Mari Mutt, JA & Bhattacharjee, RK, 1980:168) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:154

Genus GenusnovnrDicranocentrus Greenslade, P et Jordana, R, 2014:561

Species sp. Greenslade, P et Jordana, R, 2014:561

Genus Falcomurus Mandal, GP, 2018:75

Species chilikaensis Mandal, GP, 2018:77, t.t.

Falcomurus chilikaensis from India
After Mandal GP, 2018 Fig.1

Species halophilus (Mari Mutt, JA, 1985:320) Bellini, BC, de Souza, PGC & Greenslade, P, 2021:8

Species hilli Bellini, BC, de Souza, PGC et Greenslade, P, 2021:19

Species litoreus (Mari Mutt, JA, 1985:318) Bellini, BC, de Souza, PGC & Greenslade, P, 2021:9

Species pulukokosensis Bellini, BC, de Souza, PGC et Greenslade, P, 2021:14

Syn.: Falcomurus pulukokos(sic) Bellini, BC, de Souza, PGC et Greenslade, P, 2021:14 IOS

Falcomurus pulukokos(sic) from the Cocos Islands
After Bellini BC, de Souza PGC & Greenslade P, 2021 Fig.1A

Remark by Janssens, F. 2021.07.17: According to the etymology : "The epithet refers to the Malay's name for the Keeling territory". However, 'Falcomurus pulukokos' means 'the Falcomurus, Pulu Kokos', not 'the Falcomurus from the Cocos Islands'. It does not refer to a species from the Keeling territory as the authors intended. This meaning does not match the etymology. This mismatch is clear evidence of an inadvertent error that must be corrected (Art.32.5.1). 'pulukokosensis' refers to 'from the Keeling territory ("Pulu Kokos" in the Malay language)', in line with the etymology.

            Syn.: Falcomurus pulukokos(sic) Bellini, BC, de Souza, PGC et Greenslade, P, 2021:4 IOS
            Syn.: Falcomurus pulukokos(sic) Bellini, BC, de Souza, PGC et Greenslade, P, 2021:14 IOS
            Syn.: Falcomurus pulukokos(sic) Bellini, BC, de Souza, PGC et Greenslade, P, 2021:15 IOS
            Syn.: Falcomurus pulukokos(sic) Bellini, BC, de Souza, PGC et Greenslade, P, 2021:16 IOS
            Syn.: Falcomurus pulukokos(sic) Bellini, BC, de Souza, PGC et Greenslade, P, 2021:17 IOS
            Syn.: Falcomurus pulukokos(sic) Bellini, BC, de Souza, PGC et Greenslade, P, 2021:24 IOS
            Syn.: Falcomurus pulukokos(sic) Bellini, BC, de Souza, PGC et Greenslade, P, 2021:25 IOS
            Syn.: Falcomurus pulukokos(sic) Bellini, BC, de Souza, PGC et Greenslade, P, 2021:26 IOS
            Syn.: Falcomurus pulukokos(sic) Bellini, BC, de Souza, PGC et Greenslade, P, 2021:27 IOS
            Syn.: Falcomurus pulukokos(sic) Bellini, BC, de Souza, PGC et Greenslade, P, 2021:29 IOS
            Syn.: Falcomurus pulukokos(sic) Bellini, BC, Greenslade, P, Baquero, E, Jordana, R, de Souza, PGC, 2022:221 IOS
            Syn.: Falcomurus pulukokos(sic) Bellini, BC, Greenslade, P, Baquero, E, Jordana, R, de Souza, PGC, 2022:222 IOS
            Syn.: Falcomurus pulukokos(sic) Bellini, BC, Greenslade, P, Baquero, E, Jordana, R, de Souza, PGC, 2022:231 IOS
            Syn.: Falcomurus pulukokos(sic) Bellini, BC, Greenslade, P, Baquero, E, Jordana, R, de Souza, PGC, 2022:232 IOS
            Syn.: Falcomurus pulukokos(sic) Bellini, BC, Greenslade, P, Baquero, E, Jordana, R, de Souza, PGC, 2022:233 IOS
            Syn.: Falcomurus pulukokos(sic) Bellini, BC, Greenslade, P, Baquero, E, Jordana, R, de Souza, PGC, 2022:253 IOS

Genus Pseudodicranocentrus Mari Mutt, JA, 1981:91

Syn.: Dicranocentrus circulatus Mari Mutt, JA, 1979:6,gr
Syn.: Dicranocentrus circulatus Bellini, BC, Cipola, NG et Siqueira, OJR, 2020:22,gr

Species christianseni (Mari Mutt, JA, 1979:21) Mari Mutt, JA, 1981:93

Species circulatus (Mari Mutt, JA, 1979:21) Mari Mutt, JA, 1981:93, t.t.

