http://www.collembola.org/taxa/hypogast.htm - Last updated on 2022.02.20 by Frans Janssens
Checklist of the Collembola: Hypogastruridae

Familia Hypogastruridae Börner, 1906 key

Genus Number
of species
Acherongia4
Acherontides12
Acherontiella20
Acheroxenylla4
Austrogastrura3
Barbagastrura1
Biscoia1
Bonetogastrura7
Celegastrura1
Ceratophysella142
Choreutinula7
Cosberella9
Denigastrura1
Ecuadogastrura1
Gnathogastrura1
Gomphiocephalus1
Hypogastrura (Franzura)1
Hypogastrura (Hypogastrura)174
Hypogastrura175
Jacutogastrura1
Mesachorutes3
Mesogastrura5
Microgastrura8
Neobeckerella1
Octoacanthella1
Ongulogastrura1
Orogastrura8
Parawillemia1
Paraxenylla11
Pseudacherontides8
Schaefferia37
Schoettella (Knowltonella)1
Schoettella (Schoettella)10
Schoettella11
Stenogastrura1
Tafallia2
Taurogastrura1
Thibaudylla4
Triacanthella24
Typhlogastrura19
Willemgastrura1
Willemia48
Xenylla146
Xenyllogastrura8
Hypogastruridae740
 Genera with number of species
  Syn.: Aphoruridae Podurinae Scherbakov, AM, 1898:4
  Syn.: ... Achorutinae Börner, 1901 r
  Syn.: Achorutidae Achorutini(sic) Börner, C, 1901:10 IOS
  Syn.: Collembola Neogastruridae Stach, J, 1949:5
Remark by Melville, R.V. & Smith, J.D.D. 1987:23: Neogastruridae Stach, 1949. Placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology. Type genus: Neogastrura Stach, 1949. A junior objective synonym of Hypogastrurinae Börner, 1906, the respective type genera of these two families having the same nominal species as type species. Opinion 435.

  Syn.: Collembola Podurhippidae Salmon, JT, 1949:5


Remark by Melville, R.V. & Smith, J.D.D. 1987:18: Hypogastrurinae Börner, 1906. Placed on the Official List of Family-Group Names in Zoology. Type genus: Hypogastrura Bourlet, 1839. Opinion 435.
  Syn.: Collembola Hypogastrurid'(sic) Dethier, M, 1998:55 ISS
Hypogastrura sp. from the USA
Mass aggregation
2008.02.12 © Boeddeker, M.
Hypogastruridae from Peru
2009.01-03.dd © Damiano Palomino., B
Hypogastrura sp. from the USA
With uplifted abdomen and extended furcula
2009.02.26 © Cowen, R.

Remark by Greenslade, P., Stevens, M.I., Torricelli, G. & D'Haese, C.A. 2011:227: Nonmonophyly of Hypogastruridae is consistent with the results of D'Haese (2002).
Remark by Janssens, F. 2012.11.15: To partially resolve the paraphyly of the Hypogastruridae (see Greenslade & al., 2011), we tentatively propose to restore the Poduroidea as sister-group of the Neanuroidea and as follows = ((Podura+(Biscoia+Microgastrura))+(Paraxenylla+Xenylla)). Note: this is not yet reflected in the current taxonomic hierarchy.
Hypogastruridae from the USA
Short furcula with broad manubrium
2012.03.21 © Kouri, J.


  Syn.: Collembola Hypogatruridae(sic) de Lima, ECA, Stievano, LC et Zeppelini, D, 2019:142 ISS
  Syn.: Poduromorpha Hypogasturidae(sic) Cucini, C, Fanciulli, PP, Frati, F, Convey, P, Nardi, F et Carapelli, A, 2021:12 ISS
  Syn.: Collembola Hypogasturidae(sic) Palacios-Vargas, JG, Castaño-Meneses, G et Valladares, L, 2021:149 ISS
  Syn.: Collembola Hypogasturidae(sic) Palacios-Vargas, JG, Castaño-Meneses, G et Valladares, L, 2021:150 ISS
  Syn.: Collembola Hypogasturidae(sic) Palacios-Vargas, JG, Castaño-Meneses, G et Valladares, L, 2021:153 ISS





Genus Triacanthella Schäffer, 1897

F
  Syn.: Achorutidae Triacanthurus Willem, V, 1902:5
  Syn.: Hypogastrurinae Triacanthura(sic) Womersley, H, 1939:95 ISS
  Syn.: Hypogastrurinae Clingmantha Wray, DL, 1963:101


  Syn.: Collembola Tricanthella(sic) Stach, J, 1967:393 ISS
  Syn.: Collembola Tricanthella(sic) Ellis, WN et Bellinger, PF, 1973:56 ISS
  Syn.: Hypogastruridae Triacanthela(sic) Camousseight, A, 1999:3 ISS
Remark by D'Haese, C.A. 2002:1149: Triacanthella is more closely related to Odontellidae and Onychiuridae [then to Hypogastruridae].
Triacanthella sp. from Australia
2010 © Stevens, M.
Remark by Janssens, F. 2012.11.15: According to Greenslade & al. (2011), Hypogastruridae is not monophyletic and Triacanthella is the most basic poduromorph genus and sister-group of all other Poduromorpha. Therefore, to partially resolve the paraphyly of the Hypogastruridae, we tentatively propose to place Triacanthella in a monogenic family Triacanthellidae basic to all Poduromorpha. Note: this is not yet reflected in the current taxonomic hierarchy.
Remark by Janion, C., D'Haese, C.A. & Deharveng, L. 2012:58: The genus Triacanthella is phyletically isolated among Hypogastruridae both from a morphological and a molecular point of view.

Triacanthella purpurea from New Zealand
Ab.6 with 2 dorsal anal spines (das) and 1 terminal anal spine (tas)
2014.02.16 © Murray, A.
Triacanthella sp. from Tasmania
2016.09.16 © Tony.
Triacanthella sp. from Tasmania
2+1 anal spines
2016.09.16 © Tony.



Species sp. Fjellberg, 1999

Species alba Carpenter, 1909

Species andina Cassagnau & Rapoport, 1962

Species biroi Stach, J, 1924:109

            Syn.: Triacanthella Birói(sic) Stach, J, 1924:109 IOS
Triacanthella Birói(sic) from Malta
After Stach J, 1924 Taf.I Fig.15

            Syn.: Triacanthella inopinata Denis, 1924
            Syn.: Triacanthella gridellii Denis, 1938



             Syn.: Triacanthella birói(sic) Salmon, JT, 1964:241 ISS
             Syn.: Triacanthella Biroi(sic) Pfliegler, WP et Janssens, F, 2016:204 ISS
  Subspecies monacanthella Stach, J, 1924:111





Species clavata (Willem, V, 1902:5) Carpenter, 1909

Triacanthurus clavatus from Antarctica
After Willem V, 1902 Pl.I Fig.7
            Syn.: Triacanthura(sic) clavata Womersley, H, 1939:95


Species copelandi (Wray, DL, 1963:101) Christiansen, K & Bellinger, P, 1980:203

Species enderbyensis Salmon, JT, 1949:6

Triacanthella enderbeyensis from New Zealand
2009 © Te Papa Museum.

Species frigida Cassagnau, P, 1959:71

Species intermedia Dunger & Zivadinovic, 1983

Species madibai Janion, C, D'Haese, CA et Deharveng, L, 2012:60

            Syn.: Triacanthella madiba(sic) Janion, C, D'Haese, CA et Deharveng, L, 2012:60 IOS
Triacanthella madiba(sic) from South Africa
After Janion C & al, 2012 Fig.1
Triacanthella madiba(sic) from South Africa
Plurichaetosis and heterochaetosis
After Janion C & al, 2012 Fig.6A

            Syn.: Schaefferia (Typhlogastrura) sp. Sharratt & al, 2000 =?



Triacanthella madibai from South Africa
2011.05.16 © pgs.
Triacanthella madibai from South Africa
2011.05.16 © pgs.

             Syn.: Triacanthella madiba(sic) Thibaud, J-M, 2013:240 ISS
             Syn.: Triacanthella madiba(sic) Janion-Scheepers, C, Deharveng, L, Bedos, A et Chown, L, 2015:61 ISS




Species massoudi Najt, 1973

Species michaelseni Schäffer, 1897, t.t.

             Syn.: Triacanthella Michaelseni(sic) Stach, J, 1924:105 ISS
             Syn.: Triacanthella michaelsenii(sic) Jordana, R et Arbea, JI in Ramos, MA et al, 1997:231 ISS

Species najtae Izarra, 1971

Species nivalis Cassagnau & Deharveng, 1974

Species perfecta Denis, 1926

            Syn.: Triacanthella annae Börner, 1932 nn
            Syn.: Triacanthella proxima Delamare Debouttevile, 1951
            Syn.: Triacanthella proxima f. principalis Rapoport, EH, 1961:625
            Syn.: Triacanthella proxima var. montana Rapoport, EH, 1961:625
Triacanthella proxima var. montana from France
After Rapoport E-H, 1961 Fig.1

            Syn.: Triacanthella montana Cassagnau & Deharveng, 1974



Triacanthella perfecta from Spain
Pluri- & heterochaetosis
After Jordana R & 1Arbea JI, 1997 Fig.105A
Triacanthella perfecta from France
2017.10.31 © Tiky, S.
Triacanthella perfecta from France
Abd.6 with 3 anal spines
2017.10.31 © Tiky, S.
Triacanthella perfecta from France
Eyepatch with reduced posterior ocelli
2017.10.31 © Tiky, S.
Triacanthella perfecta from France
Pluri- & heterochaetosis
2018.11.18 © Tiky, S.
Triacanthella perfecta from France
2018.11.18 © Tiky, S.
Triacanthella perfecta from France
2020.12.23 © Reinach, Br.
Triacanthella perfecta from France
Eyepatch with smaller posterior ocelli
2020.12.23 © Reinach, Br.
Triacanthella perfecta from France
2021.07.12 © Picard, J.
Triacanthella perfecta from France
Abdomen in dorsal aspect
2021.07.12 © Picard, J.
Triacanthella perfecta from France
Abd.6 with 3 anal spines
2021.07.12 © Picard, J.
Triacanthella perfecta from France
Abd.6 with 3 anal spines
2021.07.12 © Picard, J.
Triacanthella perfecta from France
Head
2021.07.12 © Picard, J.

Species purpurea Salmon, JT, 1943:373

            Syn.: Triacanthella perfusa Stach, J, 1967:398 lapsus


Triacanthella purpurea from New Zealand
2009 © Te Papa Museum.
Triacanthella purpurea from New Zealand
2014.02.16 © Murray, A.
Triacanthella purpurea from New Zealand
Eyepatch with 8 ocelli of which some reduced in size
2014.02.16 © Murray, A.
Triacanthella purpurea from Tasmania
2014.03.19 © Murray, A.
Triacanthella purpurea from New Zealand
2016.02.27 © Navest, M.P.

Species rosea Wahlgren, E, 1906:7

             Syn.: Triacanthella rosae(sic) Janion, C, D'Haese, CA et Deharveng, L, 2012:59 ISS

Species rubra Salmon, JT, 1941:288

Triacanthella rubra from New Zealand
2009 © Te Papa Museum.

Species setacea Salmon, JT, 1941:289

Triacanthella setacea from New Zealand
After Salmon, J.T., 1941 Pl.39 Fig.1.
Triacanthella setacea from New Zealand
2009 © Te Papa Museum.

Species sorenseni Salmon, JT, 1949:7

             Syn.: Triacanthella serenseni(sic) Stach, J, 1967:398 ISS
Triacanthella sorenseni from Campbell Island
2009 © Te Papa Museum.



Species terrasilvatica Salmon, JT, 1943:374

            Syn.: Triacanthella transilvatica Stach, J, 1967:398 lapsus


             Syn.: Triacanthella terrasilvaticas(sic) Salmon, JT, 1943:373 ISS
Triacanthella terrasilvatica from New Zealand
2009 © Te Papa Museum.



Species travei Cassagnau & Deharveng, 1974

Species violacea Womersley, H, 1939:97

Species vogeli Weiner & Najt, 1997


Genus Hypogastrura Bourlet, C, 1839:404 key  key  key  key

F
  Syn.: Poduradæ(sic) Achorutes Templeton, R in Templeton, R et Westwood, JO, 1836:96 partim
  Syn.: ... Achoreutes(sic) Templeton, 1842 ISS
  Syn.: ... Rathumoutes Templeton, 1842, partim
  Syn.: Podurelles Podurhippus Mégnin, JP, 1878:cxiv
  Syn.: Poduridæ(sic) Schoturus(sic) MacGillivray, AD, 1893:316 IOS
  Syn.: ... Euachorutes Börner, 1901, partim
  Syn.: Achorutidae Achorutini(sic) Börner, C, 1901:13 partim IOS
  Syn.: ... Neohypogastrura Paclt, 1944
  Syn.: ... Anachorutes Bagnall, 1949
  Syn.: ... Ancistracanthella Gisin, 1949
  Syn.: ... Neogastrura Stach, 1949
  Syn.: ... Biacanthella Scott, 1961


  Syn.: ... Hypogastrurus(sic) Gervais in Walckenaer, 1844 ISS
  Syn.: Podura Hypogastrurus(sic) Elditt, HL, 1854:44 ISS
  Syn.: Hypogastrurinae hypogastrura(sic) Denis, J-R, 1924:220 ISS
  Syn.: Collembola Hypogastura(sic) Kosswig, C, 1952:91 ISS
  Syn.: Collembola Hyp(sic) Yosii, R, 1961:16 ISS
Remark by ICZN. 2007:403: Placed on the Official List of Names in Zoology. Opinion 435.


  Syn.: Hypogastruridae Hypogastura(sic) Jordana, R, Baquero, E, Ledesma, E, Sendra, A et Ortuño, VM, 2020:89 ISS
Hypogastrura sp. from Holland
Furcula with broad trapezoid manubrium and short dentes
2020.03.07 © Kamsteeg, G.

Subgenus Hypogastrura Bourlet, C, 1842 ss



Hypogastrura sp. from Germany
2006.06.28 © Stegmann, T.
Hypogastrura sp. from Czechia
2007.01.28 © Krásenský, P.
Hypogastrura sp. from Estonia
2007.03.04 © Tartes, U.
Hypogastrura sp. from Estonia
2007.03.04 © Tartes, U.
Ceratophysella sp. from the USA
2007.03.18 © Durham, J.
Hypogastrura sp. from the USA
2007.12.19 © Shaul, A.
Hypogastrura sp. from the UK
2008.02.05 © Valentine., B
Hypogastrura sp. from the USA
2008.02.06 © McDonald, G.
Hypogastrura sp. from the USA
2008.02.12 © Boeddeker, M.
Hypogastrura sp. from Estonia
2008.05.13 © Tartes, U.
Hypogastrura sp. from the UK
2008.10.21 © Spears, G.
Hypogastrura sp. from Spain
2009.01.14 © Lluch, J.
Hypogastrura sp. from the USA
With uplifted thorax
2009.02.26 © Cowen, R.
Hypogastrura sp. from the USA
2009.03.05 © Cowen, R.
Hypogastrura sp. from the USA
2010.02.15 © Murray, T.
Hypogastrura sp. from the USA
Everted anal sacs
2012.03.09 © Holland, M.
Hypogastrura sp. from the UK
Buccal cone
2012.10.03 © Murray, A.
Hypogastrura sp. from the UK
Buccal cone
2013.01.04 © Murray, A.
Hypogastrura sp. from the UK
2013.01.11 © Holt, H. & A.
Hypogastrura sp. from the UK
Lobed post-annal organ (PAO)
2013.12.13 © Phillips, E.
Hypogastrura sp. from Germany
Eyepatch with 8 ocelli
2015.09.11 © Kluck, A.
Hypogastrura sp. from the UK
With uplifted abdomen
2016.03.06 © Holla, D.
Hypogastrura sp. from France
Eyepatch with 8 ocelli (5+3)
Post-antennal organ (PAO) with 4 lobes
2017.12.31 © Picard, J.
Hypogastrura sp. from Holland
2018.10.25 © Kamsteeg, G.
Hypogastrura sp. from the UK
Post-antennal organ (PAO) with lobes
2019.02.06 © Smart, J.
Hypogastrura sp. from Holland
2019.12.17 © Kamsteeg, G.
Hypogastrura sp. from Holland
Post-antennal organ with 4 lobes
2019.12.17 © Kamsteeg, G.
Hypogastrura sp. from Holland
Abd.6 with 2 indistinctly small anal spines
2019.12.17 © Kamsteeg, G.
Hypogastrura sp. from Holland
Small anal spine on papilla
2019.12.17 © Kamsteeg, G.
Hypogastrura sp. from Holland
Small anal spine on papilla (*)
2019.12.22 © Kamsteeg, G.
Hypogastrura sp. from France
Short furca with broad manubrium
2020.12.07 © Balti, K.
Hypogastrura sp. from France
Eyepatch with 8 ocelli
2020.12.07 © Balti, K.

Species-group viatica
  Syn.: Hypogastrura viatica Christiansen, K et Bellinger, P, 1980:52,gr




Species aequepilosa (Stach, J, 1949:96) Cassagnau, P, 1953:227

            Syn.: Hypogastrura tullbergi Weiner, 1976


             Syn.: Hypogastrura (Hypogastrura) aequipilosa(sic) Gisin, H, 1960:43 ISS
             Syn.: Hypogastrura æquepilosa(sic) Gisin, H, 1963:96 ISS
             Syn.: Hypogastrura aequipilosa(sic) Weiner, WM, 1981:424 ISS
             Syn.: Hypogastrura aequipilosa(sic) Szeptycki, A et Weiner, WM in Razowski J, 1990:19 ISS

Species affinis (Lucas, H, 1846:256) Börner in Voeltzkow, 1907

            Syn.: Achorutes similis Nicolet, H, 1847:379
Achorutes similis from Chile
After Nicolet H in Gay C, 1854 Lam.1 Fig.5.

            Syn.: Proxenyllodes aberrans Cassagnau, P, 1952:309
            Syn.: Hypogastrura boldorii Cassagnau, 1955
            Syn.: Hypogastrura boldorii Cassagnau, 1956
            Syn.: Hypogastrura boldorii Gisin, 1961
            Syn.: Hypogastrura boldorii Cassagnau, 1986
            Syn.: Hypogastrura boldorii Najt, Dalens & Deharveng, 1984
            Syn.: Hypogastrura tullbergi Gisin, 1961
            Syn.: Hypogastrura tullbergi Cassagnau, 1974



Achorutes affinis from Algeria
After Lucas H, 1849 Pl.2 Fig.10

Species alta Christiansen, K et Bellinger, P, 1980:109

Species antra Christiansen, K et Bellinger, P, 1980:110

Species barguzini Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:110

Species boldorii Denis, 1931

            Syn.: Hypogastrura elegantula Butschek, 1948
            Syn.: Hypogastrura aequepilosa Raynal, 1964


             Syn.: Hypogastrura Boldorii(sic) Denis, J-R, 1931:80 ISS
             Syn.: Hypogastrura Boldorii(sic) Travé, J, Gadea, E et Delamare Deboutteville, C, 1954:207 ISS
             Syn.: Hypogastrura Boldorii(sic) Travé, J, Gadea, E et Delamare Deboutteville, C, 1954:208 ISS

Species brevifurca Skarzynski, 2000

Species bulba Christiansen & Bellinger, 1980:112

Species burkilli (Bagnall, RS, 1940:167) Weidner, 1953, B

Hypogastrura burkilli from Holland
2013.10.05 © van Duinen, J.
Det.: Berg, M.
Hypogastrura burkilli from Holland
Tibiotarsus 2 with 3 knobbed tenent setae
2013.10.05 © van Duinen, J.
Hypogastrura burkilli from Holland
Tibiotarsus 3 with 3 knobbed tenent setae
2013.10.05 © van Duinen, J.

Species caduceator (Carpenter, 1919) Salmon, JT, 1964:211

Species campbelli Womersley, 1930

Species concolor (Carpenter, 1900) Schött, 1923

Species conflictiva Jordana, R et Arbea, JI, [1992]:59

Species distincta (Axelson, 1902) Linnaniemi, WM, 1912:30

            Syn.: Hypogastrura itaya Kinoshita, 1916


Hypogastrura disctincta from the UK
2004 © Hopkin, S.P.
Spatulate seta on 6th abdominal segment
Hypogastrura distincta from Sweden
2008.08.18 © Janssens, F.
Hypogastrura distincta from Sweden
Spatulate seta on 6th abdominal segment
2008.08.18 © Janssens, F.
Hypogastrura distincta from Sweden
Anal spine curved anteriad
2008.08.18 © Janssens, F.
Hypogastrura distincta from Sweden
Foot of 1st leg
Only 1 tenent hair out of 3 in focus
2008.08.18 © Janssens, F.
Hypogastrura distincta from Sweden
Short furcula with small mucrones
2008.08.18 © Janssens, F.
 
Hypogastrura distincta from Sweden
Apical organ of 4th antennal segment
2008.08.18 © Janssens, F.
Hypogastrura distincta from Sweden
Rami of retinaculum with 3 teeth
2008.08.18 © Janssens, F.
Hypogastrura distincta from Holland
2018.10.25 © van Duinen, J.
Hypogastrura distincta from Holland
2018.10.25 © van Duinen, J.
Hypogastrura distincta from Holland
Post-antennal organ (PAO) distinct
2018.10.25 © van Duinen, J.
Hypogastrura distincta from Holland
Ant.3 and ant.4 apparently fused
2018.10.25 © van Duinen, J.
Hypogastrura distincta from Holland
Furcula
Abd.6 with spatulate setae (*)
2018.10.27 © van Duinen, J.
Hypogastrura distincta from Holland
Mouth roundish
2018.10.27 © van Duinen, J.
Hypogastrura distincta from Holland
Post-antennal organ (PAO) with 4 lobes
2018.10.27 © van Duinen, J.
Hypogastrura distincta from Holland
Abd.6 with 2 small anal spines (*)
2018.10.27 © van Duinen, J.
Hypogastrura distincta from Holland
Proleg with 3 tenent setae
2018.10.28 © van Duinen, J.

Species gracilis (Folsom, JW, 1899:263) Yosii, 1960

Achorutes gracilis from Japan
After Folsom JW, 1899 Pl.1 Fig.6

Species helena Christiansen & Bellinger, 1980:118

Species humi (Folsom, 1916:489) Snider, RJ, 1967:197

            Syn.: Achorutes gami Wray, 1952


Species leo Palacios-Vargas, JG, 1986:41

Species litoralis (Linnaniemi, 1909) Handschin, E in Dahl, F, 1929:25

            Syn.: Achorutes propinquus Carpenter & Evans, 1904
            Syn.: Achorutes purpurascens var. galiciana Börner, 1906
            Syn.: Achorutes purpurascens Womersley, 1928, nec Lubbock, 1867
             Syn.: Achorutes purpurescens(sic) Stach, J, 1949:100 nec Lubbock, 1867 ISS
            Syn.: Achorutes browni Bagnall, RS, 1940:166
             Syn.: Achorutes Browni(sic) Stach, J, 1949:45 ISS
            Syn.: Hypogastrura tullbergi Hammer, 1954
            Syn.: Hypogastrura subboldorii Delamare Deboutteville, C et Jacquemart, S, 1962:667
            Syn.: Hypogastrura concolor Pomorski & Skarzynski, 1995


Hypogastrura litoralis from the UK
Posterior abdomen without macrosetae; as=anal spines
2004 © Hopkin, S.P.

Species martiani Skarzynski, D et Kaprus, IJ, 2009:24

            Syn.: Hypogastrura tullbergi Cox, 1982

Species morbillata (Salmon, JT, 1941:292), i.s.

Achorutes morbillatus from New Zealand
After Salmon JT, 1941 Pl.41 Fig.30

Species omnigra (Salmon, JT, 1941:294), i.s.

Species perplexa Christiansen, K et Bellinger, P, 1980:124

            Syn.: Achorutes macgillivrayi Folsom, JW, 1916:488 nec Dalla Torre, 1895
            Syn.: Achorutes tullbergi Schäffer, 1900 =?