Species niger Mari Mutt, JA, 1984:816

Genus Heteromurus Wankel, 1860 nc Opinion 1064, 1976

  Syn.: Podura Ptenura Templeton, 1844
  Syn.: Podurelles(sic) Heteroma(sic) Nicolet, H, 1847:366 ISS
  Syn.: Podurides(sic) Aetheocerus Elditt, HL, 1854:41 partim
  Syn.: Poduridæ(sic) Templetonia Lubbock, J, 1862:595
  Syn.: Entomobryinae Typhlopodura Absolon, K, 1900:428
  Syn.: ... Heteromurodes Absolon, 1901
  Syn.: Heteromurini Heteromuroder(sic) Mari Mutt, JA, 1980:456 ISS
  Syn.: ... Euheteromurus Börner, 1901
  Syn.: Entomobryinae Propemesira Salmon, JT, 1942:59
  Syn.: ... Troglosinella Salmon, 1955, nec Delamare Deboutteville, 1949
  Syn.: ... Heteromurtella Mari Mutt, 1979
  Syn.: Entomobryomorpha Desoria? Wynne, JJ, Boyero Gallardo, JR, Hershauer, S, Huerta, OT, Martín, RF, Cuenca Rodriguez, J, París, M et Pardo-Igúzquiza, E, 2018:3
  Syn.: Isotomidae Desoria? Wynne, JJ, Boyero Gallardo, JR, Hershauer, S, Huerta, OT, Martín, RF, Cuenca Rodriguez, J, París, M et Pardo-Igúzquiza, E, 2018:27

  Syn.: Collembola Heteromuris(sic) Christiansen, K, 1958:475 ISS

Remark by ICZN. 2007:384: Placed on the Official List of Names in Zoology. Opinion 1064.

Heteromurus sp. from Russia
2008.10.21 © Macroclub.ru

Heteromurus major from the UK
With 5 secondary antennal segments
2009.07.26 © Barton, T.

  Syn.: Collembola Hetermurus(sic) Mejía-Recamier, BE, Palacios-Vargas, JG et Franco, A, 2012:27 ISS
  Syn.: Heteromurini Heterumurus(sic) Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:155 ISS
  Syn.: Collembola Heteromurusmajor(sic) Qazi, F et Shayanmehr, M, 2016:122 lapsus ISS

Heteromurus sp. from the USA
Abd.4 shorter than 2 times abd.3
2016.11.28 © Dobbs, Br.

Heteromurus sp. from the Canary Islands
Eating from wet yeast grain
2022.11.30 © Huskens, M.-L.

Heteromurus sp. from the Canary Islands
Exposed mouthparts
2022.11.30 © Huskens, M.-L.

Subgenus Heteromurus Mari Mutt, JA, 1979:218

Species sp.1 Shaw P, 2012:il

Syn.: Heteromurus nitidus Hopkin SP, 2004:il

Heteromurus nitidus from the UK
2004 © Hopkin, S.P.

Heteromurus nr nitidus from the UK
Descaled revealing intrinsic pigmentation
2012.04.23 © Shaw, P.

Heteromurus nr nitidus from Hungary
Dark intrinsic pigmentation
2012.05.21 © Lásló, D.

Heteromurus nr nitidus from the UK
2013.01.23 © Cobby, P.

Heteromurus nr nitidus from the UK
Dark intrinsic pigmentation
2017.04.17 © Cann, A.J.

Heteromurus nr nitidus from the UK
With dark facial mask
2017.04.17 © Cann, A.J.

Heteromurus nr nitidus from the UK
2018.11.16 © Uncle, F.

Heteromurus nr nitidus from the UK
With pale streaks on mesothorax
2018.11.16 © Uncle, F.

Heteromurus nitidus from Croatia
After Deharveng L in Lukic M, 2019 Fig.4D

Species sp.2 Janssens F & Dobbs B, 2016

Heteromurus sp. from the USA
2016.11.28 © Dobbs, Br.

Species sp.3 Greenslade, P et Jordana, R, 2014:561

Species sp.4 Greenslade, P et Jordana, R, 2014:561

Species caophongensis Nguyen, TT, 1995:563

Heteromurus caophongensis from North Vietnam
After Nguyen TT, 1995 Hinh.4.1

Species dogieli (Philiptschenko, 1926) Mari Mutt, JA, 1979:544

Syn.: Heteromuricus Dogieli(sic) Denis, J-R, 1933:259 ISS

Species fuscatus (Koch, CL & Berendt, GC in Berendt, GC, 1854:119) -

Podura fuscata from Baltic amber
After Koch CL & Berendt GC, 1845 Tab.XV Fig.127

Species gigans Mari Mutt, JA et Stomp, N, 1980:183

Species major (Moniez, 1889) Absolon, 1901, B

            Syn.: Podura teres Linnæus, C, 1746:342
            Syn.: Podura plumbea Linnaeus, C, 1761:473
            Syn.: Podura (Longæ) plumbea Geoffroy, M, 1762:610
            Syn.: Podura (Longæ) plumbea Geoffroy, M, 1764:610
            Syn.: Podura plumbeo(sic) de Geer, C, 1778:31

Podura corpore teriti nigrocaeruleo plumbeo
from Sweden
After de Geer, C., 1778 Tom.VII Pl.3 Fig.1.