Species purpurescens (Lubbock, J, 1870:302) Wolf, B, 1938:109, B

Achorutes purpurescens from the UK
After Lubbock J, 1870 Pl.XXII Fig.24
Achorutes purpurescens from the UK
Anterior foot
After Lubbock J, 1870 Pl.XXII Fig.25
Achorutes purpurescens from the UK
Saltatory appendage
After Lubbock J, 1870 Pl.XXII Fig.26
            Syn.: Achorutes purpureus Nicolet, H, 1847:379
            Syn.: Achorutes manubrialis Carpenter & Evans, 1899, nec Tullberg, 1869
            Syn.: Achorutes affinis Schäffer, 1900, nec Lucas, 1846, partim
            Syn.: Achorutes Carolinae(sic) Schäffer, 1900 IOS
            Syn.: Achorutes (Euachorutes) purpurascens(sic) var. aurea Börner, C, 1901:30
            Syn.: Achorutes (Euachorutes) purpurascens(sic) var. inermis Börner, C, 1901:30
            Syn.: Achorutes purpurascens(sic) var. pallens Willem, V, 1902:278,B
            Syn.: Achorutes purpurescens ssp. galiciana Börner, 1903
            Syn.: Achorutes propinquus Carpenter & Evans, 1904
            Syn.: Hypogastrura purpurascens(sic) var. cunicularis Latzel, R, 1917:233
            Syn.: Achorutes Nicoleti(sic) Houlbert, C, 1924:75 IOS
            Syn.: Hypogastrura pseudo-purpurascens(sic) Womersley, 1928 IOS
             Syn.: Hypogastrura peseudopurpurascens(sic) Womersley, H, 1936:327 ISS
             Syn.: Achorutes purpurascens-pseudopurpurascens Bagnall, RS, 1940:163,gr ISS
            Syn.: Hypogastrura purpurascens(sic) var. trispina Handschin, E in Dahl, F, 1929:25
            Syn.: Hypogastrura burkilli Lawrence, 1962 =?
            Syn.: Hypogastrura (viatica-group) burkilli Skarzynski, D, 2009:il
            Syn.: Ancistracanthella simoneti Gisin, 1949



             Syn.: Achorutes purpurascens(sic) Lubbock, J, 1870:709 ISS
             Syn.: Achorutes purpurascens(sic) Tullberg, 1872 ISS
Achorutes purpurescens from the UK
After Lubbock, J., 1873 Pl.41 Fig.1
Achorutes purpurescens from the UK
After Lubbock, J., 1873 Pl.41 Fig.2


             Syn.: Achorutes purpurascens(sic)? Reuter, OM, 1876:83 ISS
             Syn.: Achorutes purpurascenti(sic) Reuter, OM, 1876:83 ISS
             Syn.: Achorutes purpurascens(sic) Schäffer, 1897 ISS
             Syn.: Achorutes purpurascens(sic) Folsom, JW, 1899:264 ISS
             Syn.: Achorutes purpurascens(sic) Absolon, K, 1901:380 ISS
             Syn.: Achorutes (Euachorutes) purpurascens(sic) Börner, C, 1901:9 ISS
             Syn.: Hypogastrura purpurascens(sic) Linnaniemi, 1911 ISS
             Syn.: Hypogastrura purpurascens(sic) Linnaniemi, WM, 1912:31 ISS
             Syn.: Hypogastrura purpurascens(sic) Latzel, R, 1917:233 ISS
             Syn.: Hypogastrura purpurascens(sic) Handschin, 1924 ISS
             Syn.: Hypogastrura purpurascens(sic) Handschin, 1929 ISS
             Syn.: Hypogastrura purpurascens(sic) Bonet, F, 1930:117 ISS
             Syn.: Hypogastrura purpurascens(sic) Marlier, GJ, 1942:8,B ISS
             Syn.: Achorutes (Achorutes) purpurascens(sic) Kloet, GS et Hincks, WD, 1945:3 ISS
             Syn.: Hypogastrura purpurascens(sic) Busvine, J, 1951:421 ISS
             Syn.: Hypogastrura purpurascens(sic) Cassagnau, P in Vandel, A, 1955:1483 ISS
             Syn.: Hypogastrura purpurascens(sic) Obenberger, 1955 ISS
             Syn.: Hypogastrura purpurascens(sic) Kühnelt, W, 1961:159 ISS
             Syn.: Hypogastrura purpurascens(sic) Busvine, JR, 1966:418 ISS
             Syn.: Hypogastrura purpurascens(sic) Delhez, F et Houssa, M, 1969:207,B ISS
             Syn.: Hypogastrura purpurascens(sic) Hubart, J-M, 1982:343,B ISS
             Syn.: Hypogastrura purpurascens(sic) Szeptycki & Weiner in Razowski, 1990 ISS
             Syn.: Hypogastrura purpurascens(sic) Poole in Poole & Gentili, 1997 ISS
             Syn.: Hypogastrura purpurascens(sic) Berg, M et Aptroot, A, 2003:108 ISS
             Syn.: Hypogastrura purpurascens(sic) Pfander, I et Zettel J, 2004:576 ISS
Hypogastrura purpurescens from the UK
2004 © Hopkin, S.P.
3rd tibiotarsus with 2 clavate tenent setae in 2 rows
Hypogastrura purpurescens from the UK
2007.02.13 © Jones, D.
Hypogastrura purpurescens from the UK
2008.01.22 © Farrell, S.
Hypogastrura purpurescens from the UK, juvenile
2008.02.12 © Farrell, S.
Hypogastrura purpurescens from the UK
2008.05.01 © Farrell, S.
Hypogastrura purpurescens from the UK
2013.09.28 © Jeffery, J.
Hypogastrura purpurescens from France
2018.18.11.03 © Tiky, S.
Hypogastrura purpurescens from the UK
2019.02.25 © Smart, J.
Hypogastrura purpurescens from Holland
2020.11.07 © Kamsteeg, G.





Species quadritenenta Jiang, J et Chen, J-X, 2008:?

Species sensilis (Folsom, JW, 1919:5A) Weber, 1949

            Syn.: Hypogastrura trybomi McAlpine, 1964 =?
            Syn.: Hypogastrura trybomi McAlpine, 1965 cf =?
            Syn.: Hypogastrura trybomi Danks & Byers, 1972 =?
            Syn.: Hypogastrura trybomi Addison, 1975 =?
            Syn.: Hypogastrura trybomi Ryan, 1977 cf =?
            Syn.: Hypogastrura trybomi Christiansen &Bellinger, 1980 =?
            Syn.: Hypogastrura trybomi Fjellberg, 1986 =?
            Syn.: Hypogastrura trybomi Christiansen &Bellinger, 1998 =?
            Syn.: Hypogastrura trybomi Babenko, 2018 =?


Species subboldorii Delamare Deboutteville, C et Jacquemart, S, 1962:667

Species tullbergi (Schäffer, 1900) Schött, 1923

            Syn.: Achorutes dubius Tullberg, T, 1877:38
            Syn.: Achorutes dubius var. concolor Carpenter, 1900
            Syn.: Hypogastrura aureus Börner, 1901
            Syn.: Achorutes purperescens f. litoralis Linnaniemi, 1907
            Syn.: Achorutes sensilis Folsom, 1919
            Syn.: Hypogastrura concolor var. setosa Schött, 1923
            Syn.: Hypogastrura purpurascens Womersley, 1928, nec Lubbock, 1867
            Syn.: Hypogastrura browni Gisin, H, 1960:43
            Syn.: Hypogastrura browni Lawrence, 1962 =?
            Syn.: Hypogastrura browni Raynal, 1964
            Syn.: Hypogastrura browni Stach, J, 1967:395
            Syn.: Hypogastrura browni Thibaud, J-M, Schulz, H-J et da Gama Assalino, MM, 2004:120
            Syn.: Proxenyllodes aberrans Cassagnau, P, 1952:309
             Syn.: Proxenyllodes aberans(sic) Stach, J, 1967:395 ISS
            Syn.: Hypogastrura boldorii Delamare Deboutteville, 1954
             Syn.: Hypogastrura Boldorii(sic) Cassagnau, P in Vandel, A, 1955:1483 ISS
             Syn.: Hypogastrura Boldorii(sic) Cassagnau, P in Vandel, A, 1956:603 ISS
             Syn.: Hypogastrura Boldorii(sic) Cassagnau, P in Vandel, A, 1956:1361 ISS
             Syn.: Hypogastrura Boldorii(sic) Cassagnau, P in Vandel, A, 1958:3379 ISS
             Syn.: Hypogastrura (Hypogastrura) aequipilosa(sic) Gisin, H, 1960:43 ISS
            Syn.: Hypogastrura aequepilosa Gisin, 1961
            Syn.: Hypogastrura aequepilosa Stach, J, 1967:395
            Syn.: Hypogastrura spitsbergensis Stach, 1962
             Syn.: Hypogastrura spitzbergensis(sic) Gisin, H, 1963:77 ISS
             Syn.: Hypogastrura spitzbergensis(sic) Gisin, H, 1963:96 ISS
            Syn.: Hypogastrura subboldorii Raynal, 1964
            Syn.: Hypogastrura elegantula Stach, J, 1967:394


             Syn.: Hypogastrura (Hypogastrura) tulbergi(sic) Leinaas, HP, 1988:278 ISS
Remark by Fjellberg, A. 1998:26: A true Arctic species. Records of 'tullbergi' from Continental Europe refer to other species.


             Syn.: Hypogastrura tullbergii(sic) Skarzynski, D, 2006:85 ISS
             Syn.: Hypogastrura tulbergi(sic) Fiera, C, 2007:8 ISS
             Syn.: Hypogastrura tullbergii(sic) Skarzynski, D et Kaprus, IJ, 2009:23 ISS

Species variata Babenko, A, Efeykin, B et Bizin, M, 2020:3

Hypogastrura variata from NE Russia
After Babenko A, Efeykin B & Bizin M, 2020 Fig.20
Hypogastrura variata from NE Russia
First instar
After Babenko A, Efeykin B & Bizin M, 2020 Fig.23

Species viatica (Tullberg, 1872:50) Carpenter, 1909, t.t., B

            Syn.: Podura obscure cærulea Olafsen & Povelsen, 1772
            Syn.: Podura humicola Fabricius O., 1780
            Syn.: Achorutes dubius Templeton, R in Templeton, R et Westwood, JO, 1836:96
Achorutes dubius from Ireland
After Templeton, R, 1836 Pl.XII Fig.5

            Syn.: Hypogastrura aquatica Bourlet, C, 1839:407
Hypogastrura aquatica from France
After Bourlet C, 1839 Fig.9
Ochorute(sic) aquatique from Europe
Achorutes aquaticus
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.4

             Syn.: Hypogastrura aquatic(sic) Budha, PB et Shrestha, P, 2021:26 ISS
            Syn.: Podura (pas de crochets à l'extrémité du corps) cellaris Nicolet, H, 1842:56
            Syn.: Podura (pas de crochets à l'extrémité du corps) cyanocephala Nicolet, H, 1842:56
            Syn.: Hypogastrura fuscoviridis Bourlet, 1842
            Syn.: Hypogastrura murorum Bourlet, 1842, IZCN Opinion 435
Achorutes murorum from Europe
After Nicolet H, 1847 Pl.6 fig.11

             Syn.: Achorutes muscorum(sic) Lucas, H, 1849:376 lapsus ISS
            Syn.: Achorutes murorum Parona, 1895 =?
            Syn.: Achorutes murorum Bonet, F, 1933:125
            Syn.: Podura (pas de crochets à l'extrémité du corps) similata Nicolet, H, 1842:56
Podura similata from Switzerland
After Nicolet H, 1842 Pl.5 fig.5.

             Syn.: Achorutes similutas(sic) Carl, J, 1899:274,sd ISS
            Syn.: Hypogastrura muralis Bourlet, 1843
            Syn.: Podura Nicoleti(sic) Nicolet, 1847
            Syn.: Achorutes viatica var. inermis Axelson, 1905
            Syn.: Achorutes viaticus var. maculosa Axelson, WM, 1905:788
            Syn.: Achorutes viaticus f. trispina Axelson, WM, 1905:788
            Syn.: Achorutes pseudoviaticus Bagnall, 1941
            Syn.: Achorutes subviaticus Bagnall, 1941
             Syn.: Achorutes sub-viaticus(sic) Busvine, J, 1951:422 ISS
            Syn.: Achorutes titahiensis Salmon, JT, 1943:376
            Syn.: Hypogastrura antarctica Salmon, JT, 1962:887



Hypogastrura viatica from the UK
2004 © Hopkin, S.P.
Distinct abdominal macro- and mesosetae
Hypogastrura viatica from Antarctica
After Davis, G. in Greenslade, P., 2006 Fig.56.
Hypogastrura viatica from Sweden
2006 © Hall, K.
Hypogastrura viatica from the UK
2009.10.31 © Talbot, M.E.
Det.: Shaw, P.
Hypogastrura viatica
After Thibaud J-M & D'Haese C, 2010 Fig.31.1
Hypogastrura viatica from the UK
2012.12.16 © Murray, A.
Hypogastrura viatica from the UK
Composed eye with 8 ocelli
2012.12.16 © Murray, A.
Hypogastrura viatica from the UK
Three clavate setae distally on 3rd leg
2012.12.16 © Murray, A.
Hypogastrura viatica from the UK
Second leg with 3 clavate setae
2012.12.15 © Phillips, E.

             Syn.: Hypogastrura Viatica(sic) Santeshwari, X, Raghuraman, M et Singh, J, 2015:572 ISS
Hypogastrura viatica from India
After Santeshwari, Raghuraman M & Singh J, 2015 p.573
Hypogastrura viatica from Holland
2017.08.04 © van Duinen, J.
Det.: Berg, M., 2017.08.14
Hypogastrura viatica from Holland
2017.08.04 © van Duinen, J.
Det.: Berg, M., 2017.08.14
Hypogastrura viatica from Holland
2017.08.04 © van Duinen, J.
Det.: Berg, M., 2017.08.14
Hypogastrura viatica from France
Unguiculus (u) with basal lamella
2018.02.18 © Picard, J.
Hypogastrura viatica ♀ from France
Ventral aspect
2018.02.18 © Picard, J.
Hypogastrura viatica ♀ from France
Rami of retinaculum with 3 exad teeth
2018.02.18 © Picard, J.
Hypogastrura viatica from Belgium
2021.10.09 © Huskens, M.-L.
Hypogastrura viatica from Belgium
Middle foot with 3 tenent macrosetae in transverse row
2021.10.09 © Huskens, M.-L.
Hypogastrura viatica from Belgium
Fore foot with 2 tenent macrosetae in transverse row
2021.10.13 © Huskens, M.-L.
Hypogastrura viatica from Belgium
Dorsal anal spines
2021.10.13 © Huskens, M.-L.
Hypogastrura viatica from Belgium
Furcula
2021.10.13 © Huskens, M.-L.
Hypogastrura viatica from Belgium
2021.10.14 © Huskens, M.-L.
Hypogastrura viatica from Belgium
Eye with 8 ocelli; post-antennal organ (PAO)
Ocelli F+G osculating
2021.10.14 © Huskens, M.-L.
Hypogastrura viatica from Belgium
Hind foot with 3 tenent macrosetae in transverse row
2021.10.14 © Huskens, M.-L.



Species viatica (Bagnall, 1940) partim nec Tullberg, 1872

            Syn.: Achorutes subviaticus var. coerulescens Bagnal, 1941
            Syn.: Achorutes subviaticus var. virescens Bagnal, 1941




Species-group manubrialis             Syn.: Hypogastrura manubrialis Delamare Deboutteville, C, 1953:818,gr

Species sp.1 Delamare Deboutteville, C, 1953:818

Species sp.2 Delamare Deboutteville, C, 1953:818

Species arctandria Fjellberg, A, 1988:35

Hypogastrura arctandria from Norway
After Fjellberg, A, 1988 Fig.1A
            Syn.: Hypogastrura pannosa Fjellberg, 1985



Species assimilis (Krausbauer, 1898) Handschin, E in Dahl, F, 1929:25, B

            Syn.: Achorutes manubrialis var. obtusa Axelson, 1900
            Syn.: Hypogastrura manubrialis Womersley, 1939
            Syn.: Neogastrura manubrialis Stach, J, 1949:50
            Syn.: Hypogastrura manubrialis Maynard, 1951, nec Tullberg, 1869
            Syn.: Hypogastrura devia Christiansen & Bellinger, 1980
            Syn.: Hypogastrura (Hypogastrura) essa Christiansen, K et Bellinger, P, 1980:68
            Syn.: Hypogastrura pannosa Babenko, 1994


Hypogastrura assimilis from the UK
2002 © Hopkin, S.P.
Footcomplex with empodium
and one clavate tenent seta
Hypogastrura assimilis from the UK
2004 © Hopkin, S.P.
Maxillar lamella 1 has only a single fringe

Species breviempodialis (Stach, J, 1949:63) Stach, 1955

Species brevispina (Harvey, 1893) Gisin, 1944

             Syn.: Ceratophysella revispina(sic) Skidmore, 1995 ISS

Species christianseni Yosii, 1960

Species consanguinea (Folsom, 1924) Handschin, 1928

Species coprophila Stach, J, 1960:516

Species devia Christiansen, K et Bellinger, P, 1980:66

Species druki Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:54

Species gisini Strenzke, 1954

Species manubrialis (Tullberg, 1869:9) Linnaniemi, 1912, B

            Syn.: Podurhippus pityriasicus Mégnin, JP, 1878:cxiv
             Syn.: Podurhippus pityriasious(sic) MacGillivray, AD, 1893:316 ISS
             Syn.: Podurhippus pytyriasicus(sic) Bryk, F, 1955:1825 ISS
            Syn.: Achorutes Schötti(sic) Reuter, OM, 1895:5 IOS
             Syn.: Achorutes Schöttii(sic) Reuter, OM, 1895:28 ISS
            Syn.: Achorutes Schötti(sic) Reuter, OM, 1895:31 IOS
            Syn.: Achorutes Schötti(sic) Reuter, OM, 1895:35 IOS
            Syn.: Achorutes Schötti(sic) Carl, J, 1899:296 IOS
            Syn.: Achorutes Schötti(sic) Absolon, K, 1900:268 IOS
            Syn.: Achorutes Schötti(sic) Absolon, K, 1901:87 IOS
            Syn.: Achorutes Schötti(sic) Börner, C, 1901:7 IOS
            Syn.: Achorutes assimilis Krausbauer, 1898
            Syn.: Achorutes (Euachorutes) neglectus Börner, C, 1901:30
            Syn.: Hypogastrura yamagata Kinoshita, 1916
             Syn.: Hypogastrura yamadjata(sic) Denis, J-R, 1924:214 ISS
            Syn.: Hypogastrura yamagata Uchida, 1938
            Syn.: Hypogastrura yamagata Uchida, 1956
            Syn.: Hypogastrura yamagata Salmon, JT, 1964:237
            Syn.: Hypogastrura yamagata Yosii, R, 1977:144
            Syn.: Hypogastrura moniezi Denis, 1923
            Syn.: Hypogastrura Moniezi(sic) Denis, J-R, 1933:320 IOS
            Syn.: Achorutes consanguineus Folsom, 1924
            Syn.: Hypogastrura assimilis Austin, 1937, nec Krausbauer, 1898
            Syn.: Achorutes mucronalis Bagnall, 1941
            Syn.: Hypogastrura manubrialis var. proxenyllodiana Rapoport & Izarra, 1962


Hypogastrura manubrialis from the UK
2005 © Hopkin, S.P.
Empodium without lamella
Hypogastrura manubrialis from the USA
Head with roundish mouthcone
2016.11.20 © Loarie, D.
Hypogastrura manubrialis from the USA
Dorsal aspect; dark field illumination
2016.11.20 © Loarie, D.
Hypogastrura manubrialis from the USA
Ventral aspect; dark field illumination
2016.11.20 © Loarie, D.
Hypogastrura manubrialis from the USA
Furca with broad trapezoid manubrium and 2 short dentes with narrow pointed mucrones
2016.11.20 © Loarie, D.
Hypogastrura manubrialis from the USA
Short anal spines on papillae
2016.11.20 © Loarie, D.
Hypogastrura manubrialis from the USA
Mesofoot with outer unguis (UNG) and inner unguiculus (ung) with narrow basal lamella
2016.11.20 © Loarie, D.



Species matura (Folsom, JW, 1916:481) Christiansen, 1964

            Syn.: Achorutes schötti(sic) Guthrie, JE, 1903:93
            Syn.: Achorutes pannosa Macnamara, 1922 =?
            Syn.: Hypogastrura promatro Christiansen, K et Bellinger, P, 1980:75 =?


Species mexicana Handschin, 1928

Species nepalica Yosii, 1966

Species pahiku Christiansen, K et Bellinger, P, 1992:11

Species pannosa (Macnamara, 1922:153) Gisin, 1944

            Syn.: Hypogastrura essa from Illinois pre2013 =?
            Syn.: Hypogastrura matura from Illinois pre2013 =?


             Syn.: Hypogastrura pannosus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:48 ISS

Species promatro (Wray, 1950) Salmon, 1964

            Syn.: Achorutes utahensis Wray, 1953


Species rangkuli Martynova in Martynova & Chelnokov, 1975

            Syn.: Hypogastrura devia Fjellberg, 1985


Hypogastrura rangkuli from India
After Mandal GP & Arbea JI, 2014 Fig.1

Species satkosiaensis Mandal, GP et Arbea, J, 2019:281

Hypogastrura satkosiaensis from India
After Mandal GP & Arbea J, 2019 Fig.1a

Species serrata (Ågren, H, 1904:5) Shoebotham, 1911

            Syn.: Hypogastrura manubrialis Ripper, 1930
            Syn.: Neogastrura assimilis Stach, 1949, nec Krausbauer, 1898
            Syn.: Hypogastrura ripperi Gisin, 1952


             Syn.: Hypogastrura (Hypogastrura) serratus(sic) Gisin, H, 1960:34 ISS

Species tianshanica Martynova, 1970

Species tigridis (Brown, JM, 1926:201) Gisin, 1960:37

Species tigrina (Harvey, 1900) Gisin, 1944

            Syn.: Achorutes tigrina(sic) Harvey, 1900 IOS
            Syn.: Achorutes tigrina(sic) Folsom, JW, 1916:480 IOS
            Syn.: Anachorutes tigrina(sic) Salmon, JT, 1964:198 IOS
            Syn.: Neanura(lapsus) tigrina Massoud, Z, 1967:319,si



Species turkmenica Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:52

Species ubsunurensis Babenko, AB, 1999:885

Species vernalis (Carl, J, 1901:249) Handschin, E, 1924:91, B

            Syn.: Achorutes manubrialis Reuter, OM, 1895:30
            Syn.: Achorutes nivalis Ågren, 1903
            Syn.: Achorutes Reuteri(sic) Ågren, 1903 IOS
            Syn.: Achorutes Reuteri(sic) Ågren, H, 1904:5 IOS
            Syn.: Achorutes rossi Salmon, JT, 1941:293
            Syn.: Hypogastrura tigridis Handschin, E, 1942:401
            Syn.: Hypogastrura promatro Bernard, EC, 2015:319 =?


Hypogastrura vernalis from Holland
2020.12.30 © van Bezouw, R.
Hypogastrura vernalis from Holland
Dens with mucro
Mucro with longitudinal lamella + apical hook
2020.12.30 © van Bezouw, R.
Hypogastrura vernalis from Holland
Unguiculus with basal lamella
2020.12.30 © van Bezouw, R.
Hypogastrura vernalis from Holland
Dorsal anal valve with 2 anal spines
2020.12.30 © van Bezouw, R.
Hypogastrura vernalis from Holland
Abd.5 with a1+m1+p1 present
2020.12.30 © van Bezouw, R.



Species yosiii Stach, J, 1964:3

            Syn.: Hypogastrura yosii(sic) Stach, J, 1964:3 IOS
            Syn.: Hypogastrura sheyangensis Jiang, JG, Tang, BP et Chen, J-X, 2007:?