            Syn.: pondura(sic) plumbea de Bomare, V, 1780:296
            Syn.: Podura plumbea Fabricius, JC, 1793:66
            Syn.: Podura plumbea Fabricius, JC, 1796:138
            Syn.: Podura (Longæ) plumbea Geoffroy, M, 1799:610
            Syn.: Podura plumbea Walckenær, CA, 1802:4
            Syn.: Podura plumbea Latreille, PA, 1806:166
            Syn.: Podura (Podura) plumbea Latreille, X in Cuvier, G, 1817:162
            Syn.: Podura plumbea de Lamarck, M, 1818:21
            Syn.: Podura plumbea Latreille, Le Peletier de Saint-Fargeau, Serville et Guérin in Latreille, M, 1825:167
            Syn.: Podura (Longæ) violacea Geoffroy, M, 1762:611
            Syn.: Heteromurus caerulescens Börner, 1903
             Syn.: Heteromurus coerulescens(sic) Handschin, E, 1942:426 ISS
             Syn.: Heteromurus Coerulescens(sic) Handschin, E, 1942:426 ISS
            Syn.: Heteromurus melitensis Stach, J, 1924:115

Heteromurus melitensis from Malta
After Stach J, 1924 Taf.I Fig.13

             Syn.: Heteromurus melittensis(sic) Handschin, E, 1942:425 ISS
             Syn.: Heteromurus Melittensis(sic) Handschin, E, 1942:426 ISS
            Syn.: Heteromurus mexicanus Handschin, 1928
            Syn.: Lepidocyrtus lundbladi Agrell, 1939
            Syn.: Heteromurus caucasicus Tshelnokov in Martynova, Tshelnokov & Rasulova, 1974

Heteromurus major from Malta
After Stach J, 1967 Pl.XXXIII Fig.1

Heteromurus (Heteromurus) major
After Mari Mutt, J.A., 1980 Fig.2

Heteromurus (Heteromurus) major
After Mari Mutt, J.A., 1980 Fig.3

Heteromurus major from the UK
2004 © Hopkin, S.P.

Heteromurus major from the UK
2004 © Hopkin, S.P.

Heteromurus major from the UK
2004 © Hopkin, S.P.

Heteromurus major from the UK
2006 © Barton, T.

Heteromurus major from France
2006 © Boldsystems.org

Heteromurus major from the UK
2007 © Cornwall, N.J.

Heteromurus major from Portugal
2007.02.25 © Almeida, J.

Heteromurus major from the UK
2007.03.14 © Barton, T.

Heteromurus major from the UK
2007.09.03 © Barton, T.

Heteromurus major from the UK, juvenile
2007.09.13 © Barton, T.

Heteromurus major from the UK, juvenile
2008.04.27 © Barton, T.

Heteromurus major from the Netherlands
2008.09.16 © Baas, A.H.

Heteromurus major from the Netherlands
2008.10.03 © Baas, A.H.

Heteromurus major from Holland
2011.08.01 © van Duinen, J.

Heteromurus major from Canada
2011.09.20 © Hochhalter, R.

Heteromurus major from the UK
2012.01.25 © Barton, T.

Heteromurus major from the UK
2012.01.25 © Barton, T.

Heteromurus major from La Palma
2012.12.02 © Barton, T.

Heteromurus major from France
2012.12.30 © Sandra.

Heteromurus major juv. from Holland
2013.02.17 © van Duinen, J.

Heteromurus major juv. from Holland
2013.02.25 © van Duinen, J.

Heteromurus major from France
Annulated apical antennal segment
2013.03.16 © Sandra.

Heteromurus major from the UK
Ant.1 subdivided in two parts (1a-1b)
2014.11.14 © Barton, T.

Heteromurus major from France
Relative short 4th abdominal segment
2014.11.23 © Quintin, C.

Heteromurus major from Belgium
2016.01.28 © Huskens, M.-L.

Heteromurus major from Greece
2016.12.16 © Dimitrakopoulos, C.

Heteromurus major from Spain
Ant.1 with basal subdivision (*)
2017.01.01 © Tizon, J.

Heteromurus major from Spain
2017.01.01 © Tizon, J.

Heteromurus major from Belgium
Body scales apically rounded
2017.01.22 © Huskens, M.-L.

Heteromurus major from Belgium
Eyepatch with 4+2 ocelli
2017.01.22 © Huskens, M.-L.

Heteromurus major from Holland
Interocular area with many macrosetae
2019.02.27 © Kamsteeg, G.

Heteromurus major juv. from Holland
Interocular area with many macrosetae
2019.02.27 © Kamsteeg, G.

Heteromurus major from Holland
Descaled
Abd.4 larger then abd.3
2019.12.09 © Kamsteeg, G.

Heteromurus major from Holland
Descaled
Eyepatch with 8 ocelli A-G
2019.12.09 © Kamsteeg, G.

Heteromurus major from Holland
Body scales
2020.01.22 © Kamsteeg, G.

Heteromurus major from Spain
2020.01.31 © Zeeders, M.

Heteromurus major from the UK
2021.02.21 © Barton, T.

Heteromurus major from the UK
Distinctly large dorsal iridiscent scales
2021.02.21 © Barton, T.

Heteromurus major egg from Belgium
With distinct polar cap
and hairy outgrowths
2022.04.02 © Huskens, M.-L.

Species nitidus (Templeton, R in Templeton, R & Westwood, JO, 1836:94) Megnin, 1888, B

Podura nitida from Ireland
After Templeton, R, 1836 Pl.XI Fig.5

            Syn.: Podura crystallina Müller OF, 1776
             Syn.: Heterotoma cristallina(sic) Bourlet, C, 1839:396 ISS
             Syn.: Podura cristallina(sic) Bourlet, C, 1839:396 ISS
             Syn.: Podura Crystallina(sic) Bourlet, C, 1845:153 ISS
            Syn.: Heteroma(sic) crystallina Nicolet, H, 1847:366

Templetonia crystallina from the UK
After Lubbock, J., 1873 Pl.20.

Templetonia crystallina from the UK
After Beck J in Lubbock J, 1873 Pl.69

            Syn.: Degeeria margaritacea Nicolet, H, 1842:75
            Syn.: Heteromurus nitidus f. margaritarius Wankel, 1860
             Syn.: Heteromurus nitidus ssp. margaritaria(sic) Jordana, R et Baquero, E, 2010:364 ISS
            Syn.: Lepidocyrtus Albinos(sic) McIntire, SJ, 1867:57

Templetonia nitida from the UK
After McIntire SJ, 1867 Fig.46
After McIntire SJ, 1868

Templetonia nitida from the UK
After McIntire SJ, 1867 Fig.47
After McIntire SJ, 1868

Syn.: Lepidocyrtus albinos(sic) Packard, AS, 1869:625 ISS

Lepidocyrtus albinos(sic) from Europe
After Packard AS, 1869 Fig.620

Lepidocyrtus albinos(sic) from Europe
Scale thin with distinct markings
After Packard AS, 1869 Fig.621

             Syn.: Lepidocyrtus albinos(sic) Packard, AS, 1872:625 ISS
             Syn.: Lepidocyrtus albinos(sic) Packard, AS, 1889:625 ISS
            Syn.: Lepidocyrtus marmoratus Packard, AS, 1873:36
            Syn.: Heteromurus albus Joseph, 1881
            Syn.: Tomocerus niveus Joseph, 1882,sd
            Syn.: Tomocerus Doderii(sic) Parona, 1888,sd
             Syn.: ... doderoi(sic) Gisin, 1960 ISS
            Syn.: Templetonia americana Harvey, 1892
            Syn.: Sinella cavernicola Carpenter, 1895
            Syn.: Templetonia quadrioculata Schött, 1896
             Syn.: Heteromurus 4-oculatus(sic) Delamare Deboutteville, C in Paulian, R, 1952:68 ISS
            Syn.: Heteromurus hirsutus Absolon, K, 1899:495
            Syn.: Heteromurus margaritatus Carpenter, 1902
            Syn.: Heteromurus nitidus var. paucodentatus(sic) Stach, J, 1930:271 IOS
            Syn.: Heteromurus nitidus var. paucidentatus Stach, J, 1930:295
            Syn.: Heteromurus gradgensis Denis, 1933
            Syn.: Heteromurus nitidus ssp. albicans Durkop, 1934
            Syn.: Heteromurus nitidus ssp. brevicornis Bonet, F, 1933:180
            Syn.: Heteromurus nitidus ssp. brevicornis Mari Mutt, JA et Bellinger, PF, 1990:91
            Syn.: Heteromurus nitidus f. quadriocellata(sic) Kseneman, M, 1935:14 IOS
             Syn.: Heteromurus nitidus var. quadriocellata(sic) Gisin, H, 1960:239 ISS
             Syn.: Heteromurus (Heteromurus) nitidus var. quadriocellata(sic) Nosek, J et Paoletti, MG, 1981:68 ISS
             Syn.: Heteromurus (Heteromurus) nitidus var. quadriocellata(sic) Nosek, J et Paoletti, MG, 1981:70 ISS
             Syn.: Heteromurus nitidus var. quadriocellata(sic) Nosek, J et Paoletti, MG, 1981:70 ISS
             Syn.: Pseudosinella Kemneri(sic) Agrell, I, 1936:27 ISS
            Syn.: Pseudosinella Kemneri(sic) var. tridentata Agrell, I, 1936:29
            Syn.: Propemesira duo-oculata(sic) Salmon, JT, 1942:59 IOS

Propemesira duo-oculata(sic) from New Zealand
After Salmon JT, 1942 Plate19 Fig.18

Propemesira duo-oculata(sic) from New Zealand
Antenna
After Salmon JT, 1942 Plate19 Fig.16