            Syn.: Hypogastrura (Hypogastrura) yosii(sic) Rusek, J, 1967:184 IOS
            Syn.: Hypogastrura yosii(sic) Jiang, JG, Tang, BP et Chen, J-X, 2007:63 IOS
            Syn.: Hypogastrura (manubrialis-group) yosii(sic) Jia, J, Skarzynski, D et Konikiewicz, M, 2011:61 IOS
            Syn.: Hypogastrura yosii(sic) Babenko, A, Efeykin, B et Bizin, M, 2020:1 IOS
            Syn.: Hypogastrura yosii(sic) Babenko, A, Efeykin, B et Bizin, M, 2020:3 IOS
            Syn.: Hypogastrura yosii(sic) Babenko, A, Efeykin, B et Bizin, M, 2020:8 IOS
            Syn.: Hypogastrura yosii(sic) Babenko, A, Efeykin, B et Bizin, M, 2020:9 IOS
            Syn.: Hypogastrura (manubrialis-group) yosii(sic) Babenko, A, Efeykin, B et Bizin, M, 2020:10 IOS
            Syn.: Hypogastrura yosii(sic) Babenko, A, Efeykin, B et Bizin, M, 2021:161 IOS
            Syn.: Hypogastrura yosii(sic) Babenko, A, Efeykin, B et Bizin, M, 2021:162 IOS

Species-group trybomi key
            Syn.: Hypogastrura trybomi Christiansen, K et Bellinger, P, 1980:52,gr



Species hargrovei Skarzynski, D, 2007:201

Species maynardi Christiansen & Bellinger, 1980:102, i.s.

            Syn.: Hypogastrura montana Maynard, EA, 1951:42
            Syn.: Biacanthella neomexicana Scott, 1961 ? first instar
            Syn.: Hypogastrura (Hypogastrura) lima Christiansen & Bellinger, 1980
            Syn.: Hypogastrura (Hypogastrura) sp. Bellinger, PF, 1985:78


Species tethyca Ellis, WN, 1976:248

Species theeli (Tullberg, T, 1877:38) Yosii, 1972

            Syn.: Achorutes Theelii(sic) Tullberg, T, 1877:38 IOS
            Syn.: Achorutes Trybomi(sic) Schött, H, 1893:82 IOS

Species trybomi (Schött, H, 1893:82) Gisin, 1960:37

            Syn.: Achorutes Trybomi(sic) Schött, H, 1893:82 IOS
            Syn.: Hypogastrura copiosa Handschin, 1928



Species-subgroup eastern-palaearctic

Species analpapillata Jiang & Yin, 2012

Species heptasetata Jiang & Yin, 2010

Species hexasetata Jiang & Yin, 2010

Species manghe Jia, J, Skarzynski, D et Konikiewicz, M, 2011:57

Species maxillosa Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:70


Species-subgroup nearctic
Species gravesi Wray, 1971

Hypogastrura gravesi from the USA
After Bernard EC, 2015 Fig.1B

Species ireneae (Wray, 1953) Salmon, 1964

            Syn.: Hypogastrura oregonensis Yosii, 1960 =?


             Syn.: Hypogastrura irenae(sic) Skarzynski, D, 2007:203 ISS
             Syn.: Hypogastrura (trybomi-group-nearctic-subgroup) irenae(sic) Skarzynski, D, Kahrarian, M, Piwnik, A, et Zawisza, M, 2017:53 ISS
             Syn.: Hypogastrura (trybomi-group-nearctic-subgroup) irenae(sic) Skarzynski, D, Kahrarian, M, Piwnik, A, et Zawisza, M, 2017:54 ISS
             Syn.: Hypogastrura (trybomi-group-nearctic-subgroup) irenae(sic) Skarzynski, D, Kahrarian, M, Piwnik, A, et Zawisza, M, 2017:55 ISS

Species lima Christiansen, K et Bellinger, P, 1980:100

Species oregonensis Yosii, 1960

            Syn.: Hypogastrura trybomi Hammer, 1953
            Syn.: Hypogastrura (Hypogastrura) trybomi Christiansen, K et Bellinger, P, 1980:106
            Syn.: Hypogastrura (trybomi-group) trybomi Babenko & al, 1994
            Syn.: Hypogastrura (Hypogastrura) trybomi Skidmore, RE, 1995:47
            Syn.: Hypogastrura (Hypogastrura) trybomi Christiansen & Bellinger, 1998
            Syn.: Hypogastrura (trybomi-group) trybomi Babenko & Fjellberg, 2006
            Syn.: Hypogastrura (trybomi-group) trybomi Skarzynski, D, Kahrarian, M, Piwnik, A, et Zawisza, M, 2017:53



Species-group monticola
            Syn.: Hypogastrura monticola Skarzynski, 2009 gr



Species dasiensis Selga, 1966

            Syn.: Hypogastrura (Hypogastrura) crassaegranulata Selga, 1971


Species elevata Cassagnau, P, 1959:484

Species hatiparae Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:87

Species herrerosvelai Arbea, JI et Fernández, TP, 2020:68

Hypogastrura herrerosvelai from Spain
Modified after Arbea JI & Fernández, 2020 Fig.3AB
Hypogastrura herrerosvelai from Spain
Right eye with postantennal organ (PAO) and accessory boss (ab)
Modified after Arbea JI & Fernández, 2020 Fig.4B

Species hispanica Steiner, 1955

Species monticola Stach, J, 1946:1

            Syn.: Hypogastrura fuentei Denis, 1930 =?


Species papillata Gisin, 1949

Species subpapillata Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:77


Species-group crassaegranulata
            Syn.: Hypogastrura (Hypogastrura) crassaegranulata Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:38,gr



Species aushensis Skarzynski, D et Babenko, A, 2010:278

            Syn.: Hypogastrura crassaegranulata Babenko et al., 1994 partim

Species chouardi Cassagnau, P, 1959:487

Species crassaegranulata (Stach, J, 1949:84) Cassagnau, 1959

             Syn.: Hypogastrura crassaecranulata(sic) Kopeszki, H et Meyer, E, 1996:224 ISS
             Syn.: Hypogastrura (crassaegranulata-group) crassaegranualata(sic) s.str. Skarzynski, D et Babenko, A, 2010:275 ISS

Species franconiana (Stach, J, 1949:88) Gisin, 1960:38

            Syn.: Achorutes Schäfferi(sic) Carl, J, 1901:253
            Syn.: Hypogastrura (Hypogastrura) crassaegranulata ssp. burgundiana Gisin, 1960
            Syn.: Hypogastrura crassaegranulata ssp. estaranhensis Cassagnau, P, 1959:485
            Syn.: Hypogastrura crassaegranulata Skarzynski, 2003
            Syn.: Hypogastrura crassaegranulata Babenko et al., 1994 partim

Species ghirkani Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:93

Species himalayana Yosii, 1971

Species hohi Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:93

Species iwamurai Yosii, 1960

Species laxasensillata Lee, 1974

Species microspina Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:91

Species reticulata Börner, C, 1909:99

Hypogastrura reticulata from Japan
2014.01.05 © Hara, Y.
Hypogastrura reticulata from Japan
2014.01.05 © Hara, Y.
Hypogastrura reticulata from Japan
2014.12.23 © Hara, Y.
Hypogastrura reticulata from Japan
2014.12.23 © Hara, Y.
Hypogastrura reticulata from Japan
Distinctly large cuticular tubercles in reticulate pattern
2014.12.23 © Hara, Y.

Species tepli Skarzynski, D et Babenko, A, 2010:276

            Syn.: Hypogastrura crassaegranulata Babenko et al., 1994 partim

Species-group nivicola
            Syn.: Hypogastrura nivicola Yosii, 1960 gr
            Syn.: Hypogastrura (Hypogastrura) socialis Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:38,gr


Hypogastrura nivicola-group from Italy
Interocular tubercles present
2021.01.12 © Colombo, M.

Species bokusii Yosii, R, 1961:14

            Syn.: Hypogastrura (Hypogastrura) bokusi(sic) Yosii, R, 1961:14 IOS
            Syn.: Hypogastrura bokusi(sic) Salmon, JT, 1964:211 IOS
            Syn.: Hypogastrura (socialis-group) bokusi(sic) Skarzynski, D, 2009:il IOS
            Syn.: Hypogastrura (nivicola-group) bokusi(sic) Skarzynski, D, 2019:272 IOS
            Syn.: Hypogastrura (nivicola-group) bokusi(sic) Arbea, JI et Fernández, TP, 2020:75 IOS

Species calceolaris Latzel, R, 1917:235

            Syn.: Hypogastrura calceolaris var. minor Latzel, R, 1917:235


Remark by Gisin, H. 1960:40: Species inquirenda.

Species harveyi (Folsom, JW, 1902:320) Linnaniemi, WM, 1912:31

            Syn.: Achorutes frigidus Axelson, WM, 1905:788 =?
            Syn.: Hypogastrura (Hypogastrura) indiana Christiansen, K et Bellinger, P, 1980:91


             Syn.: Achorutes Harveyi(sic) Folsom, JW, 1902:316 ISS
Hypogastrura harveyi from Canada
2006 © Wigney., B
Hypogastrura harveyi from the USA
2007 01.28 © Murray, T.
Hypogastrura harveyi from the USA
2007.02.26 © Murray, T.
Hypogastrura harveyi from the USA
2007.03.13 © Murray, T.
Hypogastrura harveyi from the USA
2008.01.28 © Trently, D.
Hypogastrura harveyi from the USA
2010.02.09 © Lighthartstudios.com
Hypogastrura harveyi from the USA
2015.03.09 © Eiseman, C.
Hypogastrura harveyi from the USA
Exposed anal sacs
Distinctly large anal spines
2015.03.09 © Eiseman, C.
Hypogastrura harveyi from the USA
Interocular tubercles
2015.03.09 © Eiseman, C.
Hypogastrura harveyi from Canada
Distinctly long antennae
2016.03.13 © Schmidt, H.
Hypogastrura harveyi from Canada
Everted anal sacs
2016.03.13 © Schmidt, H.
Hypogastrura harveyi from Canada
Anal sacs double bubbled
2016.03.13 © Schmidt, H.
Hypogastrura harveyi from the USA
2019.12.29 © Joll, C.
Hypogastrura harveyi from the USA
2019.12.29 © Joll, C.
Hypogastrura harveyi from the USA
2020.02.23 © M, E.
Hypogastrura harveyi from the USA
Distinctly long antennae
2020.02.23 © M, E.
Hypogastrura harveyi from the USA
Distinctly long anal spines
2020.02.23 © M, E.
Hypogastrura harveyi from the USA
Distinctly long anal spines
2020.03.13 © Alloway, L.
Hypogastrura harveyi from the USA
Distinctly long antennae
2021.01.16 © Hendrix, S.



Species janetscheki Steiner, 1959

            Syn.: Hypogastrura (Hypogastrura) socialis Jordana, R et Arbea, JI in Ramos, MA et al, 1997:91
            Syn.: Hypogastrura (Hypogastrura) socialis Jordana, R, Arbea, JI, Simón, C et Luciáñez, MJ in Ramos, MA et al, 1997:759 partim
            Syn.: Hypogastrura socialis Arbea, JI, 2007:120


Species kelmendica Peja, 1985

            Syn.: Hypogastrura (Hypogastrura) socialis Jordana, R, Arbea, JI, Simón, C et Luciáñez, MJ in Ramos, MA et al, 1997:759 partim


Species lapponica (Axelson, 1902) Linnaniemi, 1912

            Syn.: Achorutes frigidus Axelson, WM, 1905:788
            Syn.: Achorutes torneensis Wahlgren, 1906


Species macrotuberculata (Hammer, 1953) Ellis, 1976

Species meridionalis Steiner, 1955

Hypogastrura meridionalis from Spain
Under horse dung
Det.: Jordana R & Baquero E
2015.10.07 © Baquero E. & Zeppelini D.

Species nemoralis Yosii, 1960

Species nivicola (Fitch, 1847) Yosii, 1960, t.t.

            Syn.: Achorutes diversiceps Lintner, 1896
             Syn.: Achorutes diversipes(sic) Handschin, E in Dahl, F, 1929:20 ISS
             Syn.: Achorutes diversipes(sic) Gisin, H, 1960:34 ISS
            Syn.: Achorutes socialis Uzel, 1890, sensu American authors
            Syn.: Achorutes spinifer Schäffer, 1896
            Syn.: Hypogastrura punctata Scott, HG, 1960:58


             Syn.: Achorutes nivicola(sic) Harvey, 1893 ISS
             Syn.: Achorutes nivicola(sic) Bertkau, P, 1894:57 ISS
Achorutes nivicola from the USA
After Folsom JW, 1902 Plate 3 Fig.1




             Syn.: Achorutes (Achorutes) nivicola(sic) Kloet, GS et Hincks, WD, 1945:3 ISS


Hypogastrura nivicola from the USA
2006 © Cossey, J.
Hypogastrura nivicola from Canada
2007 © Johnny.
Hypogastrura nivicola from the USA
With everted anal sacs
2007.03.10 © Reed, D.
Hypogastrura nivicola from the USA
2007.07.05 © Sarratt, J.
Hypogastrura nivicola from the USA
2010.02.10 © Justis, S.
Hypogastrura nivicola from the USA
2010.02.10 © Justis, S.
Hypogastrura nivicola from the USA
2010.02.10 © Justis, S.
Hypogastrura nivicola from Canada
With 3 everted anal sacs
Ready to jump
2016.03.08 © Arbour, G.
Hypogastrura nivicola juv. from the USA
2019.12.26 © Murray, T.
Hypogastrura nivicola from Canada
Ant.4 with exposed apical sensory vesicle
2020.02.24 © Arbour, G.
Hypogastrura nivicola from Canada
With 3 everted anal sacs
Ready to jump
2020.02.25 © Arbour, G.
Hypogastrura nivicola from Canada
Abd.5+6 distinctly smaller
2020.12.25 © Arbour, G.
Hypogastrura nivicola from Canada
Abd.5+6 distinctly smaller
2020.12.25 © Arbour, G.
Hypogastrura nivicola from Canada
With 3 everted anal sacs
2020.12.25 © Arbour, G.
Hypogastrura nivicola from the USA
2021.01.14 © Eiseman, C.
Hypogastrura nivicola from the USA
With interocular tubercles
2021.01.14 © Eiseman, C.
Hypogastrura nivicola from the USA
With 3 exposed anal sacs
2021.01.14 © Eiseman, C.

Species ottawana Cipola, NG, 2020:200

Hypogastrura ottawana from Canada
After Cipola NG, 2020 Fig.1A
Hypogastrura ottawana from Canada
After Cipola NG, 2020 Fig.1B

Species packardi (Folsom, JW, 1902:319) Linnaniemi, WM, 1912:31

            Syn.: Achorutes nivicola Packard, 1873
            Syn.: Achorutes lapponicus Axelson, 1902 =?
            Syn.: Achorutes packardi var. dentatus Folsom, JW, 1902:320
            Syn.: Achorutes notha Macnamara, 1922
            Syn.: Hypogastrura packardi f. furcata Maynard, EA, 1951:33


Species peloponnesica Dányi, L, 2013:158

Hypogastrura peloponnesica from Greece
After Dányi L, 2013 Fig.4
Hypogastrura peloponnesica from Greece
After Dányi L, 2013 Fig.5
Hypogastrura peloponnesica from Greece
Prominent cephalic tubercles (*)
After Dányi L, 2013 Fig.12

Species reticulata2 Hammer, 1953

Species simsi Hart & Waltz, 1995

Species socialis (Uzel, 1891) Linnaniemi, 1912

            Syn.: Achorutes diversiceps Lintner, 1896
            Syn.: Achorutes mitis Ågren, H, 1904:6,si
            Syn.: Achorutes torneensis Wahlgren, 1906
            Syn.: Hypogastrura socialis var. rufipes Latzel, R, 1917:234


Hypogastrura socialis from Germany
Furcula
2006 © Schulz H.-J. & Schubert H.D.
Hypogastrura socialis from Spain
Modified after Arbea JI & Fernández, 2020 Fig.5AB
Hypogastrura socialis from Spain
Right eye with postantennal organ (PAO) and accessory boss (ab)
Modified after Arbea JI & Fernández, 2020 Fig.6I

Species spei Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:83

Species tooliki Fjellberg, 1985

Hypogastrura tooliki from the USA
2009.04.09 © Carr, J.
Hypogastrura tooliki from the USA
2009.04.10 © Carr, J.
Hypogastrura tooliki from the USA
Cephalic interocular tubercles
2009.04.10 © Carr, J.

Species tsukubaensis Tamura, 1997

Species yongmuensis Lee, 1974

             Syn.: Hypogastrura (socialis-group) yongmunensis(sic) Skarzynski, D, 2009:il ISS

Species-group packardi
            Syn.: Hypogastrura packardi Christiansen, K et Bellinger, P, 1980:52,gr
            Syn.: Hypogastrura (Hypogastrura) sahlbergi Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:38,gr
             Syn.: Hypogastrura sahlbergii(sic) Skarzynski, D et Babenko, A, 2010:278,gr ISS
             Syn.: Hypogastrura sahlbergii(sic) Mandal, GP et Arbea, JI, 2014:132,gr ISS


Species austriaca Babenko & Thibaud, 1990

Species copiosa (Folsom, JW, 1916:480) Handschin, 1928

            Syn.: Achorutes schneideri Guthrie, JE, 1903:91
            Syn.: Hypogastrura gigas Maynard, 1951
            Syn.: Hypogastrura ulmea Maynard, EA, 1951:38
            Syn.: Xenyllodes hoffi Scott, HG, 1960:185


Species fjellbergi Babenko & Bulavintsev, 1993

            Syn.: Hypogastrura trybomi Fjellberg, 1985
            Syn.: Hypogastrura (Hypogastrura) trybomi Skidmore, RE, 1995:47
            Syn.: Hypogastrura trybomi Babenko, A, Stebaeva, S et Turnbull, MS, 2019:14


Species funesta Christiansen & Bellinger, 1980:89

Species gennargentui Dallai, R, 1971:189

Species longimucrona Lee & Choe, 1979

Species madera Christiansen & Bellinger, 1980:94

Species pomorskii Skarzynski, D, 2010:258

Species sahlbergi (Reuter, OM, 1895:31) Handschin, E, 1924:91, B

            Syn.: Achorutes Sahlbergi(sic) Reuter, OM, 1895:31 IOS

            Syn.: Achorutes theeli Tullberg, 1876
            Syn.: Hypogastrura sahlbergi var. rosea Reuter, 1895
            Syn.: Achorutes schneideri Schäffer, 1896
            Syn.: Achorutes affinis Schäffer, 1900, nec Lucas, 1846
             Syn.: Achorutes Schäfferi(sic) Carl, J, 1901:253 ISS
             Syn.: Achorutes (Achorutes) schäfferi(sic) Womersley, 1930 ISS
             Syn.: Achorutes (Achorutes) schäfferi(sic) Kloet, GS et Hincks, WD, 1945:3 ISS
            Syn.: Achorutes strenuus Brown, 1923
            Syn.: Achorutes (Achorutes) serratus Kloet, GS et Hincks, WD, 1945:3



            Syn.: Achorutes Sahlbergi(sic) Reuter, OM, 1895:35 IOS
Hypogastrura sahlbergi from the UK
2004 © Hopkin, S.P.
5th abdominal segment with only anterior and posterior row of setae
Hypogastrura sahlbergi from the UK
2021.02.03 © Cann, A.J.
Hypogastrura sahlbergi from the UK
2021.02.03 © Cann, A.J.
Hypogastrura sahlbergi from the UK
Th.2 seta m2 present
Th.3 seta m2 absent
2021.02.03 © Cann, A.J.
Hypogastrura sahlbergi from the UK
Furcula extended backwards in ventral aspect
Mucro (m) < 1/3 of dens
2021.02.05 © Cann, A.J.
Hypogastrura sahlbergi from the UK
Mucro
2021.02.05 © Cann, A.J.
Hypogastrura sahlbergi from the UK
Unguiculus (u) with basal lamella (bl)
Unguis (U) with inner tooth (it)
2021.02.05 © Cann, A.J.



Species sparta Christiansen & Bellinger, 1980:97

Species szeptyckii Skarzynski, D, 2006:83

            Syn.: Hypogastrura crassaegranulata Szeptycki, 1967
            Syn.: Hypogastrura crassaegranulata Weiner, 1981
            Syn.: Hypogastrura crassaegranulata Babenko et al., 1994 partim


Species tatrica (Stach, 1949:31) Gisin, 1960:39

Species tchabensis Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:100


Species-group unresolved

Species aethiopica (Tarsia in Curia, 1939) Salmon, JT, 1964:203

Species agaricina Bourlet, 1841

            Syn.: Achorutes agaricorum Gervais in Walckenaër, 1844
            Syn.: Podura (pas de crochets à l'extrémité du corps) cyanocephala Nicolet, H, 1842:56
Podura cyanocephala from Switzerland
After Nicolet H, 1842 Pl.5 Fig.8.




Species albella (Koch, 1840) Salmon, JT, 1964:204

            Syn.: Achoreutes Albellus(sic) Koch, 1840 IOS

Species arnei Skarzynski, D, 2019:269

Hypogastrura arnei from the USA
Ceratrimeria habitus
After Skarzynski D, 2019 Fig.1

Species aterrima Yosii, R, 1972:77

Species bilineata (Koch, 1840) Salmon, JT, 1964:211

Species capitata Cassagnau, P et Delamare Deboutteville, C, 1955:368

Species carpatica Nosek, J, 1962:13

Species crassa (Oudemans, 1890) Handschin, 1925

Species deserti Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:42

Species elegans (Parfitt, 1891) Goto, HE & Lawrence, PN in Kloet, GS & Hincks, WD, 1964:4

Species ellisi Skarzynski D et Kahrarian M in Skarzynski, D, Kahrarian, M, Piwnik, A, et Zawisza, M, 2017:46

Species exigua Gisin, 1958

            Syn.: Hypogastrura parvula Haybach, 1972 partim


Species fuentei Denis, 1930

Species hyperborea (Boheman, 1865) Bellinger, 1996 si

Species hypnorum (Fabricius O, 1783) Salmon, JT, 1964:216

Species inopinata Deharveng & Le Cong Man, 2002

Species intermedia Handschin, 1926 -

Species japonica Scott, 1961

            Syn.: Hypogastrura (Hypogastrura) sp. Bellinger, PF, 1985:79


Species magistri Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:70

Species microspinata Jiang & Yin, 2012

Species mongolica (Nosek, 1976)

Species montana (Becker, 1905), s.i.

Species myrmecophila (Womersley, 1929) Denis, J-R, 1931:80

Species norica Latzel, R, 1917:234

Species obliqua Salmon, JT, 1949:10

Species oreophila Butschek, 1948

Species paradoxa Yosii, R, 1965:3

Species persica Kahrarian M, Vafaei-Shoushtari R, Skarzynski D et Konikiewicz M in Kahrarian, M, Vafaei-Shoushtari, R, Skarzynski, D, Konikiewicz, M, Soleymannezhadyan, E, Mehr, MS et Shams, B, 2013:91

Species pityusica Ellis, WN, 1974:125

Species pizzoci Fanciulli, PP et Dallai, R, 2008:16

Species prabhooi Bhattacharjee, 1985

             Syn.: Hypogastrura (sahlbergii-group?) prabhoi(sic) Mandal, GP et Arbea, JI, 2014:132 ISS
             Syn.: Hypogastrura (packardi-group?) prabhoi(sic) Mandal, GP et Arbea, JI, 2014:132 ISS
             Syn.: Hypogastrura prabhoi(sic) Mandal, GP, 2018:119 ISS
             Syn.: Hypogastrura prabhoi(sic) Mandal, GP et Arbea, J, 2019:280 ISS
             Syn.: Hypogastrura prabhoi(sic) Mandal, GP et Arbea, J, 2019:289 ISS

Species protoviatica Handschin, 1926 -

Species pseudohispanica Loksa, 1966

Remark by Dányi, L. & Traser, G. 2008:9: As Loksa didn't provide any description or illustration, we have to consider it as nomen nudum.

Species punctata (Coleman, 1941, s.d.) Christiansen, K & Bellinger, P, 1980:60

Species pyrenaica (Cassagnau, P, 1959:500) Cassagnau, P, 1974:307

             Syn.: Hypogastrura pyrenaïca(sic) Cassagnau, P, 1974:307 ISS

Species ramia Lee & Choe, 1979

Species rehi Börner, C, 1906:166

Species similis (Absolon, 1901) Gisin, H, 1960:40, s.i.

Species sonapani Baijal, HN, 1958:349

Hypogastrura sonapani from the Himalaya
After Baijal HN, 1958 Fig.1A

             Syn.: Hypogastrura Sonapani(sic) Santeshwari, X, Raghuraman, M et Singh, J, 2015:572 ISS
Hypogastrura sonapani from India
After Santeshwari, Raghuraman M & Singh J, 2015 p.573



Species spelaea (Joseph, 1882) Handschin, 1929

Species temarpurensis (Tyagi & Baijal, 1982, i.s.) Tyagi & Baijal, 1982

Species trilobata

Species unguiculata (Mitra, 1966) Mitra, 1977

Species verruculata Rusek, J, 1967:184

Species wisei Camousseight, A, 1999:5 nn

Species zivadinovici Palissa, 1968

Subgenus Franzura Cassagnau, P et Deharveng, L, 1976:199
Species synacantha Cassagnau, P et Deharveng, L, 1976:199, t.t.


Genus Willemia Börner, 1901 key  key

F
Species sp.1 Schneider, C et Deharveng, L, 2017:2

Species acantha Bonet, 1945

Species annapurna D'Haese, C et Weiner, WM, 1998:977

Species anophthalma Börner, 1901, t.t., B

            Syn.: Willemia anophthalma var. inermis Börner, C, 1901:698


             Syn.: Willemia anophtalma(sic) Marlier, 1942,B ISS
             Syn.: Willemia anophthalna(sic) Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:236 ISS
             Syn.: Willemia anophtalma(sic) García-Gómez, A et Cutz-Pool, LQ, 2008:1 ISS
             Syn.: Willemia Anophthalma(sic) García-Gómez, A et Cutz-Pool, LQ, 2008:2,gr ISS
             Syn.: Willemia anophtalma(sic) García-Gómez, A et Palacios-Vargas, JG, 2019:564 ISS

Species antennomonstrum Bu, Y, Potapov, MB et Gao, Y, 2012:581

            Syn.: Willemia (buddenbrocki-group) Antennomonstrum(sic) Bu, Y, Potapov, MB et Gao, Y, 2012:581 IOS

Species arenicola Palacios-Vargas, JG et Vázquez, M, 1989:433

Species arida Fjellberg, A, 1991:205

Species bedosae D'Haese, C, 1998:582

Species bellingeri Palacios-Vargas, JG et Vázquez, M, 1989:435

Species brevispina Hüther, 1962

            Syn.: Willemia australis Rapoport, 1962
            Syn.: Willemia (buddenbrocki-group) australis D'Haese, C et Weiner, WM, 1998:969
            Syn.: Willemia peke Christiansen, K et Bellinger, P, 1992:28


Species buddenbrocki Hüther, 1959

            Syn.: Willemia australis Rapoport, EH et Rubio, IR, 1963:97
            Syn.: Willemia australis Mari Mutt, JA et Bellinger, PF, 1990:17
            Syn.: Willemia australis Salazar Martínez, AE, Narváez Beinhorn, MJ et Bernava Laborde, V, 2017:103
            Syn.: Willemia buddenbrocki ssp. brevispina Mari Mutt, JA et Bellinger, PF, 1990:17


Willemia buddenbrocki from Chile
After Rapoport EH & Rubio IR, 1963 Fig.7

Species bulbosa Bonet, 1945

Species christianseni D'Haese, C, 1998:586

Species deharvengi D'Haese, C et Weiner, WM, 1998:970

Species delamarei Prabhoo, 1971

Species denisi Mills, HB, 1932:263, B

            Syn.: Willemia anophthalma var. inermis Gisin, H, 1960:67
            Syn.: Willemia inermis Gisin, 1950
            Syn.: Willemia aspinata Stach, 1949
            Syn.: ... scandinavica Stach, 1949 =?
            Syn.: Willemia aspinata ssp. sensibilis Jordana, R, 1981:4
Willemia aspinata ssp. sensibilis from Spain
After Jordana R, 1981 Lam.II Fig.7

            Syn.: Willemia elisabetharum Weiner, 1986
            Syn.: Willemia elisabethum Weiner, 1986 IOS





Species dhaesei Bu, Y, Potapov, MB et Gao, Y, 2012:583

            Syn.: Willemia (buddenbrocki-group) Dhaesei(sic) Bu, Y, Potapov, MB et Gao, Y, 2012:583 IOS

Species dubia Christiansen, K et Bellinger, P, 1980:238

Species fjellbergi Potapov MB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:247

Species granulata Fjellberg, 1985

Species intermedia Mills, HB, 1934:17

            Syn.: Willemia scandinavica Stach, 1949 partim
            Syn.: Willemia biseta Christiansen, K et Bellinger, P, 1980:237,sd
            Syn.: Willemia barbarae Weiner, WM, 1986:334


Species iztaccihuatlensis García-Gómez, A et Cutz-Pool, LQ, 2008:2

             Syn.: Willemia Iztaccihuatlensis(sic) García-Gómez, A et Cutz-Pool, LQ, 2008:3 ISS

Species japonica Yosii, R, 1970:1

Species koreana Thibaud & Lee, 1994

            Syn.: Willemia (anophthalma-group) Koreana(sic) Bu, Y, Potapov, MB et Gao, Y, 2012:585 IOS

Species meybholae Palacios-Vargas, JG, 1987:6

             Syn.: Willemia meyboholae(sic) Palacios-Vargas, JG, de Lima, ECA et Zeppelini, D, 2013:1586 ISS

Species multilobata Gers & Deharveng, 1985

Species nadchatrami Yosii, R, 1959:5

Species namibiae Thibaud & Massoud, 1988

Species neocaledonica Weiner, 1991

             Syn.: Willemia neocaldonica(sic) D'Haese, C et Weiner, WM, 1998:985 ISS

Species nepalensis D'Haese, C et Weiner, WM, 1998:979

Species nosyboraha Thibaud, J-M, 2008:505

Species panamaensis García-Gómez, A et Palacios-Vargas, JG, 2019:565

Species persimilis Bonet, 1945

Species psammophila Palacios-Vargas, JG et Thibaud, J-M, 2001:162

Species scandinavica Stach, 1949

            Syn.: Willemia anophthalma var. inermis Linnaniemi, 1912


Species setonychia Prabhoo, 1971

Species shanghaiensis Yue, Q-Y, 1999:1

Species similis Mills, HB, 1934:17

            Syn.: Willemia vashtia Wray, 1950


Species subbulbosa Thibaud, 1994

Species tali Kaprus, IJ et Nevo, E, 2003:65

Willemia tali from Israel
After Kaprus I.J. & Nevo E, 2003 Fig.1a
             Syn.: Willemia talae(sic) Kaprus, IJ et Nevo, E, 2003:66 ISS


Species tondoh Zon, Bedos & D'Haese, 2015

Species trilobata Barra, J-A, 1995:125

Species trisphaerae Potapov MB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:244

Species unispina Fjellberg, A, 2007:33

Species virae Kaprus, IJ, 1997:37

Willemia virae from Ukraine
After Kaprus, I.J, 1997 Fig.1-2

Species wandae Tamura & Zhao, 1997

Species zeppelinii D'Haese & Thibaud, 2011

            Syn.: Willemia zeppelini(sic) D'Haese & Thibaud, 2011 IOS


             Syn.: Willemia zeppelini(sic) Bu, Y, Potapov, MB et Gao, Y, 2012:582 ISS



Species zhaoi Tamura, Yin & Weiner, 2000


Genus Orogastrura Deharveng & Gers, 1979 key

F
Species dilatata (Cassagnau, P, 1959:73) Deharveng & Gers, 1979, t.t.

            Syn.: Mesachorutes espanoli Cassagnau, 1961 =?
            Syn.: Mesachorutes espanoli Deharveng & Gers, 1979
             Syn.: Mesachorutes espagnoli(sic) Arbea, JI et Jordana, R, 1990:26 =? ISS
            Syn.: Mesachorutes espanoli Thibaud, J-M, Schulz, H-J et da Gama Assalino, MM, 2004:152


Species emucronata Deharveng & Gers, 1979

             Syn.: Orogastrura dilatata-emucronata(sic) Gers, C, 1980:306 ISS

Species espanoli (Selga, 1959) Jordana, R & Arbea, JI in Ramos, MA & al, 1997:193

            Syn.: Mesachorutes españoli(sic) Selga, 1959 IOS
            Syn.: Orogastrura fusca Gers, C, 1980:304
Orogastrura fusca from France
Modified after Gers C, 1980 Fig.1C

            Syn.: Orogastrura fusca Arbea, JI et Jordana, R, 1990:26 =?
            Syn.: Orogastrura fusca Jordana, R et Arbea, JI in Ramos, MA et al, 1997:193 =?
            Syn.: Orogastrura fusca Arbea, JI, 2020:282




            Syn.: Mesachorutes españoli(sic) Salmon, JT, 1964:240 IOS
             Syn.: Mesachorutes (Denigastrura) espagnoli(sic) Selga, 1971 ISS
             Syn.: Mesachorutes espagnoli(sic) Arbea, JI et Jordana, R, 1990:26 ISS
Orogastrura espanoli juv. from Spain
After Arbea JI, 2020 Fig.2
Orogastrura espanoli juv. from Spain
Right eye
Tentative designation of ocelli
After Arbea JI, 2020 Fig.2





Species octoseta Arbea, JI et Jordana, R, 1990:18

Species pallida (Cassagnau, P, 1954:69) Deharveng & Gers, 1979

Species parva (Gisin, 1949) Deharveng & Gers, 1979

            Syn.: Hypogastrura monticola Gisin, 1947, nec Stach, 1946
            Syn.: Hypogastrura parvula Haybach, 1972 partim
            Syn.: Hypogastrura parvula Skarzynski, D, 2011:63 partim
            Syn.: Hypogastrura parvula Deharveng, L, Bedos, A et Duran, V, 2015:191


Species stebaevae Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:206

Species tetrophthalma Deharveng, L, Bedos, A et Duran, V, 2015:181

Orogastrura tetrophthalma from France
After Deharveng L, Bedos A & Duran V, 2015 Fig.2A


Genus Mesachorutes Absolon, K, 1900:266

M
  Syn.: Achorutidae Achorutini(sic) Börner, C, 1901:13 partim IOS
  Syn.: ... Beckerella Linnaniemi, 1912
  Syn.: ... Troglogastrura Stach in Demel, 1918
  Syn.: ... Beckerellodes Salmon, 1945
  Syn.: ... Subbeckerella Stach, 1949




Species sp. Lukic, M, 2019:309

Mesachorutes undescr. from Croatia
After Lukic M, 2019 Fig.1B

Species quadriocellatus Absolon, K, 1900:267, t.t., B

            Syn.: Mesachorutes 4-ocellatus(sic) Absolon, K, 1900:267,tt IOS
            Syn.: Mesachorutes cionii Denis, 1934
            Syn.: Mesachorutes marlieri Delamare Deboutteville, 1947,B


             Syn.: Mesachorutes 4-ocellatus(sic) Absolon, K, 1901:581 ISS
             Syn.: Mesachorutes 4-ocellatus(sic) Bonet, F, 1930:119 ISS
             Syn.: Mesachorutes 4-oculatus(lapsus) Bonet, F, 1930:121 ISS
             Syn.: Mesachorutes 4-ocullatus(lapsus) Bonet, F, 1930:128 ISS
             Syn.: Mesachorutes 4-oculatus(lapsus) Bonet, F, 1930:131 ISS
             Syn.: Mesachorutes 4-ocellatus(sic) Bonet, F, 1930:131 ISS
             Syn.: Hypogastrura (Mesachorutes) 4-ocellata(sic) Bonet, F, 1930:131 ISS
             Syn.: Mesachorutes 4-ocellatus(sic) Bonet, F, 1930:135 ISS
Mesachorutes quadriocellatus from Spain
After Arbea JI, 2013 Fig.3





Species thomomys (Chamberlain, 1943) Christiansen & Bellinger, 1980:200


Genus Taurogastrura Vargovitsh, RS, 2007:30

F
Species skelica Vargovitsh, RS, 2007:30, t.t.

Taurogastrura skelica from the Ukraine
After Vargovitsh, R.S., 2007 Fig.2.


Genus Celegastrura Palacios-Vargas, JG, Mendoza, S et Villalobos, FJ, 2000:140

F
Species aldebaranis Palacios-Vargas, JG, Mendoza, S et Villalobos, FJ, 2000:140, t.t.

             Syn.: Celegastrura aldebarani(sic) Palacios-Vargas, JG, Mendoza, S et Villalobos, FJ, 2000:142 ISS



Genus Ceratophysella Börner in Brohmer, 1932 key  key  key

F
  Syn.: Hypogastrura Ceratophysélla(sic) Börner in Brohmer, 1932 IOS
  Syn.: ... Cystioceras Börner in Schille, 1912, nn
  Syn.: ... Mitchellania Wray, 1953
  Syn.: ... Achurotes(sic) Baijal, HN, 1958:350 ISS
  Syn.: ... Cyclograna Yosii, 1961
  Syn.: ... Spinachorutes Scott, 1962
  Syn.: Hypogastrurinae Spinifacies Scott, HG, 1963:108
  Syn.: Isotomidae Cryptopygus(sic) Falahati HosseinAbad, A, Mehr, MS et Khyroodin, A, 2013:258 lapsus


  Syn.: Collembola Caratophysella(sic) Yosii, R, 1965:3 ISS
Ceratophysella sp. from Sweden
2006 © Hall, K.
Remark by ICZN. 2007:156: Placed on the Official List of Names in Zoology. Opinion 1193.

Ceratophysella sp. from the USA
2007 © Feldman, M.
Ceratophysella sp. from the USA
2007.02.16 © Stone, W.
Ceratophysella sp. from the USA
2007.03.27 © Murray, T.
Ceratophysella sp. from the UK
2007.04.01 © Cornwall, N.J.
Ceratophysella sp. from Canada
2007.12.17 © McCann, S.
Ceratophysella sp. from the USA
2008.01.26 © Gross, J.
Ceratophysella sp. from the UK
2008.02.11 © Farrell, S.
Ceratophysella sp. from the USA
2008.02.23 © Boeddeker, M.
Ceratophysella sp. from the UK
2009.03.09 © Talbot, M.
Ceratophysella sp. from Tasmania
Crawling leg sequence
2009 © Daley, A.
Ceratophysella sp. from Tasmania
Everting antennal sacs just before jump
2009 © Daley, A.
Ceratophysella sp. from Tasmania
Drinking from drop of water
2009 © Daley, A.
Ceratophysella sp. from the USA
2009.10.27 © Ten Eyck, C.
Ceratophysella sp. from Norway
2011 © Coulson, S.J.
Ceratophysella sp. from the UK
2012.09.26 © Murray, A.
Ceratophysella sp. from Spain
With distinct anal spines
2012.11.11 © Montoro, A.
Ceratophysella sp. from the UK
Composed eye (with 8 ocelli) and post-antennal organ
2012.12.25 © Murray, A.
Ceratophysella sp. from South-France
Post-antennal organ with 4 lobes
2013.02.13 © Delhem, R.
Ceratophysella sp. from the UK
Mouthcone
2013.03.23 © Murray, A.
Ceratophysella sp. from the UK
Foot of 3rd leg with outer large unguis (U) and inner small unguiculus (u)
2015.02.01 © Garlick, T.
Ceratophysella succinea from Belgium
Furca with broad basal manubrium, short thick dentes and twisted mucronal teeth
2017.11.08 © Huskens, M.-L.
Ceratophysella succinea from Belgium
Post-antennal organ (PAO) with 4 lobes
2017.11.08 © Huskens, M.-L.
Ceratophysella sp. from France
Ready to jump
With everted antennal + anal sacs
2018.05.11 © Picard, J.
Ceratophysella sp. from the USA
2019.02.02 © Westfall, Bo.
Ceratophysella sp. from the USA
2019.02.06 © Bairstow, A.
Det.: Potapov, M.
Ceratophysella sp. from Germany
Landing on sticky anal sac after jump
2020.02.15 © Fabian, Be.
Ceratophysella sp. from the UK
Mucro with twisted teeth
2020.03.06 © Deed, Be.
Ceratophysella sp. from Belgium
Distinctly different long + short setae
2021.01.18 © Huskens, M.-L.
Ceratophysella sp. from Belgium
Distinctly large anal spines
2021.01.18 © Huskens, M.-L.
Ceratophysella sp. ♂ from Taiwan
2021.08.12 © Cheng, H.-J.
Ceratophysella sp. ♂ from Taiwan
2021.08.12 © Cheng, H.-J.
Ceratophysella bengtssoni from Belgium
Abd.4 with longitudinal lateral fold
2022.01.22 © Huskens, M.-L.





Species sp.1 (Denis, J-R, 1924:214)

Species sp.2 Schneider, C et Deharveng, L, 2017:13

Species adexilis Stach, J, 1964:4

Species ainu (Yosii, R, 1972:75)

Species alachuensis Skarzynski, D, 2007:139

Species alani (Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB & Stebaeva, SK, 1994:188) Skarzynski, D & Christiansen, K, 2008:990

Species annae (Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB & Stebaeva, SK, 1994:180) Park, K-H & Park, N-Y, 2006:492

Species anshanensis (Wu & Xie, 2007) Jia, J, Skarzynski, D & Li, Y, 2010:61

Species armata (Nicolet, H, 1842:57) Börner, 1932, t.t., Bhabitus lateral

Podura armata from Switzerland
After Nicolet H, 1842 Pl.5 fig.6

            Syn.: Hypogastrura fusco-viridis(sic) Bourlet, 1843 IOS
            Syn.: Achorutes marmoratus Packard, AS, 1873:30
            Syn.: Achorutes Texensis(sic) Packard, AS, 1873:30
            Syn.: Achorutes longispinus Schäffer, 1897
             Syn.: Achorutes longispinis(sic) Salmon, JT, 1964:206 ISS
            Syn.: Achorutes armatus ab. pallens Krausbauer, 1898
            Syn.: Achorutes filiformis Wahlgren, 1906, sensu Salmon, 1964
            Syn.: Achorutes sigillatus Liniger, 1919, sensu Salmon, 1964
            Syn.: Hypogastrura luteospina Stach, 1920
            Syn.: Achorutes buxtoni Brown, JM, 1926:203
            Syn.: Proxenyllodes folsomi Denis, 1926
             Syn.: Proxenyllodes Folsomi(sic) Denis, J-R, 1931:81 ISS
            Syn.: Hypogastrura armata var. decolor Bonet, F, 1933:127
            Syn.: Hypogastrura armata var. unicolor Bonet, 1934 sensu Mari Mutt & Bellinger, 1990
            Syn.: Ceratophysella crassispina Cassagnau, P, 1952:307
             Syn.: Hypogastrura crasispina(sic) Jordana, R, 1980:6 ISS
Ceratophysella crassispina
Broad cylindrical anal spines
2006 © Boldsystems.org
Ceratophysella armata f. crassispina from France
Broad cylindrical anal spines
2016.11.11 © Réal, J.-A.
Ceratophysella armata f. crassispina from France
2016.11.11 © Réal, J.-A.
Ceratophysella armata f. crassispina from France
Post antennal organ (PAO)
2016.11.11 © Réal, J.-A.
Ceratophysella armata f. crassispina from France
2018.09.23 © Dejoux, P.
Ceratophysella armata f. crassispina from France
2020.09.20 © Dejoux, P.
Ceratophysella armata f. crassispina from France
2020.09.20 © Dejoux, P.

            Syn.: Hypogastrura (Ceratophysella) armatissima Gisin, 1958
             Syn.: Hypogastrura armatisima(sic) Gisin, Bougeois & Cassagnau, 1972 ISS
            Syn.: Hypogastrura (Ceratophysella) acanthospina Steiner, 1958
            Syn.: Ceratophysella macronycha Steiner, 1958
            Syn.: Ceratophysella attenuata Cassagnau, P, 1959:496
            Syn.: Ceratophysella armata var. triaena Rapoport, 1959
            Syn.: Hypogastrura (Ceratophysella) armata f. brunea Bourgeois, A et Cassagnau, P in Vandel, A, 1973:1197
            Syn.: Hypogastrura sinetertiaseta Lee, 1974
             Syn.: Hypogastrura aniiala(sic) Bhagat, 2013 ISS
            Syn.: Hypogastrura granulata Skarzynski, D, Smolis, A, Kovác, L et Porco, D, 2021:4 partim,cf



             Syn.: Podura armatus(sic) Lucas, 1844 ISS
Achorutes armatus from the UK
Saltatory appendage
After Lubbock J, 1870 Pl.XXII Fig.22
Achorutes armatus from the UK
After Lubbock J, 1873 Pl.40 Fig.1
Achorutes armatus from the UK
After Lubbock J, 1873 Pl.40 Fig.2

             Syn.: Achorutes armata(sic) Moniez, 1890 ISS
             Syn.: Achorutes armata(sic) Schille, 1906 ISS
             Syn.: Achoreutes(sic) armatum(sic) Popenoe, 1912 ISS
             Syn.: Achoreutes(sic) armatum(sic) Weiss, 1916 ISS
             Syn.: Hypogastrura armatus(sic) Denis, J-R, 1924:215 ISS
             Syn.: Hypogastrura armatus(sic) Denis, J-R, 1924:215,gr ISS
             Syn.: Hypogastrura armatus(sic) Denis, J-R, 1931:81 ISS
             Syn.: Achorutes armata(sic) van den Bruel, WE in de Jonghe d'Ardoye, E, 1937:321 ISS
             Syn.: Hypogastrura (armata-group) armatus Denis, J-R, 1947:1 ISS
             Syn.: Achorutes (Hypogastrura) armata(sic) Poole, 1959 ISS
             Syn.: Ceratophysella armatus(sic) Yosii, R, 1965:3,gr ISS
             Syn.: Hypogastrura armatus(sic) Snider, RJ, 1967:196 ISS
             Syn.: Hypogastrura armatus(sic) Peterson, AJ, 1971:81 ISS
Ceratophysella armata from Spain
2002 © Bacquero, E. & Jordana, R.
Ceratophysella armata from Italy
2006 © Paolucci, P.

             Syn.: Hypogastrura armatus(sic) Glime, JM, 2015:3 ISS
Ceratophysella armata from Holland
Anal spines on large contiguous papillae
2015.02.13 © van Duinen, J.
Ceratophysella armata from France
2016.12.17 © Quintin, C.
Ceratophysella "armata" from the USA
2017.01.08 © Abela, A.
Ceratophysella "armata" from the USA
Anal papillae contiguous basally
2017.01.08 © Abela, A.
Ceratophysella armata from France
2017.03.19 © Quintin, C.
Ceratophysella armata from France
Abd.4: p1 > p2
2018.12.03 © Tiky, S.
Ceratophysella armata from France
Post-antennal organ (PAO)
2019.06.15 © Garcelon, P.

  Subspecies atypica Cassagnau, P, 1954:67

  Subspecies helvispina Latzel, R, 1917:233

  Subspecies inermis Axelson, WM, 1905:788

  Subspecies trispina Womersley, H, 1934:445





Species ateruii (Tamura, 2001) Nakamori, T, 2013:377

Species attenuata (Gisin, 1960) Christian & Schönmann, 1983

Species baichengensis Wu & Yin, 2007

Species baltica (Tyagi & Baijal, 1982) Mandal, GP & Arbea, J, 2019:279

Hypogastrura baltica from India
After Santeshwari, Raghuraman M & Singh J, 2015 p.573

Species bengtssoni (Ågren, H, 1904:2) Stach, J, 1949:142, B

            Syn.: Achorutes Bengtssoni(sic) Ågren, H, 1904:2 IOS
            Syn.: Podura rufescens Nicolet, 1842
            Syn.: Achorutes sigillata Uzel, 1891
            Syn.: Achorutes navicularis Schött, H, 1893:83
            Syn.: Achorutes sigillatus var. stygius Absolon, K, 1900:268
            Syn.: Achorutes (Achorutes) sigillatus Axelson, 1900, nec Uzel, 1890
            Syn.: Hypogastrura bengtssoni var. fennica Agrell, 1932
            Syn.: Agreniella angularis Bagnall, 1949
            Syn.: Agreniella collingei Bagnall, 1949


             Syn.: Achorutes Bengtssoni(sic) Wahlgren, 1906 ISS
             Syn.: Hypogastrura Bengtssoni(sic) Linnaniemi, WM, 1912:17 ISS
             Syn.: Hypogastrura Bengtssoni(sic) Cassagnau, P in Vandel, A, 1956:1362 ISS


Ceratophysella bengtssoni from the UK
2002 © Hopkin, S.P.
Ceratophysella bengtssoni from the UK
2004 © Hopkin, S.P.
Ceratophysella bengtssoni from the UK
2007.12.10 © Valentine., B
Ceratophysella bengtssoni from the UK
2007.12.28 © Valentine., B
Ceratophysella bengtssoni from the UK
2008.01.01 © Valentine., B
Ceratophysella bengtssoni from the UK
2008.01.14 © Valentine., B
Ceratophysella bengtssoni from the UK
2008.11.19 © Valentine., B
Ceratophysella bengtssoni from the UK
2008.12.20 © Valentine., B
Ceratophysella bengtssoni from the UK
2008.12.21 © Valentine., B
Ceratophysella bengtssoni from the UK
With everted anal sacs and antennal vesicles
2009.01.25 © Valentine., B
Ceratophysella bengtssoni from the UK
Facial pattern
2009.01.29 © Valentine., B
Ceratophysella bengtssoni from the UK
2012.01.18 © Jones, A.
 