             Syn.: Propemesira duo-oculata(sic) Salmon, JT, 1945:71 ISS
             Syn.: Propemesira duoculata(sic) Mari Mutt, JA, 1980:37 ISS
            Syn.: Heteromurus nitidus ssp. armapollerae Massera, 1952
            Syn.: Heteromurus nitidus var. callaticola Gruia, 1965
            Syn.: Heteromurus tetrophthalmus Mari Mutt, JA, 1980:36
            Syn.: Pseudosinella suboculata Arbea, JI, 2017:408 partim
            Syn.: Desoria? sp. Wynne, JJ, Boyero Gallardo, JR, Hershauer, S, Huerta, OT, Martín, RF, Cuenca Rodriguez, J, París, M et Pardo-Igúzquiza, E, 2018:3

Desoria(?) sp. from Spain
After Wynne JJ & al, 2018 Fig.1C

             Syn.: Podura Nitida(sic) Bourlet, C, 1845:160 ISS
             Syn.: Isotoma Nitidum(sic) Kolenati, FA, 1858:133 ISS
            Syn.: Heteromurus nitidus Stach, J, 1924:109
             Syn.: Heteromurus nitida(sic) Salmon, JT, 1945:70 ISS
             Syn.: Heteromurus nididus(sic) Godissart, J et Delhez, F, 1969:38,B ISS
             Syn.: Isotoma nitidens(sic) Mari Mutt, JA, 1980:36 ISS
             Syn.: Heteromurus nitida(sic) Poole in Poole & Gentili, 1997 ISS

Heteromurus nitidus from Belgium
2004 © Vuijlsteke, M.

Heteromurus nitidus from Belgium
2006 © Vuijlsteke, M.

Heteromurus nitidus from Germany
2006.12.26 © Kohl, F.

Heteromurus nitidus from Germany
2006.12.26 © Kohl, F.

Heteromurus nitidus from the UK
2007.11.10 © Valentine., B

Heteromurus nitidus from the UK
2008.01.17 © Valentine., B

Heteromurus nitidus from the UK
2008.01.17 © Valentine., B

Heteromurus nitidus from the UK
2008.04.09 © Spears, G.

Heteromurus nitidus from Belgium
2008.06.09 © Vuijlsteke, M.

Heteromurus nitidus from the UK
2008.11.15 © Valentine., B

Heteromurus nitidus from the UK
2008.11.19 © Valentine., B

Heteromurus nitidus from the UK
2009.09.26 © Valentine., B

Heteromurus nitidus from the UK
2011.02.26 © Valentine., B

Heteromurus nitidus from the UK
2012.06.12 © Murray, A.

Heteromurus nitidus from the UK
Small reddish eyepatch
2012.07.18 © Jones, A.

Heteromurus nitidus from the UK
Antenna
2012.07.30 © Murray, A.

Heteromurus nitidus from the UK
2012.08.23 © Murray, A.

Heteromurus nitidus from the UK
2012.09.12 © Jones, A.

Heteromurus nitidus from the UK
Ribbed scales
2012.12.20 © Prior, F.

Heteromurus nitidus from Slovakia
Annulated apical antennal segment
2013.11.05 © Rahmé, N.

Heteromurus nitidus from Spain
Ant.1 subdivided; reddish eyes
2015.06.30 © Miralles, A.

Heteromurus nitidus from Holland
Predated by Acari
2015.10.11 © Heemskerk, R.

Heteromurus nitidus from the USA
Ribbed scales
2016.07.07 © Raymond-Léonard, J.L.

Heteromurus nitidus from the USA
Mucro with back-tilted subapical tooth, and basal spine
2016.07.08 © Raymond-Léonard, J.L.

Heteromurus nitidus from Belgium
2016.12.04 © Huskens, M.-L.

Heteromurus nitidus from Holland
Ant.1 subdivided (*)
2017.07.17 © Kamsteeg, G.

Heteromurus nitidus from the UK
Mucro with back-tilted subapical tooth, and basal spine
2019.02.05 © Cann, A.J.

Heteromurus nitidus from Holland
Ant.1 subdivided in short basal (1a) and long apical (1b) part
2019.10.22 © Kamsteeg, G.

Heteromurus nitidus juv. from Holland
2020.09.28 © Kamsteeg, G.