Ceratophysella bengtssoni from the UK
2012.01.18 © Jones, A.
Ceratophysella bengtssoni from the UK
2012.01.20 © Jones, A.
Ceratophysella bengtssoni from the UK
Short anal spines on papillae
2012.08.31 © Murray, A.
Ceratophysella bengtssoni from the UK
With egg clusters
2014.01.11 © Valentine., B
Ceratophysella bengtssoni from Belgium
Abd.4: p1>p2
2017.02.25 © Huskens, M.-L.
Ceratophysella bengtssoni from the UK
2018.11.25 © Uncle, F.
Ceratophysella bengtssoni from the UK
Furca
2018.11.25 © Uncle, F.
Ceratophysella bengtssoni from the UK
Anal spines
2018.11.25 © Uncle, F.
Ceratophysella bengtssoni from Belgium
2019.01.27 © Huskens, M.-L.
Ceratophysella bengtssoni from Belgium
Abd.6 with 2 small anal spines (*)
2019.01.27 © Huskens, M.-L.
Ceratophysella bengtssoni from the UK
Abd.4 : p1 > p2; m2 absent
2019.02.06 © Valentine, Br.
Ceratophysella bengtssoni from Belgium
2019.06.08 © Huskens, M.-L.
Ceratophysella bengtssoni from Belgium
Two small anal spines (*)
2019.06.08 © Huskens, M.-L.
Ceratophysella bengtssoni ♂ from Belgium
Manubrium distinctly bent
2019.11.04 © Huskens, M.-L.
Ceratophysella bengtssoni ♂ from Belgium
Small anal spines (*)
Manubrium with broad basis
2019.11.04 © Huskens, M.-L.
Ceratophysella bengtssoni ♂ from Belgium
Furcula and retinaculum
Collophore with extended vesicles
2019.11.04 © Huskens, M.-L.
Ceratophysella bengtssoni ♂ from Belgium
Retinaculum with 2 rami
2019.11.04 © Huskens, M.-L.
Ceratophysella bengtssoni ♂ from Belgium
Dentes short with apical swelling (*)
2019.11.04 © Huskens, M.-L.
Ceratophysella bengtssoni from the UK
Preparing for escape jump
Exposed sticky antennal + anal vesicles (*)
2021.12.25 © Valentine, Br.
Ceratophysella bengtssoni from Belgium
Feet with 1 tenent macroseta (*)
2022.01.22 © Huskens, M.-L.
Ceratophysella bengtssoni from Belgium
Footcomplex with 1 clavate sticky macroseta
2022.01.22 © Huskens, M.-L.
Ceratophysella bengtssoni from Belgium
Abd.4 m2 missing
2022.01.22 © Huskens, M.-L.



Species biclavata Park, K-H et Park, N-Y, 2006:490

Species biloba (Christiansen & Bellinger, 1980:150) Park, K-H & Park, N-Y, 2006:492

Species bispinata Loksa, 1977

Remark by Thibaud, J.-M., Schulz, H.-J. & da Gama Assalino, M.M. 2004:40,45: Insufficient description.

Remark by Dányi, L. & Traser, G. 2008:7: Species inquirenda. Maybe junior synonym of Ceratophysella silvatica Rusek, 1964.

Species boletivora (Packard, AS, 1873:30) Lippert & Butler, 1976

Species borealis Martynova, 1977

Species brevis (Christiansen, K & Bellinger, P, 1980:155) Babenko & al, 1994

Species brevisensillata Yosii, 1961

            Syn.: Achorutes pseudarmatus Pearse, 1946 =?
            Syn.: Achorutes pseudarmatus Knowlton & al., 1950 =?
            Syn.: Hypogastrura (Ceratophysella) pseudarmata Maynard, EA, 1951:46 =?
            Syn.: Hypogastrura pseudarmata Yosii, 1960
            Syn.: Hypogastrura pseudarmata Salmon, JT, 1964:225 partim
            Syn.: Hypogastrura pseudarmata O'Neill & Pedigo, 1969 =?
            Syn.: Hypogastrura sigillata Hammer =?


Species californica (Bacon, 1914) Skarzynski, D, 2007:262

Species caucasica Martynova, 1964

            Syn.: Ceratophysella stercoraria Martynova, 1964


Species cavicola (Börner, 1901) Thibaud, J-M, 1970:88, B

            Syn.: Schaefferia hystrix Handschin, 1919
            Syn.: Hypogastrura luteospina Stach, 1922


Ceratophysella cavicola from Holland
2021.03.08 © Kamsteeg, G.
Ceratophysella cavicola juv. from Holland
2021.03.08 © Kamsteeg, G.
Ceratophysella cavicola juv. from Holland
Eye with 8 ocelli
2021.03.08 © Kamsteeg, G.
Ceratophysella cavicola juv. from Holland
2 distinctly curved anal spines (*)
2021.03.08 © Kamsteeg, G.

Species citri (Bacon, 1914), s.i.

Species communis (Folsom, JW, 1898:52) Stach, 1949

Achorutes communis from Japan
After Folsom JW, 1898 Fig.1
Achorutes communis from Japan
After Folsom JW, 1898 Fig.2

            Syn.: Achorutes armatus Yosii, R, 1940:114
            Syn.: Hypogastrura armata Scott, 1961
            Syn.: Hypogastrura armata Salmon, JT, 1964:207 partim
            Syn.: Hypogastrura armata Yosii, R, 1977:143
            Syn.: Hypogastrura yuasai Yosii, 1954
            Syn.: Hypogastrura yuasai Salmon, JT, 1964:209
            Syn.: Hypogastrura yuasai Thibaud, J-M, Schulz, H-J et da Gama Assalino, MM, 2004:47
            Syn.: Ceratophysella communis f. principalis Yosii, 1960
            Syn.: Hypogastrura armata ssp. communis f. principalis Salmon, JT, 1964:209
            Syn.: Hypogastrura communis ssp. principalis Salmon, JT, 1964:209
            Syn.: Ceratophysella armata Yosii, R, 1977:143



Ceratophysella armata var. communis from Argentina
After Rapoport EH, 1959 Fig.1
Ceratophysella communis from Japan
2006 © Kitobi, M.
Ceratophysella communis from Japan
2014.01.25 © Hara, Y.
Ceratophysella communis from Taiwan
2021.11.23 © Cheng, H.-J.
Ceratophysella communis from Taiwan
Anal spines on papillae
2021.11.25 © Cheng, H.-J.
Ceratophysella communis from Taiwan
Egg cluster
2021.11.25 © Cheng, H.-J.
Ceratophysella communis from Taiwan
2021.11.26 © Cheng, H.-J.
Ceratophysella communis from Taiwan
Distinctive anal spines
on large papillae
2021.11.26 © Cheng, H.-J.
Ceratophysella communis juv. from Taiwan
Reddish form
2021.11.28 © Cheng, H.-J.
Ceratophysella communis from Taiwan
Hatchling
2021.12.03 © Cheng, H.-J.
Ceratophysella communis from Taiwan
Egg cluster
Eyes of embryo visible
2021.12.05 © Cheng, H.-J.
Ceratophysella communis from Taiwan
Yeast suspension feeding
2021.12.07 © Cheng, H.-J.

Species communis2 Denis, 1924

            Syn.: Hypogastrura (Ceratophysella) engadinensis Greenslade P & Ireson, 1986 cf


Species comosa Nakamori, T, 2013:373

Ceratophysella comosa from Japan
With cephalic spines
After Nakamori T ©, 2013 Fig.5

Species cylindrica Cassagnau, P, 1959:499

Species czelnokovi Martynova, 1978

Species czukczorum Martynova & Bondarenko, 1978

             Syn.: Hypogastrura (Ceratophysella) czuckczorum(sic) Fjellberg, A, 1984:93 ISS

Species denisana (Yosii, R, 1956:16) Skarzynski, D, [2005]:152

            Syn.: Ceratophysella denisi Yosii, 1954, nec Gisin, 1944
            Syn.: Ceratophysella denisi Yosii, R, 1956:16
            Syn.: Ceratophysella denisi Yosii, R, 1977:143
            Syn.: Ceratophysella denisi Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:155
            Syn.: Ceratophysella denisi Thibaud, J-M, Schulz, H-J et da Gama Assalino, MM, 2004:48
            Syn.: Hypogastrura (Cyclograna) loricata Yosii, 1938 in Nakamori T, 2013:372 =?
            Syn.: Hypogastrura (Cyclograna) loricata Yosii, 1970 in Nakamori T, 2013:372 =?


Species densornata (Maynard, EA, 1951:31) Skarzynski, D, 2005:2

Species denticulata (Bagnall, 1941) Stach, 1949, B

            Syn.: Achorutes armatus Oudemans, 1890
            Syn.: Achorutes scotica Carpenter & Evans, 1899=?
            Syn.: Achorutes armatus var. cuspidatus Axelson, WM, 1905:788
            Syn.: Achorutes (Ceratophysella) cuspidata Axelson, 1905
             Syn.: Achorutes armata var. cuspidate(sic) Weiner, WM, Xie, Z-J, Li, Y et Sun, X, 2019:72 ISS
            Syn.: Achorutes distinguendus Bagnall, 1941
            Syn.: Hypogastrura engadinensis Gisin, 1949
            Syn.: Ceratophysella communis ssp. exilis Yosii, 1954
            Syn.: Hypogastrura (Ceratophysella) communis ssp. exilis Yosii, R, 1956:14
            Syn.: Ceratophysella exilis Yosii, 1956
            Syn.: Ceratophysella communis ssp. exilis Yosii, R, 1977:143
            Syn.: Ceratophysella exilis Yosii, R, 1977:143
            Syn.: Hypogastrura (Ceratophysella) afghanistanensis Stach, 1963
            Syn.: Hypogastrura sp. communis-group King, Hutchinson & Greenslade P, 1976
            Syn.: Hypogastrura granulata Skarzynski, D, Smolis, A, Kovác, L et Porco, D, 2021:4 partim
            Syn.: Hypogastrura granulata Skarzynski, D, Smolis, A, Kovác, L et Porco, D, 2021:4 partim,cf


             Syn.: Ceratophysella denticula(sic) Cassagnau, P, 1962:226 ISS
Ceratophysella denticulata from France
Right eye; red = longitudinal axis
Modified after Thibaud J-M & Massoud Z, 1973 Fig.A

             Syn.: Ceratophysella denticulate(sic) Chahartaghi, M, Langel, R, Scheu, S et Ruess, L, 2005:1722 ISS
             Syn.: Hypogastrura deniculata(sic) Nakamori, T et Suzuki, A, 2005:262 ISS
Ceratophysella denticulata from the UK
2005 © Brocklehurst, K.
Ceratophysella denticulata from the UK
Footcomplex with empodium
2005 © Hopkin, S.P.
Ceratophysella denticulata from the USA
2009.03.20 © Justis, S.
 
Ceratophysella denticulata from the USA
2009.03.20 © Justis, S.
Ceratophysella denticulata from the USA
2009.03.20 © Justis, S.
Ceratophysella denticulata adult and 1st instar
After Thibaud J-M & D'Haese C, 2010 Fig.52.A

             Syn.: Ceratophysella denticulada(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:108 ISS
Ceratophysella denticulata from the UK
2012.03.31 © Nurcombe, E.
Ceratophysella denticulata from the UK
2012.03.31 © Nurcombe, E.
Ceratophysella denticulata from Holland
2012.07.15 © van Duinen, J.
Ceratophysella denticulata from Holland
2012.07.30 © van Duinen, J.
Ceratophysella denticulata from the UK
2013.02.11 © Nurcombe, E.
Ceratophysella denticulata from the UK
Abd.4: 3 setae nr midline; p1 short, p2 long. Abd.5: transvers array of 3 setae
2013.02.11 © Nurcombe, E.
Ceratophysella denticulata from the UK
Sense organ of 3rd antennal segment
2013.02.11 © Nurcombe, E.
Ceratophysella denticulata from the UK
Head
2013.02.11 © Nurcombe, E.
Ceratophysella denticulata from Holland
Abd.4: 3 setae nr midline; p1 short, p2 long. Abd.5: transvers array of 3 setae
2013.02.16 © van Duinen, J.
Ceratophysella denticulata from the UK
2013.04.01 © Nurcombe, E.
Ceratophysella denticulata from Belgium
Abd.4: p1<p2
2017.01.19 © Huskens, M.-L.
Ceratophysella denticulata from Belgium
Abd.4: p1<p2
2018.02.05 © Huskens, M.-L.
Ceratophysella denticulata from Belgium
2018.02.05 © Huskens, M.-L.
Ceratophysella denticulata from France
Exposed sticky antennal (*)+ anal (**) sacs before jump
2018.03.21 © Picard, J.
Ceratophysella denticulata from France
Exposed sticky anal sacs
2018.03.21 © Picard, J.
Ceratophysella denticulata from France
Exposed sticky antennal sacs (*)
Inflatable sacs at joint of ant.3/4
2018.03.21 © Picard, J.
Ceratophysella denticulata from France
With uplifted adomen to catch wind
Sailing on watersurface
2018.03.25 © Picard, J.
Ceratophysella denticulata from Belgium
2019.08.11 © Huskens, M.-L.
Ceratophysella denticulata from Belgium
2019.08.20 © Huskens, M.-L.
Ceratophysella denticulata from Belgium
2020.02.15 © Huskens, M.-L.
Ceratophysella denticulata from Belgium
Abd.4 with p1 < p2
2020.02.15 © Huskens, M.-L.
Ceratophysella denticulata from Belgium
Abd.5 with transverse ridge of strong cuticular tubercles
2020.02.15 © Huskens, M.-L.
Ceratophysella denticulata from Belgium
Distinctly large anal spines (*)
2020.02.15 © Huskens, M.-L.
Ceratophysella denticulata-group from the USA
2021.04.01 © Pavy, R.
Ceratophysella denticulata-group from the USA
Abd.3 with p1 < p2
2021.04.01 © Pavy, R.



Species dolsana (Lee, B-H & Kim, J-T, 1995:135) Park, K-H & Park, N-Y, 2006:489

Species duplicispinosa (Yosii, 1954) Babenko & al., 1994

            Syn.: Chinogastrura punctata Rusek, J, 1967:186
            Syn.: Chinogastrura quinidentis Jia, Shi & Chen, 2005


             Syn.: Hypogastrura (Ceratophysella) duplisispinosa(sic) Fjellberg, A, 1984:96 ISS

Species empodialis (Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB & Stebaeva, SK, 1994:142) Skarzynski, D, 2019:152

Species engadinensis (Gisin, 1949) Cassagnau, 1959

            Syn.: Ceratophysella denticulata Jordana, 1980
            Syn.: Ceratophysella denticulata Ardanaz & Jordana, 1983
            Syn.: Ceratophysella denticulata Arbea & Jordana, 1985
            Syn.: Ceratophysella denticulata Ardanaz, 1986
            Syn.: Ceratophysella denticulata Ardanaz & Jordana, 1986
            Syn.: Ceratophysella denticulata Villanueva, 1986
            Syn.: Ceratophysella (denticulata-group) denticulata Najt, J, Thibaud, J-M et Jacquemart, S (+), 1991:154 =?


Ceratophysella engadinensis from the UK
Dorsal chaetotaxy of 5th abdominal segment
2002 © Hopkin, S.P.
Ceratophysella engadinensis
2006 © Boldsystems.org
Ceratophysella engadinensis from Spain
Aggregation on surface of swimming pool
2007.05.03 © Jordana, R.
             Syn.: Ceratophysella engadiniensis(sic) Arbea, JI et Ariza, E, 2012:206 ISS
Ceratophysella engadinensis from Spain
After Arbea JI, 2018 Fig.5



Species engeli Ellis, WN, 1968:89

Ceratophysella engeli from Guatemala
After Ellis WN, 1968 Fig.1

Species falcifer Cassagnau, P, 1959:497

Species flectoseta Lin & Xia, 1983

             Syn.: Ceratophysella flectochaeta(sic) Weiner, WM, Xie, Z-J, Li, Y et Sun, X, 2019:75 lapsus ISS

Species franzi (Butschek & Gisin, 1949) Skarzynski, D, 2006:293

Species fujisana Itoh, 1985

Species gibbomucronata (Hammer, 1953) Skarzynski, D & Christiansen, K, 2008:990

Species gibbosa (Bagnall, RS, 1940:165) Stach, J, 1946:8

            Syn.: Hypogastrura (Ceratophysella) occidentalis Gisin, 1958
            Syn.: Ceratophysella azteca Yosii, 1962


             Syn.: Hypogastrura gibbosus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:47 ISS
Ceratophysella gibbosa from the UK
Dorsal chaetotaxy of 4th abdominal segment
2002 © Hopkin, S.P.
Ceratophysella gibbosa from the USA
2009.06.07 © Justis, S.
Ceratophysella gibbosa from the USA
2009.06.07 © Justis, S.
Ceratophysella gibbosa from Holland
2014.03.14 © van Duinen, J.
Det.: Berg, M.



Species glancei Hammer, 1953

Species granulata Stach, J, 1949:133

             Syn.: Ceratophysella granulate(sic) Rendos, M, Miklisová, D, Kovác, L et Mock, A, 2020:22 ISS
Ceratophysella granulata juv. from Belgium
2020.11.28 © Huskens, M.-L.
Ceratophysella granulata juv. from Belgium
Abd.4 with p1>p2
2020.11.28 © Huskens, M.-L.
Ceratophysella granulata juv. from Belgium
2020.11.28 © Huskens, M.-L.
Ceratophysella granulata juv. from Belgium
Large anal spines on distinct papillae
2020.11.28 © Huskens, M.-L.
Ceratophysella granulata juv. from Belgium
Anal spines on large papilla
Osculating at bases
2020.11.28 © Huskens, M.-L.
Ceratophysella granulata juv. from Belgium
Abd.4 with 3 fields of coarser granulation
Abd.5 with rectangular area of finer granulation
2020.11.28 © Huskens, M.-L.
Ceratophysella granulata juv. from Belgium
Unguiculus shorter than unguis
2020.11.28 © Huskens, M.-L.
Ceratophysella granulata from Belgium
Abd.4 with 3 area's of coarse granulation
2021.02.23 © Huskens, M.-L.
Ceratophysella granulata from Belgium
Ingesting fungal hyphen
2021.02.24 © Huskens, M.-L.
Ceratophysella granulata from Belgium
Posterior margin of abd.5 marked with coarse granulae
2021.02.25 © Huskens, M.-L.
Ceratophysella granulata from Belgium
2021.03.05 © Huskens, M.-L.
Ceratophysella granulata from Belgium
Abd.4 with p1 > p2
2021.03.05 © Huskens, M.-L.
Ceratophysella granulata from Belgium
Post-antennal organ (PAO) with several lobes
2021.03.05 © Huskens, M.-L.
Ceratophysella granulata from Belgium
Abd.5 with narrow field of finer granulation
Large anal spines
2021.03.08 © Huskens, M.-L.



Species granulifera Yosii, 1962

Species guthriei (Folsom, JW, 1916:489) Stach, J, 1946:8

            Syn.: Achorutes longispinus Guthrie, JE, 1903:90


Ceratophysella guthriei from the USA
2017.09.09 © Johnson, S.T.

Species hermosa (Wray, 1953) Skarzynski, D & Christiansen, K, 2008:990

            Syn.: Spinifacies oregonensis Scott, 1963 =?


Ceratophysella hermosa from the USA
After Bernard EC, 2015 Fig.1A

Species hiawatha (Yosii, 1962) Skarzynski, D & Christiansen, K, 2008:990

Species horrida (Yosii, 1960) Skarzynski, D & Christiansen, K, 2008:990

Species hystrix (Handschin, E, 1924:93) Skarzynski, D, 2007:261

Species impedita Skarzynski, D, 2002:171

Species indica Salmon, 1956

Species indovaria Salmon, 1970

Ceratophysella indovaria from India
After Yadav RS et al, 2018 Fig.1

Species ionescui (Börner, 1922)

Species isabellae Fjellberg, 1985

Species jondavi (Wray, 1946) Salmon, 1964

             Syn.: Ceratophysella jondaui(sic) Salmon, 1964 ISS

Species jonescoi (Bonet, F, 1930:125, s.d.) Thibaud, J-M, Schulz, H-J & da Gama Assalino, MM, 2004:41, s.d.

            Syn.: Hypogastrura cavicola Stach, J, 1949:30
            Syn.: Hypogastrura cavicola Bonet, F, 1930:125 nec Börner


Species kapoviensis (Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB & Stebaeva, SK, 1994:185) Skarzynski, D, Kaprus, I & Shrubovych, J, 2001:413

Species katraensis (Tyagi & Baijal, 1982) Mandal, GP & Arbea, J, 2019:279

Species kennethi Skarzynski, D, 2007:139

Species kolchidica (Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB & Stebaeva, SK, 1994:137) Skarzynski, D, 2003:4

Species krafti (Scott, 1962) Skarzynski, D & Christiansen, K, 2008:990

Species kutyrevae Babenko, 1994

Species laricis Martynova, 1977

Species lawrencei (Gisin, H, 1963:97)

            Syn.: Hypogastrura gibbosa Gisin, 1949 nec Bagnall, 1940 cf
            Syn.: Hypogastrura gibbosa Gisin, H, 1963:97,cf


Species liguladorsi (Lee, 1974) Huang & al, 1992

            Syn.: Hypogastrura (Ceratophysella) tergilobata Yoshii, R et Suhardjono, YR, 1989:24
            Syn.: Hypogastrura (Ceratophysella) tergilobata Suma, 2003
            Syn.: Hypogastrura (Ceratophysella) tergilobata Suma & Karasawa, 2005
            Syn.: Ceratophysella tergilobata Jantarit & al, 2016 =?
            Syn.: Hypogastrura (Ceratophysella) tergilobata Nakamori, T, Ichisawa, K, Pham, HN-D et Terashima, Y, 2020:12


Ceratophysella liguladorsi from Japan
Modified after Nakamori T & al, 2020 Fig.a+b
Ceratophysella liguladorsi from Japan
Abd.5 with central projection
Modified after Nakamori T & al, 2020 Fig.b

Species lobata Babenko, A et Skarzynski, D, 2011:260

            Syn.: Hypogastrura (Ceratophysella) brevisensillata Fjellberg, 1985
            Syn.: Hypogastrura (Ceratophysella) brevisensillata Babenko & al., 1994

Species longispina (Tullberg, T, 1877:37) Stach, J, 1946:8

            Syn.: Hypogastrura longispina f. trispina Carpenter, 1927
            Syn.: Hypogastrura hirsuta Valpas, 1967
            Syn.: Ceratophysella arctica Martynova, Gorodkov & Tschelnokov, 1973


             Syn.: Achorutes longispinis(sic) Reuter, OM, 1901:130 ISS
             Syn.: Hypogastrura longispinus(sic) Poole in Poole & Gentili, 1997 ISS
Ceratophysella longispina from Norway
2011 © Fjellberg, A.



Species loricata (Yosii, 1960) Skarzynski, D & Christiansen, K, 2008:990

Species lucifuga (Packard, 1886)

            Syn.: Lipura? lucifugus(sic) Packard, 1886 IOS

Species macrocantha Stach, J, 1946:4

            Syn.: Ceratophysella (armata-group) recta Skarzynski, D, 2019:152 =?
            Syn.: Ceratophysella (armata-group) attenuata Skarzynski, D, 2019:152 nec Cassagnau, 1959 =?


Species macrospinata (Maynard, 1951, s.i.) Maynard, 1951

Species maheuxi (Butler, 1966) Skidmore, RE, 1995:48

Species maya Yosii, 1962

Species mediolobata Nakamori T in Nakamori, T, Ichisawa, K, Pham, HN-D et Terashima, Y, 2020:13

Ceratophysella mediolobata from Japan
After Nakamori T & al, 2020 Fig.e+f
Ceratophysella mediolobata from Japan
Abd.5 with central projection
Modified after Nakamori T & al, 2020 Fig.f

            Syn.: Hypogastrura (Ceratophysella) tergilobata Suma, 1985
            Syn.: Ceratophysella tergilobata Suma, 2017
            Syn.: Ceratophysella liguladorsi Babenko, 1994



Species michalinae Skarzynski, D, 2005:1

Species microchaeta (Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:130)

Species moroni Villalobos & Palacios-Vargas, 1986:7

Species morula Deharveng & Bourgeois, 1991

Species mosambicensis Cardoso, MA, 1973:7

            Syn.: Hypogastrura moçambicensis(sic) Cardoso, MA, 1973:7 IOS
            Syn.: Hypogastrura moçambicensis(sic) Thibaud, J-M, 2013:239 IOS

Species mosquensis (Becker, 1905)

            Syn.: ... armatus Becker, 1902
            Syn.: Ceratophysella monstruosa Gisin, 1949


Species multilobata Skarzynski, D, 2006:1

Species mucronata Deharveng & Bourgeois, 1991

Species najtae Villalobos & Palacios-Vargas, 1986:5

Species narkandae (Baijal, HN, 1955:531) Mandal, GP & Arbea, JI, 2014:133

Achorutes narkandae from the Himalaya
After Baijal HN, 1955 p.532 fig.1.

Species nataliae Cipola NG, Bellini BC et Palacios-Vargas JG in Palacios-Vargas, JG, Cipola, NG et Bellini, BC, 2017:12

Ceratophysella nataliae from Brazil
After Palacios-Vargas JG, Cipola NG & Bellini BC, 2017 Fig.19

Species neomeridionalis (Nosek, J & Cervek, S, 1970:207) Skarzynski, D & Smolis, A, 2002:304

            Syn.: Hypogastrura (Ceratophysella) meridionalis Nosek & Cervek, 1967 nec Steiner, 1955
            Syn.: Hypogastrura (Ceratophysella) meridionalis Nosek, J et Cervek, S, 1970:207


Species norensis Cassagnau, P, 1964:414

             Syn.: Ceratophysella (communis-group) norica(sic) Ellis, WN, 1968:92 lapsus ISS

Species orizabae Yosii, 1962

Species palustris Martynova, 1978

Species paraliguladorsi Nguyen, 2001

Species pauciseta Greenslade, Ireson & Skarzynski, 2014

Species pecki (Christiansen & Bellinger, 1980:163) Skidmore, 1995:48

Species penicillifer Cassagnau, P, 1964:468

             Syn.: Ceratophysella (armata-group) penicilifer(sic) Skarzynski, D, 2019:152 ISS

Species pilosa (Yosii, R, 1956:15) Skarzynski, D & Christiansen, K, 2008:990

Species planipila Yosii, 1966

Species platyna Park, K-H et Park, N-Y, 2006:490

Species pratorum (Packard, AS, 1873:493:31) Villalobos & Palacios-Vargas, 1986

Ceratophysella pratorum from the USA
2007.04.30 © Murray, T.
Ceratophysella pratorum from the USA
2007.04.30 © Murray, T.

Species proserpinae (Yosii, R, 1956:14) Ellis, WN, 1968:92

Species quinquesetosa Gisin, 1958

Species recta Cassagnau, P, 1959:497

Species richardi Skarzynski, D et Christiansen, K, 2008:990

Species robustiseta (Skarzynski, D & Smolis, A, 2006:299) Skarzynski, D, 2007:137

Species rogerarlei Palacios-Vargas JG, Bellini BC et Cipola NG in Palacios-Vargas, JG, Cipola, NG et Bellini, BC, 2017:3

Ceratophysella rogerarlei from Brazil
After Palacios-Vargas JG, Cipola NG & Bellini BC, 2017 Fig.1

Species sakayorii Tamura, 1997

             Syn.: Ceratophysella sakayori(sic) Nakamori, T, 2013:377 ISS

Species scotica (Carpenter & Evans, 1899) Ellis, WN, 1967:95

            Syn.: Achorutes armatus var. cuspidatus Axelson, WM, 1905:788
             Syn.: Achorutes cuspidata(sic) Bagnall, RS, 1940:164 ISS
             Syn.: Achorutes armatus ssp. cuspidata(sic) Salmon, JT, 1964:209 ISS
            Syn.: Achorutes filiformis Wahlgren, 1906
            Syn.: Ceratophysella gotoi Murphy, 1960
            Syn.: Ceratophysella macrocantha Skarzynski, D, 2019:151 =?


             Syn.: Hypogastrura denticulata-scotica(sic) Nelson, JM, 1981:65,cf ISS
Ceratophysella scotica from the UK
Footcomplex with claw and empodium
2002 © Hopkin, S.P.



Species scotti Yosii, 1962

Species sedecimocellata (Yosii, 1962) Palacios-Vargas, 1983

            Syn.: Ceratophysella duodecimocellata f. 16-ocellata(sic) Yosii, 1962 IOS
             Syn.: Ceratophysella 16-ocellata(sic) Jordana, R, Baquero, E, Ledesma, E, Sendra, A et Ortuño, VM, 2020:89 ISS



Species semnacantha (Börner in Schille, 1912, n.n.) Bellinger, PF & Ellis, WN, 1976:274

Remark by Bellinger, P.F. & Ellis, W.N. 1976:274: Manuscript name.

Species sextensis Cassagnau, 1968

Species sibirica Martynova, 1974:14

Species sigillata (Uzel, 1891) Stach, J, 1949:147

            Syn.: Podura (deux crochets à l'extrémité du corps) rufescens Nicolet, H, 1842:57
Podura rufescens from Germany
After Nicolet, H, 1842 Pl.5 fig.7.

            Syn.: Achorutes larvatus Gervais in Walckenaër, 1844
            Syn.: Achorutes Schupplii(sic) Haller, 1880
             Syn.: Achorutes Schupplii(sic) Carl, J, 1899:289 ISS
             Syn.: Achorutes Schupplii(sic) Absolon, K, 1901:579 ISS
            Syn.: Achorutes pluvialis Vogler, 1893
            Syn.: Achorutes navicularis Schött, H, 1893:83
            Syn.: Achorutes pseudarmatus Folsom, JW, 1916:490
             Syn.: Neogastrura pseudarmatus(sic) Stach, J, 1949:132 ISS
             Syn.: Hypogastrura (Ceratophysella) pseudoarmata(sic) Voegtlin, 1982 ISS
             Syn.: Hypogastrura pseudoarmata(sic) Parsons & al, 1991 ISS
             Syn.: Ceratophysella pseudoarmata(sic) Zettel, J, Zettel, U et Egger, B, 2000:41 ISS
            Syn.: Hypogastrura minuta Denis, J-R, 1931:80



Ceratophysella sigillata from Austria
Mass migration
After Christian E & Meyer E, 1997 Abb.1
Ceratophysella sigillata from Germany
2006 © Kohl, F.
Ceratophysella sigillata from Switzerland
Purple subadults and blue-grey adults
After Zettel J & Zettel U, 2008 Abb.3
Ceratophysella sigillata from Switzerland
Climbing a tree using eversible sticky antennal and anal vesicles
After Zettel J & Zettel U, 2008 Abb.17
Ceratophysella sigillata from Germany
2013.03.dd © Böhling, N.
Ceratophysella sigillata from Germany
With everted antennal vesicles (*)
2013.03.dd © Böhling, N.
Ceratophysella sigillata from Germany
Abd.4 seta p1 long; anal spines (*) not terminal
2016.02.07 © Melzer, L.

Species silvatica Rusek, 1964

            Syn.: Ceratophysella bispinata Loksa, 1977 =?
            Syn.: Hypogastrura granulata Skarzynski, D, Smolis, A, Kovác, L et Porco, D, 2021:4 partim
            Syn.: Ceratophysella granulata Skarzynski, D, Smolis, A, Kovác, L et Porco, D, 2021:4


             Syn.: Ceratophysella sylvatica(sic) Fiera, C, 2014:229 ISS
             Syn.: Ceratophysella sylvatica(sic) Fiera, C, 2014:231 ISS

Species sinensis Stach, J, 1964:5

Species sinetertiaseta (Lee, 1974) Park, K-H & Park, N-Y, 2006:489

Species skarzynskii Weiner WM et Sun X in Weiner, WM, Xie, Z-J, Li, Y et Sun, X, 2019:69

             Syn.: Ceratophysella skarzynski(sic) Weiner, WM, Xie, Z-J, Li, Y et Sun, X, 2019:71 ISS

Species snideri (Skarzynski, D, 2007:262) Skarzynski, D & Christiansen, K, 2008:990

Species sphagni (Becker, 1905)

Species stachi Skarzynski D, Smolis A et Porco D in Skarzynski, D, Smolis, A, Kovác, L et Porco, D, 2021:5

            Syn.: Ceratophysella granulata Fjellberg, 1998
            Syn.: Ceratophysella granulata Skarzynski, D, Smolis, A, Kovác, L et Porco, D, 2021:5


Species stercoraria (Stach, 1963) Martynova, 1964

Ceratophysella stercoraria from Poland
After Skarzynski D, 2000 Fig.1

Species subhorrida (Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB & Stebaeva, SK, 1994:194) Skarzynski, D & Christiansen, K, 2008:990

Species succinea (Gisin, 1949:387) Ellis, WN, 1968:89

Ceratophysella succinea from the USA
2007.10.22 © McClarin, J.
Ceratophysella succinea from the USA
2007.10.22 © McClarin, J.
Ceratophysella succinea from the UK
2012.08.23 © Murray, A.
Ceratophysella succinea from the UK
Eyepatch with 8 ocelli
2012.08.23 © Murray, A.
Ceratophysella succinea from the UK
Anal spines
2012.08.23 © Murray, A.
Ceratophysella succinea from France
Abd.4: seta p1<p2; m1 present
2017.10.06 © Picard, J.
Ceratophysella succinea from France
Yellowish with brown spots
2017.10.06 © Picard, J.
Ceratophysella succinea from France
2017.10.06 © Picard, J.
Ceratophysella succinea from Belgium
2017.11.08 © Huskens, M.-L.
Ceratophysella succinea from Belgium
Abd.5 with transverse band of larger cuticular tubercles
2017.11.08 © Huskens, M.-L.
Ceratophysella succinea from Belgium
2019.06.14 © Huskens, M.-L.
Ceratophysella succinea from Belgium
Abd.4 p2 > p1
Abd.5 with transverse cuticular band (*)
2019.06.14 © Huskens, M.-L.
Ceratophysella succinea from Belgium
Distinctly large anal spines
Abd.5 with transverse cuticular band
2019.06.14 © Huskens, M.-L.

Species taiguensis Jia, J, Skarzynski, D et Li, Y, 2010:58

Species tergilobata (Cassagnau, P, 1954:239) Deharveng, L, 1983:21

Ceratophysella tergilobata from Spain
With tergal lob at 5th abdominal segment (*)
After Arbea JI, 2013 Fig.2

Species toemoesvaryi (Loksa, I, 1964:83)

            Syn.: Hypogastrura (Ceratophysella) tömösvaryi(sic) Loksa, I, 1964:83 IOS


             Syn.: Hypogastrura tömösvaryi(sic) Loksa, 1966 ISS
             Syn.: Ceratophysella tömösvaryi(sic) Skarzynski, D, 2002:172 ISS
             Syn.: Ceratophysella tömösvaryi(sic) Thibaud, J-M, Schulz, H-J et da Gama Assalino, MM, 2004:66 ISS
Remark by Thibaud, J.-M., Schulz, H.-J. & da Gama Assalino, M.M. 2004:40,66: Insufficient description.





Species tolteca Yosii, 1962

Species troglodytes (Yosii, R, 1956:11)

            Syn.: Hypogastrura troglodites(sic) Yosii, R, 1956:11 IOS
             Syn.: Hypogastrura (Ceratophysella) troglodites(sic) Yosii, R, 1977:144 ISS
             Syn.: Ceratophysella troglodites(sic) Thibaud, J-M, Schulz, H-J et da Gama Assalino, MM, 2004:66 ISS
             Syn.: Ceratophysella troglodites(sic) Skarzynski, D, 2007:275 ISS
             Syn.: Ceratophysella troglodites(sic) Nakamori, T, 2013:377 ISS



Species tuberculata Cassagnau, P, 1959:494

Species tupamara Palacios-Vargas, JG et Bocanegra, T, 2012:418

Ceratophysella tupamara from Peru
After Palacios-Vargas JG & Bocanegra T, 2012 Fig.1+7

Species vargovychi Skarzynski, D, Kaprus, I et Shrubovych, J, 2001:411

Species varians (Stach, J, 1967:396) Dallai, R, Malatesta, E & Ramellini, P in Minelli, A, Ruffo, S & La Posta, S, 1995:4

            Syn.: Hypogastrura melitensis Stach, J, 1967:394 lapsus


Species virga (Christiansen, K & Bellinger, P, 1980:183) Skarzynski, D & Christiansen, K, 2008:990

            Syn.: Spinifacies oregonensis Scott, HG, 1963:108
Spinifacies oregonensis from the USA
After Scott HG, 1963 Fig.1 upper
Spinifacies oregonensis from the USA
After Scott HG, 1963 Fig.1 lower

             Syn.: Spinifacies orogonensis(sic) Roback, 1981 ISS



Species vulgaris (Yosii, 1960) Skarzynski, D & Christiansen, K, 2008:990

Species wallmoi (Fjellberg, 1985) Skarzynski, D & Christiansen, K, 2008:990

             Syn.: Mitchellania vallmoi(sic) Skarzynski, D, 2007:264 ISS

Species wrayia (Uchida, H et Tamura, H, 1968:341)

             Syn.: Ceratophysella wraya(sic) Skarzynski, D, 2007:261 ISS

Species xiaoi (Tamura in Tamura & Zhao, 1998) Jia, J, Skarzynski, D & Li, Y, 2010:62, s.i.

Species yakushimana Yosii, R, 1965:2

Species yinae (Yue & Fu, 2000) Jia, J, Skarzynski, D & Li, Y, 2010:62

Species zhangi (Zhao in Tamura & Zhao, 1998) Jia, J, Skarzynski, D & Li, Y, 2010:62, s.i.


Genus Schaefferia Absolon, K, 1900:265 key

F
  Syn.: Poduridae Schäfferia(sic) Absolon, K, 1900:265 IOS

  Syn.: Achorutidae Achorutini(sic) Börner, C, 1901:13 partim IOS
  Syn.: Neanurini Octomma Willem, V, 1902:279
  Syn.: ... Octama(sic) Pomorski & Skarzynski, 1992 ISS
  Syn.: ... Xenyllodes Womersley, 1930, nec Axelson, 1903
  Syn.: ... Spelaeogastrura Bonel, 1945
  Syn.: ... Typhlogastrura Poole, RW in Poole, RW et Gentili, P, 1997:49


  Syn.: Achorutini(sic) Schäfferia(sic) Börner, C, 1901:13 ISS
Schaefferia sp. from the USA
2007.10.13 © McClarin, J.


  Syn.: Collembola Schaeffer(sic) Suetsugu, N, Satoh, F et Koike, T, 2010:58 ISS
Schaefferia sp. from Holland
2013.11.08 © van Duinen, J.
Schaefferia sp. from Holland
2015.11.14 © van Leeuwen, J.

  Syn.: Collembola Schaeffaria(sic) Raymond-Léonard, LJ, Gravel, D et Handa, IT, 2019:5 ISS




Species ariegica Cassagnau, P, 1959:70

            Syn.: Schaefferia emucronata ssp. quadrioculata Jordana, R, 1981:3
Schaefferia emucronata ssp. quadrioculata from Spain
After Jordana R, 1981 Lam.I Fig.1


            Syn.: Schaefferia emucronata Arbea & Jordana, 1985



Schaefferia ariegica from France
Right eye; red = longitudinal axis
Modified after Thibaud J-M & Massoud Z, 1973 Fig.G
Schaefferia ariegica from France
2006 © Boldsystems.org

Species baschkirica Kniss, 1985

            Syn.: Schaefferia baschkirica ssp. casteri Kniss, 1985


Species bidentata (Cassagnau, P, 1954:68) Gisin, H, 1960:55

Species babenkoi Jordana R et Baquero E in Jordana, R, Baquero, E, Ledesma, E, Sendra, A et Ortuño, VM, 2020:90

            Syn.: Schaefferia vandalica Babenko & al, 1994


Species bermani Babenko, 2019

Species canigouensis Deharveng & Thibaud, 1980

Species cheoha Wray, DL, 1963:103

Species coeca Cassagnau, P, 1959:71

Schaefferia coeca from France
Right eye
After Thibaud J-M & Massoud Z, 1973 Fig.H
             Syn.: Schaefferia caeca(sic) Palacios-Vargas & Thibaud, 1985 ISS
             Syn.: Schaefferia caeca(sic) Bernard, EC, 2015:321 ISS


Species czernovi (Martynova, 1978) Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB & Stebaeva, SK, 1994:214

             Syn.: Schaefferia chernovi(sic) Skarzynski, D, 2007:204 ISS

Species decemoculata (Stach, 1939) Thibaud, J-M, 1970:88

Schaefferia decemoculata from Europe
Right eye; red = longitudinal axis
Modified after Thibaud J-M & Massoud Z, 1973 Fig.Cc
Schaefferia decemoculata from Europe
Right eye; red = longitudinal axis; ab. with 4 ocelli
Modified after Thibaud J-M & Massoud Z, 1973 Fig.Cd
Schaefferia decemoculata from Europe
Right eye; red = longitudinal axis; ab. with 6 ocelli
Modified after Thibaud J-M & Massoud Z, 1973 Fig.Cb
Schaefferia decemoculata from Europe
Right eye; red = longitudinal axis; ab. with 6 ocelli
Modified after Thibaud J-M & Massoud Z, 1973 Fig.Ca
Schaefferia emucronata decemoculata from Japan
After Suetsugu N, Satoh F & Koike T, 2010 Photo1
Schaefferia decemoculata from France
2022.01.28 © Usami, R.
Schaefferia decemoculata from France
Anal spines on papillae
2022.01.28 © Usami, R.
Schaefferia decemoculata from France
2022.02.09 © Usami, R.
Schaefferia decemoculata from France
Eye with 5 ocelli
2022.02.09 © Usami, R.

Species deharvengi (Jordana, R & Arbea, JI, 1990:167) Thibaud, J-M, Schulz, H-J & da Gama Assalino, MM, 2004:176

            Syn.: Ceratophysella duodecimoculata Deharveng, 1976


             Syn.: Schaefferia dehanvergi(sic) Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:47 ISS

Species duodecimocellata Bonet, 1945

            Syn.: Ceratophysella duodecimocellata f. 16-ocellata(sic) Yosii, 1962 IOS
            Syn.: Ceratophysella duodecimocellata f. 16-ocellata(sic) Jordana, R, Baquero, E, Ledesma, E, Sendra, A et Ortuño, VM, 2020:89 IOS
            Syn.: Schaefferia cheoha Thibaud & al, 2004
            Syn.: Schaefferia vandalica Thibaud & al, 2004


Schaefferia duodecimocellata from Mexico
Right eye; red = longitudinal axis
Modified after Thibaud J-M & Massoud Z, 1973 Fig.B
Schaefferia duodecimocellata from the USA
2009.03.28 © Justis, S.
Schaefferia duodecimocellata from the USA
2009.12.01 © Justis, S.
With 6+6 eyes
Schaefferia duodecimocellata from the USA
2009.12.01 © Justis, S.



Species duodecimoculata (Steiner, 1955) Thibaud, J-M, Schulz, H-J & da Gama Assalino, MM, 2004:177

Species elegans (Jordana, R & Arbea, JI in Ramos, MA & al, 1997:111) Thibaud, J-M, Schulz, H-J & da Gama Assalino, MM, 2004:177

            Syn.: Hypogastrura cassagnaui Salmon, JT, 1964:211


Species emucronata Absolon, K, 1900:266, t.t., B

Schaefferia emucronata from Europe
Right eye; red = longitudinal axis
Modified after Thibaud J-M & Massoud Z, 1973 Fig.E
Schaefferia emucronata from the UK
2013.01.08 © Murray, A.
Schaefferia emucronata from the UK
Reduced composed eye
2013.01.08 © Murray, A.
Schaefferia emucronata from the UK
Prominent anal spines
2013.01.08 © Murray, A.
Schaefferia emucronata from the UK
Strongly reduced furca
2013.01.08 © Murray, A.
Schaefferia emucronata from the UK
Eyepatch with at most 4 ocelli
2013.05.13 © Murray, A.
Schaefferia emucronata juv. from the UK
Eyepatch with 4 ocelli: 3+1
2013.11.26 © Nurcombe, E.
Schaefferia emucronata from the UK
Reduced furca
2019.02.27 © Smart, J.

Species fjellbergi Jordana R et Baquero E in Jordana, R, Baquero, E, Ledesma, E, Sendra, A et Ortuño, VM, 2020:90

            Syn.: Schaefferia duodecimocellata Fjellberg, 1985


Species fukugakuchiana (Yosii, R, 1956:13) Thibaud, J-M, Schulz, H-J & da Gama Assalino, MM, 2004:178

Species guerrerensis (Bonet, 1945) Thibaud, 1972

            Syn.: Spelaeogastrura guerrerense(sic) Bonet, 1945 IOS


             Syn.: Schaefferia guerrerense(sic) Thibaud, 1972 ISS
             Syn.: Schaefferia guerrerense(sic) Thibaud, J-M et Massoud, Z, 1973:162 ISS
             Syn.: Schaefferia guerrerense(sic) Hoffmann et al., 1980 ISS
             Syn.: Schaefferia (Schaefferia) guerrerense(sic) Christiansen & Bellinger, [1992] ISS
             Syn.: Schaefferia guerrerense(sic) Palacios-Vargas, JG, 2006:9 ISS
             Syn.: Schaefferia guerrerense(sic) Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:43 ISS



Species hubbardi Thibaud, 1995

Species jarae (Jordana, R & Arbea, JI, 1990:168) Thibaud, J-M, Schulz, H-J & da Gama Assalino, MM, 2004:179

            Syn.: Hypogastrura (Ceratophysella) duodecimoculata Selga, 1971, partim
            Syn.: Hypogastrura (Ceratophysella) elegans Selga, 1971


Species kitakamiana Yosii, 1991

Species lindbergi da Gama, 1963

            Syn.: Schaefferia emucronata ssp. emucronata Jordana & Beruete, 1983
            Syn.: Schaefferia emucronata Jordana & Beruete, 1983


Species maxima Deharveng & Thibaud, 1980

Species oaxacana Palacios-Vargas, JG et Thibaud, J-M, 1985:5

Species oculea Babenko, AB, 1999:887

Species pouadensis Delamare Deboutteville, 1945

Schaefferia pouadensis from France
Right eye; red = longitudinal axis
Modified after Thibaud J-M & Massoud Z, 1973 Fig.F

Species profundissima Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:41

Species quadrioculata (Stach, 1939) Thibaud, J-M, 1970:88

Species quinqueoculata (Yosii, R, 1956:12) Yoshii, 1991

            Syn.: Typhlogastrura shtanchaevae Abdurakhmanov & Babenko, 1991


Species raulmugnizi Palacios-Vargas, JG et Castaño-Meneses, G, 2008:123

Schaefferia raulmugnizi from Mexico
After Palacios-Vargas JG & Castaño-Meneses G, 2008 Fig.1a2a
             Syn.: Schaefferia raulmugnezi(sic) Palacios-Vargas, JG et Castaño-Meneses, G, 2008:124 ISS



Species scossirolii Dallai, R et Sabatini, MA, 1981:196

             Syn.: Schaefferia scossiroli(sic) Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:46 ISS

Species sendrai Jordana R et Baquero E in Jordana, R, Baquero, E, Ledesma, E, Sendra, A et Ortuño, VM, 2020:87

Schaefferia sendrai from Spain
Modified after Jordana R & Baquero E in Jordana R & al, 2020 Fig.4I

Species sexoculata (Gisin, 1947) Cassagnau, P, 1973:127

Species subcoeca Deharveng & Thibaud, 1980

             Syn.: Schaefferia subcaeca Palacios-Vargas & Thibaud, 1985 ISS

Species vandalica (Jordana, R & Arbea, JI, 1990:172) Jordana R & Baquero E in Jordana, R, Baquero, E, Reboleira, S & Sendra, A, 2012:46

Species vandalica Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:213

Species willemi (Bonet, F, 1930:125) Denis, J-R, 1936:82, B

            Syn.: Octomma longispina Willem, V, 1902:279 nec Tullberg, 1876,B
            Syn.: Octomma longispina Bonet, F, 1930:125 nec Tulberg, 1876
            Syn.: Xenyllodes cavernicola Womersley, 1930
            Syn.: Hypogastrura octoculata Womersley, 1930


             Syn.: Schaefferia Willemi(sic) Denis, J-R, 1936:82 ISS
             Syn.: Hypogastrura (Schäfferia(sic)) Willemi(sic) Denis, J-R, 1937:2,B ISS
             Syn.: Hypogastrura (Schaefferia) Willemi(sic) Leruth, R, 1939:197,B ISS
             Syn.: Schaefferia willema(sic) Thibaud, J-M et Massoud, Z, 1973:162 ISS
Schaefferia willemi from Europe
Right eye; red = longitudinal axis
Modified after Thibaud J-M & Massoud Z, 1973 Fig.D
Schaefferia willemi from Belgium
2004 © Janssens, F.