Species noseki Mari Mutt, JA et Stomp, N, 1980:186

Heteromurus noseki from Portugal
After Mari Mutt JA & Stomp N, 1980 Fig.3C

Species peyerimhoffi Denis, 1937

Species pulcher (Koch, CL & Berendt, GC in Berendt, GC, 1854:119) -

Podura pulchra from Baltic amber
After Koch CL & Berendt GC, 1845 Tab.XV Fig.128

Species sexoculatus Brown, JM, 1926:213

Syn.: Heteromurus hexophthalmus Denis, 1948

Species sinensis Liu & Li, 1999

Species tetrophthalmus Börner, 1903

Syn.: Heteromurus nitidus Zoughailech, A, 2017:92,cf

Heteromurus cf. nitidus from Algeria
After Zoughailech A, 2017 Fig.48

Heteromurus cf. nitidus from Algeria
2+2 ocelli
After Zoughailech A, 2017 Fig.48

Syn.: Heteromurus tatraphthalmus(sic) Denis, 1930 ISS

Heteromurus tetrophthalmus from Algeria
2+2 ocelli
After Bendjaballah M & al, 2018 Fig.19

Species uzicensis (Lucic & Curcic, 2002) Lucic & al., 2002

Species variabilis Martynova in Martynova, Tshelnokov & Rasulova, 1974

Syn.: Heteromurus variabilis ssp. martynova(sic) Nguyen, TT, 1995:564 lapsus

Heteromurus (Heteromurus) variabilis
After Mari Mutt, J.A., 1980 Fig.9

Subgenus Verhoeffiella Absolon, K, 1900:429 Syn.: Entomobryinae Typhlopodura Absolon, K, 1900:428

Verhoeffiella anagastumensis from Croatia
2010.11.19 © Lukic, M.

  Syn.: Heteromurus Verhoeffiela(sic) Bellini, BC, de Souza, PGC et Greenslade, P, 2021:2 ISS
  Syn.: Heteromurus Verhoeffiela(sic) Bellini, BC, de Souza, PGC et Greenslade, P, 2021:27 ISS
  Syn.: Heteromurus Verhoeffiela(sic) Bellini, BC, de Souza, PGC et Greenslade, P, 2021:28 ISS

Species absoloni (Kseneman, 1938) Gisin, H, 1960:237

Species anagastumensis Lucic LR in Lucic, LR, Tomic, VT, Brajkovic, MM et Curcic, SB, 2008:298

Heteromurus (Verhoeffiella) anagastumensis from Montenegro
After Lucic LR et al, 2008 Fig.1

Verhoeffiella anagastumensis from Croatia
After Lukic M & al, 2019 Fig.2up

Species cavicola (Absolon, K, 1900:430) Handschin, E in Dahl, F, 1929:93, t.t.

Verhoeffiella cavicola from Bosnia and Herzegovina
After Lukic M in Lukic M & al, 2018 Fig.1D

Verhoeffiella cavicola from Bosnia and Herzegovina
After Lukic M in Lukic M & al, 2018 Fig.2D

Species constantinellus Curcic BPM et Lucic LR in Lucic, LR, Curcic, BPM et Mitic, BM, 2007:67P

Species constantius Curcic BPM et Lucic LR in Lucic, LR, Curcic, BPM et Tomic, VT, 2007:72P

Heteromurus (Verhoeffiella) constantius from Herzegovina
After Lucic LR & al, 2007 Fig.1

Species dallaii Nosek, J et Paoletti, MG, 1985:271

            Syn.: Heteromurus (Verhoeffiella) dallaii Nosek, J et Paoletti, MG, 1981:70 nn
            Syn.: Verhoeffiella dallaii Nosek, J et Paoletti, MG, 1981:70 nn
             Syn.: Verhoeffiella dallai(sic) Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:155 ISS

Species gamae Lukic, Porco, Bedos & Deharveng, 2015

Species hispanicus Bonet, F, 1931:354

            Syn.: Heteromurus (Verhoeffiella) hispanica(sic) Bonet, F, 1931:354 IOS

             Syn.: Heteromurus (Verhoeffiella) hispanica(sic) Lucic, LR, Curcic, BPM et Tomic, VT, 2007:71P ISS
             Syn.: Heteromurus (Verhoeffiella) hispanica(sic) Lucic, LR, Tomic, VT, Brajkovic, MM et Curcic, SB, 2008:298 ISS

Species longicornis (Absolon, K, 1900:428) Gisin, H, 1960:237

Heteromurus (Verhoeffiella) longicornis from Austria
After Absolon K, 1900

Verhoeffiella longicornis from Croatia
2010.11.19 © Bedek, J.

Verhoeffiella longicornis from Austria
After Absolon K, 1900 in Cipola NG & al, 2016 Fig.2

Verhoeffiella longicornis from Bosnia and Herzegovina
After Lukic M in Lukic M & al, 2018 Fig.1A

Verhoeffiella longicornis from Bosnia and Herzegovina
After Lukic M in Lukic M & al, 2018 Fig.2A

Verhoeffiella longicornis from Croatia
Predated by Stalagtia hercegovinenensis
After Bedek J in Lukic M, 2019 Fig.1C

Species margusi Lucic M et Deharveng L in Lukic, M, Delic, T, Zagmajster, M et Deharveng, L, 2018:1145

Verhoeffiella margusi from Croatia
After Lukic M in Lukic M & al, 2018 Fig.1F

Verhoeffiella margusi from Croatia
After Lukic M in Lukic M & al, 2018 Fig.2F

Syn.: Verhoeffiella sp. Margus & al, 2012
Syn.: Verhoeffiella sp. Margus & al, 2013