Genus Typhlogastrura Bonet, F, 1930:126 key

F
  Syn.: ... Achorutes Carl, 1906 partim
  Syn.: Hypogastrurinae Achorutes Bonet, F, 1930:126
  Syn.: ... Hypogastrura Handschin, 1925 partim
  Syn.: Hypogastrurinae Hypogastrura Bonet, F, 1930:126


Typhlogastrura sp. from Russia
2006 © Boldsystems.org

Species alabamensis Thibaud, 1975

Species asymmetrica Christiansen, KA et Wang, H, 2006:90

Species atlantea (Gisin, 1951) Thibaud, J-M & Massoud, Z, 1973:164

Species breuili Thibaud, 1967

Species christianseni Thibaud, 1975

Species elsarzolae Palacios-Vargas, JG et Thibaud, J-M, 1997:324

Species fousheensis Christiansen, KA et Wang, H, 2006:91

Species helleri Christiansen, KA et Wang, H, 2006:92

Species korenevskyi Babenko, 1987

             Syn.: Typhlogastrura korenevski(sic) Christiansen, KA et Wang, H, 2006:85 ISS
Typhlogastrura korenevskyi from Russia
2006 © Boldsystems.org



Species mendizabali (Bonet, F, 1930:127) Ellis, WN & Bellinger, PF, 1973:58, t.t.

Hypogastrura (Typhlogastrura) mendizabali from Spain
After Bonet F, 1930 Fig.1

            Syn.: Hypogastrura (Typhlogastrur) mendizabali ssp. sensibilis Bonet, F, 1930:130
            Syn.: Schaefferia (Typhlogastrura) mendizabali ssp. sensibilis Gisin, H, 1960:54



Species morozovi Babenko, 1987

Species preobrazhenskyi Babenko, 1987

Species shtanchevae Abdurakhmanov & Babenko, 1991

Species steinmanni Christiansen, KA et Wang, H, 2006:93

Typhlogastrura steinmanni
Det. Steinmann, D.
2006 © Scott, V.
Typhlogastrura steinmanni
Det. Steinmann, D.
2006 © Scott, V.

Species thibaudi Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:228

Species topali (Loksa & Bogojevic, 1967) Thibaud, J-M & Massoud, Z, 1973:165

Typhlogastrura topali from Croatia
2010.11.19 © Bedek, J.

Species unica Christiansen, KA et Wang, H, 2006:94

Species valentini Thibaud, 1996

Species veracruzana Palacios-Vargas, JG et Thibaud, J-M, 1985:7


Genus Bonetogastrura Thibaud, 1974

F
Species balazuci (Delamare Deboutteville, 1951) Thibaud, 1975, t.t.

Bonetogastrura balazuci from France
After Thibaud J-M & D'Haese C, 2010 Fig.31.2
Bonetogastrura balazuci adult and 1st instar
After Thibaud J-M & D'Haese C, 2010 Fig.52.B

Species delhezi (Stomp & Thibaud, 1974)

Species nivalis (Martynova, 1973)

Species soulensis Thibaud, 1975

Species spelicola (Gisin, H, 1964:675) Christian, 1987

Species subterranea (Carl, 1906) Radu & Rogojanu, 1972

            Syn.: Hypogastrura cavernarum(lapsus) Handschin, E in Dahl, F, 1929:26 nn


             Syn.: Hypogastrura subterrannea(sic) Obenberger, J, 1955:88 ISS
Schaefferia variabilis from Alaska
After Christiansen, K., 1951 Pl.11 Fig.7.


Species variabilis (Christiansen, 1951:125) Fjellberg, A, 1984:91



Genus Schoettella Schäffer, 1896 key

F
  Syn.: ... Schöttella(sic) Schäffer, 1896 IOS
  Syn.: ... Schöttella(sic) Börner, 1901 ISS
  Syn.: Achorutes Schöttella(sic) Axelson, 1905 ISS
  Syn.: Hypogastrura Schöttella(sic) Linnaniemi, 1912 ISS
  Syn.: Achorutes Schöttella(sic) Folsom, JW, 1916:494 ISS
  Syn.: Collembola Shottella(sic) Denis, J-R, 1924:216 ISS
  Syn.: Collembola Schottella(sic) Denis, J-R, 1924:220 ISS
  Syn.: Hypogastrura Schöttella(sic) Bonet, F, 1930:131 ISS
  Syn.: Hypogastrura Schöttella(sic) Bonet, F, 1930:132 ISS
  Syn.: Hypogastrura Schötella(sic) Bonet, F, 1930:137 ISS
  Syn.: Hypogastrura Schöttella(sic) Bonet, F, 1930:137 ISS
  Syn.: ... Schöttella(sic) Bonet, 1931 ISS
  Syn.: ... Schöttella(sic) Denis, 1931 ISS
  Syn.: Collembola Schottella(sic) Denis, J-R, 1933:318 ISS
  Syn.: Hypogastruridae Schottella(sic) Snider, RJ, 1967:195 ISS
  Syn.: Hypogastruridae Schottella(sic) Díaz, A et Palacios-Vargas, JG, 1983:84 ISS
  Syn.: ... Schoetella(sic) Skidmore, RE, 1995:48 ISS
  Syn.: Hypogastruridae Schoetella(sic) Busmachiu, G, Popa, I et Weiner, WM, 2014:551 ISS
  Syn.: Hypogastruridae Schotella(sic) Glime, JM, 2015:3 ISS
  Syn.: Hypogastruridae Schoetella(sic) Özata, MA, Sevgili, H et Kaprus, IJ, 2017:237 ISS
  Syn.: Hypogastruridae Schoetella(sic) Palacios-Vargas, JG, Castaño-Meneses, G et Valladares, L, 2021:156 ISS

Subgenus Schoettella s.s.



Species alba Folsom, JW, 1932:54

Species andina Díaz & Palacios-Vargas, 1983:82

Species celiae Fernandes, LH et de Mendonça, MC, 1998:453

Species distincta (Denis, 1931) Bonet, 1931

            Syn.: Hypogastrura (Schoettella) denisi Gisin, 1944


Species glasgowi (Folsom, JW, 1916:494) Bonet, 1931

            Syn.: Achorutes magnoliana Wray, 1952


             Syn.: Schoettella glascowi(sic) Salmon, JT, 1964:200 ISS
Schoettella glasgowi from the USA
After Bernard EC, 2015 Fig.13A



Species janiae Palacios-Vargas, JG, 1979:63

             Syn.: Schoettella janie(sic) Fernandes, LH et de Mendonça, MC, 1998:453 ISS

Species neomexicana (Scott, 1961)

Species novajaniae Palacios-Vargas, JG et Castaño-Meneses, G, 1998:779

             Syn.: Schoettella novajanieae(sic) Palacios-Vargas, JG et Castaño-Meneses, G, 1998:779 ISS

Species tristani (Denis, J-R, 1931:69) Bonet, 1931

            Syn.: Beckerella Tristani(sic) Denis, J-R, 1931:69 IOS
            Syn.: Schoettella sp. Palacios-Vargas, JG, 1993:162
            Syn.: Schoettella sp. Zeppelini Filho, D et Castaño Meneses, G, 1995:10


             Syn.: Schöttella(sic) Tristani(sic) Denis, J-R, 1933:318 ISS
             Syn.: Schoettella Tristani(sic) Fernandes, LH et de Mendonça, MC, 1998:453 ISS



Species ununguiculata (Tullberg, 1869) Schäffer, 1896, t.t., B

             Syn.: Podura Tullbergii(sic) Lubbock, 1873 ISS
             Syn.: Podura tullbergii(sic) Foster, 1908 ISS


             Syn.: Achorutes uniunguicalatus(sic) Meinert, 1896 ISS
             Syn.: Schöttella(sic) uniunguiculata(sic) Schäffer, 1896 ISS
             Syn.: Schöttella(sic) uniunguiculata(sic) Schäffer, 1900 ISS
             Syn.: Schöttella(sic) uniunguiculata(sic) Krausbauer, 1901 ISS
             Syn.: Achorutes (Schöttella) uniunguiculatus(sic) Axelson, 1905 ISS
             Syn.: Achorutes uniunguicalatus(sic) Linnaniemi, 1907 ISS
             Syn.: Achorutes (Schöttella(sic)) uniunguiculata(sic) Folsom, JW, 1916:494 ISS
             Syn.: Schöttella(sic) uniunguiculata(sic) Bonet, F, 1930:121 ISS
             Syn.: Hypogastrura (Schöttella(sic)) uniunguiculata(sic) Bonet, F, 1930:132 ISS
Schoettella ununguiculata from the USA
Det. Christiansen, K.
2006 © Daun, A.
Schoettella ununguiculata from Germany
Eyepatch with 8 ocelli
2014.08.16 © Beck, K.
Schoettella ununguiculata from Germany
Eyepatch with 8 ocelli; cross-shaped PAO
2014.08.16 © Kluck, A.
Schoettella ununguiculata from Germany
Two anal spines
2014.08.16 © Kluck, A.
Schoettella ununguiculata from Germany
2014.08.22 © Kluck, A.
Schoettella ununguiculata from France
2018.01.28 © Erhardt, M.
Schoettella ununguiculata from France
2018.01.28 © Erhardt, M.

Subgenus Knowltonella Wray, 1958


Species idahoensis (Wray, 1958) Fjellberg, 1990, t.t.


Genus Austrogastrura Thibaud, J-M et Palacios-Vargas, JG, 1999:28

Species lobata (Yosii, R, 1959:3) Thibaud, J-M, Schulz, H-J & da Gama Assalino, MM, 2004:69

Species marambaia Fernandes, Bellini & de Mendonça, 2010

Species travassosi (Arlé, 1939) Rapoport, 1962, t.t.

            Syn.: Choreutinula edaphica Rapoport, 1962


             Syn.: Schoetella travassossi(sic) Fernandes, LH et de Mendonça, MC, 1998:453 ISS
             Syn.: Austrogastrura travassossi(sic) da Silveira, TC et de Mendonça, MC, 2020:211 ISS


Genus Choreutinula Paclt, 1944 key

F
  Syn.: Hypogastrurinae Beckerella Linnaniemi, WM, 1912:39
  Syn.: ... Schoettella Denis, 1931
  Syn.: Hypogastrura Mesogastrura Bonet, 1931, partim
  Syn.: Achorutinae Beckerellodes Salmon, JT, 1945:68


Species alpina Babenko AB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:199

Species americana Palacios-Vargas, JG, Castaño-Meneses, G et Gao, Y, 2010:7

Choreutinula americana from Canada
Unguiculus absent (*); inner ungual tooth present (**)
2016.01.12 © Rousseau, L.
Choreutinula americana from Canada
Two clavate tenent setae(*)
2016.01.12 © Rousseau, L.

Species gauquelini Gers & Deharveng, 1985

Species inermis (Tullberg, 1871) Stach, 1955, t.t., B

            Syn.: Achorutes (Schöttella(sic)) sensibilis Schött, 1902
            Syn.: Beckerella yanoi Kinoshita, 1932
            Syn.: Beckerella yanoi Uchida, 1938
            Syn.: Xenylla yanoi Kinoshita & Uchida, 1950
            Syn.: Beckerella yanoi Uchida, 1956
            Syn.: Beckerella yanoi Yosii, 1961
            Syn.: Choreutinula yanoi Salmon, JT, 1964:200
            Syn.: Beckerella yanoi Yosii, R, 1977:144
            Syn.: Choreutinula yanoi Thibaud, J-M, Schulz, H-J et da Gama Assalino, MM, 2004:69,sd
            Syn.: Choreutinula montana Martynova, 1968


             Syn.: Hypogastrura (Schoettella) intermis(sic) Loksa, I, 1978:53 ISS
Choreutinula inermis from France
2006 © Boldsystems.org
Choreutinula inermis from Japan
2013.03.11 © Hara, Y.
Choreutinula inermis from Japan
Ant.3 organ (A3O) with 2 microsensilla, 2 guard sensilla + laterodorsal microsensillum
2013.03.11 © Hara, Y.
Choreutinula inermis from Japan
Postantennal organ (PAO) roundish
2013.03.11 © Hara, Y.
Choreutinula inermis from Japan
Tibiotarsus with 2 knobbed tenent hairs (*)
2013.03.11 © Hara, Y.
Choreutinula inermis from Japan
Postantennal organ (PAO) roundish with 4 lobes; 8 ocelli
2013.03.11 © Hara, Y.



Species kulla Fjellberg, A, 2007:331

Species nodiseta (Handschin, 1928) Palacios-Vargas, JG, 1979:63

Species yinae Palacios-Vargas, JG, Castaño-Meneses, G et Gao, Y, 2010:9


Genus Mesogastrura Bonet, F, 1930:132

F
  Syn.: ... Beckerella Caroli, 1914
  Syn.: ... Schöttella(sic) Denis, 1931 partim
  Syn.: ... Mesachorutes Stach, 1919 partim
  Syn.: Hypogastrurinae Mesachorutes Bonet, F, 1930:132 partim
  Syn.: ... Mesachorutes Delamare Deboutteville, 1947
  Syn.: Neogastruridae Mesachorutes Stach, J, 1949:188


Species boneti (Tarsia in Curia, 1941) Delamare Deboutteville, 1950

             Syn.: Mesogastrura Boneti(sic) Delamare Deboutteville, 1950 ISS

Species coeca Cassagnau, P, 1959:70

             Syn.: Mesogastrura caeca(sic) Cassagnau, P, 1974:308 ISS

Species libyca (Caroli, 1914) da Gama, 1963

             Syn.: Mesachorutes libyca(sic) Goto, 1951 ISS
             Syn.: Schoettella lybica(sic) Rapoport, EH et Rubio, IR, 1963:97 ISS
             Syn.: Mesogastrura lybica(sic) Cassagnau, P, 1973:129 ISS
             Syn.: Mesogastrura lybica(sic) Cassagnau, P, 1974:308 ISS
Mesogastrura libyca from Holland
2018.03.19 © Kamsteeg, G.
Mesogastrura libyca from Holland
Eyepatch with 5 clear ocelli
2018.03.19 © Kamsteeg, G.
Mesogastrura libyca from the UK
After Shaw P & Trewhella S, 2019 Fig.13



Species ojcoviensis (Stach, 1919) Nosek, J, 1962:15

            Syn.: Beckerella quadriocellata Ionesco, 1922, nec Absolon, 1900
             Syn.: Beckerella 4-ocellata(sic) Bonet, F, 1930:137 ISS
            Syn.: Xenylla spelaea Ionesco, 1922
            Syn.: Hypogastrura (Mesogastrura) levantina Bonet, 1930
            Syn.: Hypogastrura (Mesogastrura) carpetana Bonet, F, 1930:133
            Syn.: Hypogastrura (Mesogastrura) intermedia Denis, 1931
             Syn.: Mesachorutes intermedia(sic) Delamare Deboutteville & Gisin, 1951 ISS
            Syn.: Hypogastrura (Mesogastrura) quadripunctata Gisin, 1944
            Syn.: Mesachorutes (Mesogastrura) quadripunctatus Gisin, H, 1960:58,
            Syn.: Mesachorutes ionescoi Delamare Deboutteville, 1947
            Syn.: Mesogastrura quadripunctata ssp. tirolensis Törne, 1955
            Syn.: Mesogastrura anthrohungarica Loksa, 1959
             Syn.: Mesogastrura antrohungarica(sic) Nosek, J, 1962:15 ISS
            Syn.: Mesachorutes tiliophilus Loksa, I et Rubio, I, 1966:140
Mesachorutes tiliophilus from Hungary
After Loksa I & Rubio I, 1966 Abb.11
Remark by Thibaud, J.-M., Schulz, H.-J. & da Gama Assalino, M.M. 2004:128: Mesachorutes tiliophilus Loksa & Rubio, 1966 is a species dubia. The typus has disappeared and the written description does not correspond with the drawing. This taxon belongs without doubt to Mesogastrura ojcoviensis.


             Syn.: Hypogastrura inermis(sic) Dallai, R et Malatesta, E, [1983]:174 ISS
            Syn.: Mesachorutes levantinus Jordana & Beruete, 1983



             Syn.: Mesogastrura ojcowiensis(sic) Bonet, F, 1930:121 ISS
             Syn.: Mesogastrura ojcowiensis(sic) Cassagnau, 1952 ISS
             Syn.: Mesogastrura ojcowiensis(sic) Salmon, JT, 1964:203 ISS
             Syn.: Mesogastrura ojcovensis(sic) Fjellberg, A, 1984:91 ISS
Mesogastrura ojcoviensis from Spain
After Arbea JI, 2013 Fig.4



Species sp. Leruth, R, 1939:197, B


Genus Denigastrura Stach, 1949

F
Species tetrophthalma (Denis, 1928) Stach, J, 1949:46, t.t.


Genus Gnathogastrura Díaz & Najt, 1983

F
Species paramoensis Díaz & Najt, 1983, t.t.

            Syn.: Gnathogastrura paramoense Díaz & Najt, 1983,tt IOS


Genus Barbagastrura Massoud, Najt & Thibaud, 1975

F
Species palpigera Massoud, Najt & Thibaud, 1975, t.t.


Genus Xenyllogastrura Denis, J-R, 1932:358 key

F
Species affinis (Steiner, 1955) Cassagnau, 1959

Species afurcata Deharveng & Gers, 1979

Species arenaria Fjellberg, A, 1992:449

Xenyllogastrura arenaria from the Canary Islands
After Fjellberg A, 1992 Fig.5B

Species octoculata (Steiner, 1955) Cassagnau, 1959

Species pruvoti Denis, J-R, 1932:358, t.t.

            Syn.: Xenyllogastrura Pruvoti(sic) Denis, J-R, 1932:358,tt IOS
Xenyllogastrura Pruvoti(sic) from France
After Denis J-R, 1932 Fig.I.1

  Subspecies minutavesicula Thibaud & Peja, 1996



Species reducta Fjellberg, A, 1992:449

Species steineri Jordana, R et Arbea, JI, 1992:13

Species venezueliensis Thibaud, J-M et Diaz, A, 1998:124


Genus Willemgastrura Pereira de Oliveira & Thibaud, 1988

F
Species coeca Oliveira & Thibaud, 1988, t.t.


Genus Tafallia Bonet, [1947]

F
Remark by Ellis, W.N. & Bellinger, P.F. 1973:54: The date of publication is given on the authority of the Zoological Record 86, Ins.: 398, errata, where entry 330 of volume 84 is corrected so that "1946" is to be read as "[1947]".

  Syn.: Onychiuridae Hoffia Scott, 1961


  Syn.: Hypogastrurinae Taffalia(sic) Poole in Poole & Gentili, 1997 ISS

Species insularis Bonet, [1947] tt

             Syn.: Tafallia insulares(sic) Gómez-Anaya, JA, Palacios-Vargas, JG et Castaño-Meneses, G, 2010:103 ISS

Species robusta (Scott, 1961) Christiansen, K & Bellinger, P, 1980:246

            Syn.: Tafallia sp. Bellinger, PF, 1985:79



Genus Parawillemia Izarra, 1975

F
Species pampeana Izarra, 1975, t.t.


Genus Biscoia Salmon, 1962

F
Species sudpolaris Salmon, 1962, t.t.

Biscoia sudpolaris from Antarctica
2008 © Univ of Waikato.


Genus Microgastrura Stach, 1922

F
  Syn.: Hypogastruridae Microgastrurella Massoud & Bellinger, 1963


Microgastrura sp. from Belgium
2021.10.08 © Huskens, M.-L.
Microgastrura sp. from Belgium
2021.10.08 © Huskens, M.-L.
Microgastrura sp. from Belgium
Eye with 6 ocelli
2021.10.08 © Huskens, M.-L.

Species sp.1 Janssens F sensu Ospina-Sánchez, 2019

            Syn.: Microgastrura sp.nov Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:5

Species duodecimoculata Stach, 1922, t.t.

Species jamaicensis (Massoud & Bellinger, 1963) Mari Mutt & Bellinger, 1990:19

Species massoudi Deharveng, L et Najt, J in Tillier, S, 1988:17

Species minutissima (Mills, HB, 1934:14) Stach, 1949:256

            Syn.: Microgastrura selgae Simon Benito & Pozo Martínez, 1984


             Syn.: Microgastrura minutissma(sic) Christiansen, KA in Dindal, DL, 1990:975 ISS
             Syn.: Microgastrura minutisima(sic) Vázquez & Palacios-Vargas, [1997] ISS
Microgastrura minutissima from Japan
After Nakamori, T. et al., 2009 Fig.1



Species nanacatlica Vázquez, M et Palacios-Vargas, JG, 1997:60

Species sensiliata Jordana, R, 1981:27

Species sofiae Vázquez, M et Palacios-Vargas, JG, 1997:62


Genus Acherongia Massoud & Thibaud, 1985 key

F
Species huetheri Fjellberg, A, 1992:455

Acherongia huetheri from the Canary Islands
After Fjellberg A, 1992 Fig.8A

Species minima Massoud & Thibaud, 1985, t.t.

Species palatinensis (Hüther, 1969) Massoud & Thibaud, 1985

Species steineri Christian, E et Thibaud, J-M, 1996:95


Genus Acherontides Bonet, 1945 key

M
Remark by Ellis, W.N. & Bellinger, P.F. 1973:1: Gender treated as neuter by the author [Bonet, 1945].
Species atoyacensis Bonet, 1945 tt

            Syn.: Acherontides atoyacense(sic) Bonet, 1945 tt IOS
             Syn.: Acherontides atoyacense(sic) Delamare Deboutteville, 1948 ISS
             Syn.: Acherontides atoyacense(sic) Stach, 1959 ISS
             Syn.: Acherontides atoyacense(sic) Salmon, JT, 1964:189 ISS
             Syn.: Acherontides atoyacense(sic) Palacios-Vargas, JG, 2006:9 ISS



Species bullocki Palacios-Vargas, JG et Gómez-Anaya, JA, 1996:377

             Syn.: Acherontides bullock(sic) de Lima, ECA, Stievano, LC et Zeppelini, D, 2019:143 ISS
             Syn.: Acherontides bullock(sic) de Lima, ECA, Stievano, LC et Zeppelini, D, 2019:148 ISS

Species edaphicus Yosii, 1971

Species eleonorae Palacios-Vargas, JG et Gnaspini-Netto, P, 1992:443

             Syn.: Acherontides leo(sic) Palacios-Vargas, JG et Zeppelini, D, 1995:173 lapsus ISS
Remark by Deharveng, L. 2013.12.22: Palacios-Vargas & Zeppelini, 1995 refer in error to Acherontides leo Palacios-Vargas & Gnaspini-Netto, 1992 for the species A. eleonorae Palacios-Vargas & Gnaspini-Netto, 1992 from Brazil.