Species media (Loksa & Bogojevic, 1967) Mari Mutt, JA, 1985:197

Syn.: Verhoeffiella medius(sic) Lukic, M, Delic, T, Zagmajster, M et Deharveng, L, 2018:1146 ISS
Syn.: Verhoeffiella medius(sic) Lukic, M, Delic, T, Zagmajster, M et Deharveng, L, 2018:1147 ISS

Species milosi Lucic M et Deharveng L in Lukic, M, Delic, T, Zagmajster, M et Deharveng, L, 2018:1153

Verhoeffiella milosi from Montenegro
After Lukic M in Lukic M & al, 2018 Fig.1E

Verhoeffiella milosi from Montenegro
After Lukic M in Lukic M & al, 2018 Fig.2E

Species polychaetosa Lucic M et Deharveng L in Lukic, M, Delic, T, Zagmajster, M et Deharveng, L, 2018:1155

Verhoeffiella polychaetosa from Bosnia and Herzegovina
After Lukic M in Lukic M & al, 2018 Fig.1B

Verhoeffiella polychaetosa from Bosnia and Herzegovina
After Lukic M in Lukic M & al, 2018 Fig.2B

Species verdemontana Lucic M et Deharveng L in Lukic, M, Delic, T, Zagmajster, M et Deharveng, L, 2018:1151

Verhoeffiella verdemontana from Bosnia and Herzegovina
After Lukic M in Lukic M & al, 2018 Fig.1C

Verhoeffiella verdemontana from Bosnia and Herzegovina
After Lukic M in Lukic M & al, 2018 Fig.2C

Genus Alloscopus Börner, C, 1906:177

Syn.: ... Indoscopus Prabhoo, 1971

Species arborealis Alviola, MS, Lucañas, CC, Jit Jr, IL, Soto-Adames, FN et Jantarit, S, 2024:281

Alloscopus arborealis from the Philippines
After Alviola MS & al, 2024 Fig.cover

Species aspinosus (Prabhoo, 1971) Yoshii, R & Suhardjono, YR, 1989:64

Species bannaensis Zhang F in Zhang, F, Cipola, NG, Pan, Z-X et Ding, Y-H, 2020:7

Alloscopus bannaensis from China
After Zhang F in Zhang F & al, 2020 Fig.3

Species deharvengi (Mari Mutt, JA, 1985:338) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:160

Syn.: Alloscopus tetracanthus Yosii, 1959 =?

Species fallax Yoshii, R et Suhardjno, YR, 1992:35

Species liuae Zhang F in Zhang, F, Cipola, NG, Pan, Z-X et Ding, Y-H, 2020:10

Species multispinatus (Mari Mutt, JA, 1982:86) Yoshii, R & Suhardjno, YR, 1992:34

Syn.: Heteromurus (Alloscopus) tenuicornis Handschin, 1925 =?

Species spinosus (Prabhoo, 1971) Yoshii, R & Suhardjono, YR, 1989:64

Species strebeli Winter, C, 1966:165

Syn.: Alloscopus strebeli Winter, C, 1963:509 nn

Species tenuicornis (Börner, C, 1906:177) Handschin, 1928, t.t.

            Syn.: Templetonia sp. Oudemans, 1890 =?
            Syn.: Alloscopus tetracanthus Yosii, 1960 =?
            Syn.: Alloscopus tetracanthus Gapud, 1971
            Syn.: Heteromurus yosiius Yoshii, R et Suhardjono, YR, 1989:64

            Syn.: Heteromurus (Alloscopus) tenuicornis Stach, J, 1924:109

Species tetracanthus (Börner, C, 1906:177) Handschin, 1928

            Syn.: Heteromurus (Alloscopus) tetracantha(sic) Börner, C, 1906:177 IOS
            Syn.: Templetonia sp. Oudemans, JT in Weber, M, 1890:88

             Syn.: Heteromurus tetracantha(sic) Imms, 1912 ISS
             Syn.: Heteromurus (Alloscopus) tetracantha(sic) Stach, J, 1924:109 ISS
             Syn.: Alloscopus tetracantha(sic) Winter, C, 1966:166 ISS
             Syn.: Alloscopus tetracantha(sic) Winter, C, 1966:168 ISS
             Syn.: Alloscopus tetracantha(sic) Prabhoo, 1969 ISS
             Syn.: Alloscopus tetracantha(sic) Yoshii & Yayuk, 1989 ISS
             Syn.: Heteromurus (Alloscopus) tetracantha(sic) Mandal, GP et Hazra, AK, 2009:68 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:68 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:72 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:76 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:80 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:115 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:125 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:127 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:128 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:205 ISS

Alloscopus tetracantha(sic) from India
After Mandal GP & Hazra AK, 2009 Fig.75a

Syn.: Alloscopus tetrachantus(sic) Bellini, BC, Greenslade, P, Baquero, E, Jordana, R, de Souza, PGC, 2022:221 ISS

Species thailandensis (Mari Mutt, JA, 1985:336) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:161

Species yosiius (Mari Mutt, JA, 1985:336) Yayuk, RS (sic), 1989:19

Syn.: Alloscopus tenuicornis Yoshii, R et Suhardjno, YR, 1992:35

Genus Heteromurtrella Mari Mutt, JA, 1979:214

Syn.: Heteromurini Alloscopus Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:157 lapsus

Syn.: Collembola Heteromurustrella(sic) Nguyen, TT, 1995:563 ISS

Heteromurtrella sp. from the Philipinnes
2009.03.26 © Tamayo, A.

Heteromurtrella sp. from the Philipinnes
2009.03.26 © Tamayo, A.

Heteromurtrella sp. from the Philipinnes
2009.03.26 © Tamayo, A.

Syn.: Heteromurini Heterumurtrella(sic) Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:165 ISS
Syn.: Heteromurini Heteromutrella(sic) Mandal, GP, 2018:83 ISS

Heteromurtrella sp. from Taiwan
2021.12.07 © Cheng, H.-J.

Heteromurtrella sp. from Taiwan
2021.12.07 © Cheng, H.-J.

Heteromurtrella sp. from Taiwan
2021.12.08 © Cheng, H.-J.

Heteromurtrella sp. from Taiwan
Descaled
2021.12.09 © Cheng, H.-J.

Heteromurtrella sp. from Taiwan
Dental spines absent
2021.12.09 © Cheng, H.-J.

Species abrupta (Salmon, 1951) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:165

Syn.: Heteromurus? sp. Yayuk, RS(sic), 1989:19

Species affinis (Mari Mutt, JA, 1987:575) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:166

Species anae Cipola NG in Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:175

Heteromurtrella anae from Brazil
After Cipola NG in Cipola NG & al, 2016 Fig.22

Species barrai (Mari Mutt, JA, 1979:221) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:166

Syn.: Heteromurus mirificus Barra, 1968 nec Salmon, 1954

Species caeca (Najt, J, Thibaud, J-M & Mari Mutt, JA, 1988:560) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:166

Species diommata (Denis, 1931) Palacios-Vargas, 1992

Species dubia (Delamare Deboutteville, C in Paulian, R, 1952:68) Castaño-Meneses, G, Palacios-Vargas, JG & Carmo, AFR, 2015:392

Heteromurus dubius from Ivory Coast
After Delamare Deboutteville C, 1952 Fig.22

Heteromurus dubius from Ivory Coast
Head
After Delamare Deboutteville C, 1952 Fig.24

            Syn.: Heteromurus dubius Delamare Deboutteville, 1948a nn
            Syn.: Heteromurus dubius Delamare Deboutteville, 1948b nn
            Syn.: Heteromurus dubius Delamare Deboutteville, 1951a nn
            Syn.: Heteromurus dubius Delamare Deboutteville, 1951b nn

             Syn.: Ptenura dubius(sic) Salmon, JT, 1964:479 ISS

Species echinata (Mari Mutt, JA, 1983:158) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:165

Syn.: Heteromurtrella echinatus(sic) Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:167 ISS

Species greensladeae (Mari Mutt, JA, 1987:572) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:167

Species livida (Salmon, 1954) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:172

Species mirifica (Salmon, 1954) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:172

Species nitens (Yosii, R, 1964:14) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:172

Heteromurus (Heteromurtrella) nitens from the Galagapos
After Katz AD et al, 2016 Fig.5A

Species pruinosa (Mari Mutt, JA, 1983:163) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:172

Species puertoricensis (Mari Mutt, JA, 1979:220) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:173, t.t.

Heteromurtrella puertoricensis from Puerto Rico
After Mari Mutt JA, 1979 in Cipola NG & al, 2016 Fig.3

Species schoetti (Denis, 1931) Palacios-Vargas, 1992

            Syn.: Heteromurus schoetti f. tridentata Denis, 1931
            Syn.: Heteromurus schoetti f. quadridentata Denis, 1931
            Syn.: Heteromurus bidentatus Denis, 1931
            Syn.: Heteromurus nitidus Winter, C, 1963:509

             Syn.: Heteromurus Schötti(sic) Denis, J-R, 1933:254 ISS
             Syn.: Heteromurus Schötti(sic) Denis, J-R, 1933:319 ISS
             Syn.: Heteromurus Schötti(sic) Delamare Deboutteville, C in Paulian, R, 1952:68 ISS

Species similis (Barra, 1968) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:173

Species stannardi (Mari Mutt, JA, 1976:326) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:173

Species subdubia (Barra, 1968) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:174

Syn.: Heteromurus subduvius(sic) Mari Mutt, JA, 1976:326 ISS

Species tihuiensis (Mari Mutt, JA, 1985:145) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:174

Species zairensis (Chelnokov, 1977) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:174

Tribus Mastigocerini Mari Mutt, 1980

Genus Mastigoceras Handschin, 1924

Species camponoti Handschin, 1924, t.t.

Mastigoceras camponoti from Brazil
After Cipola NG & al, 2019 Fig.5

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