Species huetheri Deharveng, L et Diaz, A, 1984:340

Species juxtlahuacaensis Palacios-Vargas, JG et Gómez-Anaya, JA, 1996:375

Species pappogeomysae Palacios-Vargas, JG et Gómez-Anaya, JA, 1996:379

Species peruensis Hüther, 1975

Species potosinus Bonet, 1946

Species serrasapoensis de Lima, ECA, Stievano, LC et Zeppelini, D, 2019:142

Acherontides serrasapoensis from Brazil
After de Lima ECA, Stievano LC & Zeppelini D, 2019 Fig.1
Acherontides serrasapoensis from Brazil
Dorsal chaetotaxy
Modified after de Lima ECA, Stievano LC & Zeppelini D, 2019 Fig.3+4

Species spinus (Christiansen, K & Reddell, JR, 1986:155) Mari Mutt, JA & Bellinger, PF, 1990:17

Species tanasachiae (Gruia, 1969) Gruia, 1970

             Syn.: Acherontides tanasachie(sic) de Lima, ECA, Stievano, LC et Zeppelini, D, 2019:143 ISS


Genus Acherontiella Absolon, 1913

F
  Syn.: ... Acherontiellina Delamare Deboutteville, 1948
  Syn.: ... Xenyllina Delamare Deboutteville, 1948 nn


Species aokii Tamura & Yue, 1999

Species bougisi Cassagnau, P et Delamare Deboutteville, C, 1955:370

            Syn.: Acherontiella Bougisi(sic) Cassagnau, P et Delamare Deboutteville, C, 1955:370 IOS
            Syn.: Acherontiella bougisi ssp. rhodia Ellis, WN, 1974:109


             Syn.: Acherontiellina bouigisi(sic) Fjellberg, A, 1992:452 ISS
Acherontiellina bougisi from the Canary Islands
After Fjellberg A, 1992 Fig.6A





Species candida (Delamare Deboutteville, 1952) Deharveng & Diaz, 1984

Species carusoi Dallai, R, 1978:345

Species cassagnaui Thibaud, 1967

Species cavernicola (Tarsia in Curia, 1941) Ellis, 1974

Species colotlipana Palacios-Vargas, JG et Thibaud, J-M, 1985:9

Species dentata Djanaschvili, 1971

            Syn.: Acherontiella cryptobia Djanaschvili, 1971


Species epigea Bonet, 1945

Species globulata Thibaud & Massoud, 1980

Species kowalskiorum Weiner, WM et Najt, J, 1998:217

Species mac (Palacios-Vargas, JG & Thibaud, J-M, 1985:1986:11) Thibaud, J-M, 1990:407

Species massoudi Thibaud, 1963

Species onychiuriformis Absolon, 1913, t.t.

Species prominentia Thibaud, J-M et Weiner, WM in Najt, J et Matile, L, 1997:64

Acherontiella prominentia from New Caledonia
After Thibaud J-M & Weiner WM, 1997 Fig.3

Species sabina Bonet, 1945:31

Species thai Thibaud, J-M, 1990:408

Species thibaudi Barra, J-A, 1994:181

Species variabilis Delamare Deboutteville, 1948

Species xenylliformis Gisin, 1952


Genus Acheroxenylla Ellis, WN, 1976:236 key

F
Species canariensis Fjellberg, A, 1992:451

Acheroxenylla canariensis from the Canary Islands
After Fjellberg A, 1992 Fig.6D

Species cretensis Ellis, WN, 1976:238, t.t.

Acheroxenylla cretensis from Crete
After Ellis WN, 1976 Fig.6

Species furcata Fjellberg, A, 1992:454

Acheroxenylla furcata from the Canary Islands
After Fjellberg A, 1992 Fig.7A

Species lipsae Palacios-Vargas, JG, 2020:112

Acheroxenylla lipsae from Peru
© Montejo M in Palacios-Vargas JG, 2020 Fig.3
Acheroxenylla lipsae from Peru
© Arango M in Palacios-Vargas JG, 2020 Fig.4
Acheroxenylla lipsae from Peru
Modified after Palacios-Vargas JG, 2020 Fig.6
Designation of ocellus E incorrect
Ocellus E should be designated as F (red)


Genus Paraxenylla Murphy, DH, 1965:392 key  key

F
  Syn.: Hypogastruridae Haloxenylla da Gama, MM et Deharveng, L, 1984:131


  Syn.: Hypogastruridae Paraxenyla(sic) Skarzynski, D, 2007:201 ISS

Species affiniformis (Stach, J, 1930:280) Jordana & Arbea, 1989, t.t.

            Syn.: Xenylla (Paraxenylla) mangle Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:306 =?


             Syn.: Haloxenylla affiniforrnis(sic) da Gama, MM, 1988:75 ISS
Paraxenylla affiniformis from the Canary Islands
After Fjellberg A, 1992 Fig.3U



Species arenosa (Uchida, H & Tamura, H, 1967:234) Palacios-Vargas, JG & Janssens, F, 2006:42

Xenylla arenosa from Japan
After da Gama MM, 1981 Fig.2
Dorsal chaetotaxy of head, th.I-II and abd. I, IV-VI

Species cubana Palacios-Vargas, JG et Janssens, F, 2006:39

Species lapazana Palacios-Vargas, JG et Vázquez, M, 1989:436

            Syn.: Paraxenylla sp. Palacios-Vargas, 1987


Species mahahualana Palacios-Vargas, JG et Vázquez, M, 2018:1057

Paraxenylla mahahualana from Mexico
After Palacios-Vargas JG & Vázquez M, 2018 Fig.7+10

Species mangle (Murphy, DH, 1965:392) Palacios-Vargas, JG & Janssens, F, 2006:37, t.t.

Species norvegica Fjellberg, A, 2010:65

Paraxenylla norvegica from the UK
After Fjellberg A, 2010 Fig.1

Species oceanica (Yosii, R, 1960:10) Palacios-Vargas, JG & Janssens, F, 2006:42

Species peruensis Palacios-Vargas, JG et Janssens, F, 2006:41

Species piloua Thibaud, J-M et Weiner, WM in Najt, J et Matile, L, 1997:66

Paraxenylla piloua from New Caledonia
After Thibaud J-M & Weiner WM, 1997 Fig.13
Paraxenylla piloua from New Caledonia
After Thibaud J-M & D'Haese C, 2010 Fig.27

Species sooretamensis Queiroz, GC et Deharveng, L, 2008:?


Genus Pseudacherontides Djanaschvili, 1971

M
  Syn.: Hypogastruridae PseudoAcherontides(sic) de Lima, ECA, Stievano, LC et Zeppelini, D, 2019:141 ISS
  Syn.: Hypogastruridae PseudAcherontides(sic) de Lima, ECA, Stievano, LC et Zeppelini, D, 2019:141 ISS

Species aspinatus (Stach, 1959)

Species bisetosus (Stach, 1959)

Species bulgaricus (Martynova in Martynova & al., 1971)

Species crassus (Stach, 1959)

Species spelaeus (Ionesco, 1922) Arbea, JI & Jordana, R, 1991:21

             Syn.: Acherontides spelaea(sic) Delamare Deboutteville, 1948 ISS
             Syn.: Beckerella spalaea(sic) Stach, J, 1949:167 ISS
             Syn.: Acherontides spelaea(sic) Salmon, JT, 1964:189 ISS
             Syn.: Mesachorutes spelæus(sic) Dethier, M, 1998:55 ISS

Species stachi (Ljovushkin, 1972)

Species vivax (Yosii, 1956)

             Syn.: Acherontides vivax Salmon, JT, 1964:189 ISS

Species zenkevitchi Djanaschvili, 1971, t.t.

            Syn.: Pseudacherontides birsteri Djanaschvili, 1971
            Syn.: Pseudacherontides trisetosus Djanaschvili, 1971



Genus Thibaudylla Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:21

Species anniae (Najt, J & Weiner, WM, 1991:98) Najt, J & Weiner, WM in Najt, J & Matile, L, 1997:23

Species danieleae (Deharveng, L & Najt, J in Tillier, S, 1988:17) Najt, J & Weiner, WM in Najt, J & Matile, L, 1997:23, t.t.

Species palpata (Deharveng, L & Najt, J in Tillier, S, 1988:25) Najt, J & Weiner, WM in Najt, J & Matile, L, 1997:23

Species thibaudi (Massoud, 1965) Najt, J & Weiner, WM in Najt, J & Matile, L, 1997:24


Genus Xenylla Tullberg, 1869 key  key  key  key

F
  Syn.: Achorutidae Achorutini(sic) Börner, C, 1901:13 partim IOS
  Syn.: ... Propexenylla Salmon, 1944
  Syn.: ... Proxenylla(sic) Pomorski & Skarzynski, 1991 ISS
  Syn.: Collembola Entomobryomorpha Meyer-Rochow, VB, 2013:2
Entomobryomorpha(lapsus) from Nord Korea
Hexagonal epicuticular ultrastructure
After Meyer-Rochow VB, 2013 Fig.2
Entomobryomorpha(lapsus) from Nord Korea
Retinaculum
After Meyer-Rochow VB, 2013 Fig.5
Entomobryomorpha(lapsus) from Nord Korea
Setae on labium
After Meyer-Rochow VB, 2013 Fig.6
Entomobryomorpha(lapsus) from Nord Korea
Eyepatch with 5 ocelli
Negative image
Modified after Meyer-Rochow VB, 2013 Fig.7




  Syn.: Achorutini(sic) Xennylla(sic) Börner, C, 1901:13 ISS
  Syn.: ... Xennylla(sic) Börner, 1906 ISS
  Syn.: ... Xynella(sic) Prinzing, A, 1996:20 sp ISS
Xenylla sp. from Hungary
2008.08.10 © Keresztes, G.
Xenylla sp. from the UK
2012.08.17 © Murray, A.
Xenylla sp. juv. from the UK
2012.11.11 © Nurcombe, E.
Xenylla sp. juv. from the UK
Eyepatch with 5 ocelli
2012.11.11 © Nurcombe, E.
Xenylla sp. juv. from the UK
Body dorso-ventrally flattened
2012.11.11 © Nurcombe, E.
Xenylla sp. from New Zealand
2013.10.12 © Sullivan, J.
Xenylla sp. juv. from Holland
2014.01.24 © van Duinen, J.
Xenylla sp. juv. from Germany
2014.11.04 © Beck, K.
Xenylla sp. from the UK
2015.02.14 © Barton, T.


  Syn.: Hypogastruridae Xenyla(sic) Queiroz, GC et Mendonça, MC, 2016:36 ISS
Xenylla sp. from Belgium
Eyepatch with 5 ocelli
2016.02.16 © Huskens, M.-L.
Xenylla sp. from Belgium
2016.12.28 © Huskens, M.-L.
Xenylla sp. from the USA
Abd.6 with minute anal spines (*)
2017.04.04 © Fritzl.
Xenylla sp. from the USA
Furca with mucro fused to dens
2017.04.04 © Fritzl.
Xenylla sp. from France
Unguiculus absent (*)
2017.11.04 © Picard, J.
Xenylla sp. from Australia
Eyepatch with 5 ocelli
2017.12.29 © Vale, E.
Xenylla sp. from Australia
2017.12.29 © Vale, E.
Xenylla sp. from Holland
Unguiculus absent (*)
2019.02.07 © Kamsteeg, G.
Xenylla sp. from Spain
Cuticula with leather look
2020.10.25 © Kamsteeg, G.
Xenylla sp. from the UK
Skin appears lile leather
2021.02.01 © Barton, T.
Xenylla sp. from the UK
Eyepatch with 5 ocelli
2021.02.01 © Barton, T.





Species sp.1 Janssens F sensu Bellini & Zeppelini, 2004

            Syn.: Xenylla sp.undescr. Bellini, BC et Zeppelini, D, 2004:588

Species sp.2 Janssens F sensu Cutz-Pool & al, 2008

            Syn.: Xenylla sp.nov Cutz-Pool, LQ, Palacios-Vargas, JG et Castaño-Meneses, G, 2008:4

Species sp.3 Janssens F sensu Palacios-Vargas, 2008

            Syn.: Xenylla sp.nov. Palacios-Vargas, JG, 2008:46

Species sp.4 Janssens F sensu Zeppelini & al, 2009

            Syn.: Xenylla sp.n. Zeppelini, D, Bellini, BC, Creao-Duarte, AJ et Hernández, MIM, 2009:1166

Species sp.5 Janssens F sensu Bernard, 2015

            Syn.: Xenylla duchesnea Christiansen, K et Bellinger, P, 1980:220
            Syn.: Xenylla duchesnea Christiansen & Bellinger, 1998
            Syn.: Xenylla sp.undescr. Bernard, EC, 2015:330

Species sp.6 Janssens F sensu Ospina-Sánchez, 2019

            Syn.: Xenylla sp.nov.1 Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:6

Species sp.7 Janssens F sensu Ospina-Sánchez, 2019

            Syn.: Xenylla sp.nov.2 Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:6

Species abichiana

Species acauda Gisin, 1947:344

Species aculeata Babenko, A, Efeykin, B et Bizin, M, 2020:10

Xenylla aculeata from NE Russia
Modified after Babenko A, Efeykin B & Bizin M, 2020 Fig.30-31

Species alba Folsom, JW, 1932:54

Species andrzeji Busmachiu & Weiner, 2008

Species aristides Fernando, 1959

Species arnei Babenko, A, Efeykin, B et Bizin, M, 2020:13

Xenylla arnei from NE Russia
After Babenko A, Efeykin B & Bizin M, 2020 Fig.36-37

Species atrata (Salmon, JT, 1944:125) da Gama, MM, 1969:15

            Syn.: Xenylla longiseta Cassagnau & Rapoport, 1962


Species auka Christiansen, K et Bellinger, P, 1992:19

Species australiensis da Gama, 1974

            Syn.: Xenylla mucronata Womersley, 1939
             Syn.: Achorutes speciosa(sic) Womersley, H, 1939:92 ISS


             Syn.: Xenylla australis(sic) Greenslade P, 1981 ISS
  Subspecies westraliensis da Gama, 1974



Species babenkoi Stebaeva SK et Potapov MB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:300

Species badakhshanica Yosii, 1966

             Syn.: Xenylla badakhsanica(sic) Jia, J-L et Skarzynski D, 2019:581 ISS

Species bellingeri da Gama, 1969:7

             Syn.: Xenylla Bellingeri(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:109,cf ISS

Species betulae Fjellberg, 1985

Species boerneri Axelson, WM, 1905:789

            Syn.: Xenylla börneri(sic) Axelson, WM, 1905:789 IOS
            Syn.: Xenylla pseudoboerneri Stach, J, 1949:241



            Syn.: Xenylla börneri(sic) Axelson, 1905 IOS
            Syn.: Xenylla börneri(sic) Axelson, 1907 IOS
            Syn.: Xenylla börneri(sic) Linnaniemi, 1912 IOS
            Syn.: Xenylla börneri(sic) Handschin, E, 1924:94 IOS
            Syn.: Xenylla börneri(sic) Brown, 1926 IOS
            Syn.: Xenylla börneri(sic) Brown, 1929 IOS
            Syn.: Xenylla börneri(sic) Handschin, E in Dahl, F, 1929:30 IOS
            Syn.: Xenylla börneri(sic) Womersley, 1930 IOS
            Syn.: Xenylla börneri(sic) Durkop, 1932 IOS
            Syn.: Xenylla börneri(sic) Tarsia in Curia, 1936 IOS
            Syn.: Xenylla börneri(sic) Frenzel, 1937 IOS
            Syn.: Xenylla börneri(sic) Kseneman, 1938 IOS
            Syn.: Xenylla börneri(sic) Agrell, 1939 IOS
            Syn.: Xenylla börneri(sic) Agrell, 1941 IOS
            Syn.: Xenylla börneri(sic) Agrell, 1943 IOS
            Syn.: Xenylla börneri(sic) Gisin, 1943 IOS
            Syn.: Xenylla börneri(sic) Gisin, 1944 IOS
            Syn.: Xenylla börneri(sic) Kloet, GS et Hincks, WD, 1945:3 IOS
            Syn.: Xenylla börneri(sic) Weis-Fogh, 1948 IOS
            Syn.: Xenylla börneri(sic) Strenzke, 1949 IOS
            Syn.: Xenylla börneri(sic) Goto, 1955 IOS
            Syn.: Xenylla börneri(sic) Stach, 1955 IOS
            Syn.: Xenylla börneri(sic) Gisin, 1957 IOS
            Syn.: Xenylla börneri(sic) Pschorn-Walcher & Gunhold, 1957 IOS
            Syn.: Xenylla börneri(sic) Grinbergs, 1958 IOS
            Syn.: Xenylla börneri(sic) von Törne, 1958 IOS
            Syn.: Xenylla börneri(sic) Dallai, R, Malatesta, E et Ramellini, P in Minelli, A, Ruffo, S et La Posta, S, 1995:4 IOS

Species brasiliensis da Gama, 1978

Species brevicauda Tullberg, 1869, B

            Syn.: Xenylla microchaeta Cassagnau, 1968


Species brevispina Kinoshita, 1916

            Syn.: Xenylla humicola Uchida, 1954
            Syn.: Xenylla humicola Yosii, 1954
            Syn.: Xenylla humicola Salmon, JT, 1964:194 partim
            Syn.: Xenylla humicola Yosii, R, 1977:144


Xenylla brevispina from Japan
2019.02.27 © Hara, Y.
Xenylla brevispina from Japan
Eyepatch with 5 ocelli
2019.02.27 © Hara, Y.

Species californica da Gama, MM, 1976:325

Species canadensis Hammer, 1953

Species capensis Weiner & Najt, 1991

Species capitata Thibaud & Massoud, 1980

Species capixaba Fernandes, LH et de Mendonça, MC, 2010:124

Species cassagnaui da Gama, 1983

Species caudata Jordana, 1993

Species cavarai Caroli, 1914

Species cavernarum Jackson, 1927

  Subspecies salomonensis da Gama, MM, 1967:6

Species changchunensis Wu, DH et Yin, WY, 2007:380

             Syn.: Xenylla Changchunensis(sic) Wu, DH et Yin, WY, 2007:383 ISS

Species changlingensis Wu, DH et Yin, WY, 2007:378

             Syn.: Xenylla Changlingensis(sic) Wu, DH et Yin, WY, 2007:383 ISS

Species christianseni da Gama, MM, 1974:323

             Syn.: Xenylla chistianseni(sic) Palacios-Vargas, JG et Mejía-Recamier, BE, 2008:98 ISS

Species claggi Wise, KAJ, 1970:185

Species collis Bacon, 1914, s.i.

Species constricta von Olfers, 1907

Species continentalis Stebaeva SK et Potapov MB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:282

Species convexopyga Lee, B-H, Park, N-Y et Park, K-H, 2005:266

Species corticalis Börner, C, 1901:696

            Syn.: Xenylla pallida Ågren, 1903 cf
            Syn.: Xenylla planipila Stach, J, 1949:235


Species deharvengi da Gama, 1983

Species dotata Lee, B-H, Park, N-Y et Park, K-H, 2005:268

Species duchesnea Wray, 1958

             Syn.: Xenylla duchensnea(sic) Poole, RW in Poole, RW et Gentili, P, 1997:49 ISS

Species fernandesi da Gama, MM, 1974:331

Species franzi Steiner, 1955

Species gamae Cardoso, MA, 1967:187

Species gisini Cardoso, MA, 1968:1

Species gomerensis Fjellberg, A, 1992:443

Species granulosa da Gama, 1966

Species greensladeae da Gama, 1974

            Syn.: Xenylla prisca Womersley


Species grisea Axelson, 1900, B

            Syn.: Xenylla gracilis Guthrie, JE, 1903:95
            Syn.: Xenylla grisea var. decolor Axelson, 1904
            Syn.: Xenylla cavarai Caroli, 1914
            Syn.: Xenylla cavernarum Jackson, 1927
            Syn.: Xenylla grisea f. brevifurcata Denis, 1931
            Syn.: Xenylla grisea f. principalis Denis, 1931
            Syn.: Xenylla maritima Womersley, 1933
            Syn.: Xenylla maritima Paclt, 1959, nec Tullberg, 1869


             Syn.: Xeniella(sic) grisella(sic) Weinreich, 1968 ISS
             Syn.: Xeniella(sic) grisella(sic) Messr, C, Walther, J, Dettner, K et Schulz, S, 2000:217 ISS
Xenylla grisea from the UK
2012.05.dd © Murray, A.
Xenylla grisea from the UK
2012.05.dd © Murray, A.
Xenylla grisea from the UK
Eyepatch with 5 ocelli
2012.12.12 © Nurcombe, E.
Xenylla grisea juv. from the UK
2013.05.08 © Murray, A.
Xenylla grisea juv. from the UK
2013.05.08 © Murray, A.
Xenylla grisea juv. from the UK
Composed eye with 5 ocelli
2013.05.13 © Murray, A.
Xenylla grisea juv. from the UK
With indistinctly small anal spines
2013.05.13 © Murray, A.
Xenylla ?grisea? from the UK
2014.10.26 © Barton, T.
Xenylla grisea from Holland
Eyepatch with 5 ocelli
2014.11.24 © van Duinen, J.
Xenylla grisea from the UK
2016.10.03 © Barton, T.
Xenylla grisea from France
2017.09.02 © Picard, J.
Xenylla grisea from the UK
Eyepatch with 5 ocelli
2019.08.13 © Smart, J.
Xenylla grisea from Switzerland
2020.01.16 © Bornand, P.
Xenylla grisea from Switzerland
Eyepatch with 5 ocelli
Ant.3+4 fused
2020.01.16 © Bornand, P.
Xenylla grisea from Switzerland
2020.01.16 © Bornand, P.
Xenylla grisea from Switzerland
Eyepatch with 5 ocelli
Ant.3+4 fused
2020.01.16 © Bornand, P.
Xenylla grisea from France
2022.01.09 © Usami, R.



Species gwiazdowiczi Sternalski, J et Pasnik, G, 2019:186

Xenylla gwiazdowiczi from Peru
After Sternalski J & Pasnik G, 2019 Fig.1

Species hadialii Baijal, HN, 1955:532

Species hawaiiensis da Gama, MM, 1969:48

Species helena Scott, 1937

Species hexagona Fjellberg, A, 1992:443

Species hodori Neves, ACR et de Mendonça, MC, 2017:188

Xenylla hodori from Brazil
After Neves ACR & de Mendonça MC, 2017 Fig.1 top
Xenylla hodori from Brazil
After Neves ACR & de Mendonça MC, 2017 Fig.1 bottom
Xenylla hodori from Brazil
After Neves ACR & de Mendonça MC, 2017 Fig.2cef

Species humicola (Fabricius O, 1780) Reuter, 1876, B

            Syn.: Xenylla maritima Tullberg, 1871
            Syn.: Xenylla maritima Tullberg, 1872
            Syn.: Xenylla maritima Reuter, OM, 1895:32 partim
            Syn.: Achorutes maritima Meinert, 1896
            Syn.: Xenylla maritima Caspers, 1942
            Syn.: Achorutes viaticus Tullberg, 1872
            Syn.: Achorutes murorum Lubbock, 1873
            Syn.: Xenylla affinis Schäffer, 1897
            Syn.: Xenylla humicola var. olivacea Axelson, WM, 1905:789
            Syn.: Xenylla baconae Folsom, JW, 1916:496
            Syn.: Xenylla orientalis Handschin, 1932
            Syn.: Xenyllodes hoffi Scott, HG, 1960:185
Xenyllodes hoffi from New Mexico
After Scott HG, 1960 Fig.2

            Syn.: Xenylla neomexicana Scott, HG, 1960:60
Xenylla neomexicana from New Mexico
After Scott HG, 1960 Fig.2

            Syn.: Anurida violacea Scott, HG, 1960:186
Anurida violacea from New Mexico
After Scott HG, 1960 Fig.4

            Syn.: Schoettella santiaguensis Rapoport, EH et Rubio, IR, 1963:95
Schoettella santiaguensis from Chile
After Rapoport EH & Rubio IR, 1963 Fig.1




Xenylla humicola from the UK
Distinctly long pointed mucro; mucro apparently fused with dens
2020.03.09 © Deed, Be.

Species inermis Olfers, 1907

Species jamaicensis da Gama, MM, 1969:37

Species jocquei André, M, 1988:508

Species kenyensis da Gama, 1983

Species kirgisica Martynova, 1976

Species laurisilvae Fjellberg, A, 1992:445

Species lawrencei da Gama, MM, 1967:20

Species lesnei Denis, 1935

Species littoralis Womersley, 1933

Xenylla littoralis from Australia
After da Gama MM, 1980 Fig.1
Dorsal chaetotaxy of head, th.I-II and abd. I, IV-VI
Xenylla littoralis from Tasmania
2014.04.26 © Murray, A.
Det.: Greenslade, P.
Xenylla littoralis from Tasmania
2014.04.26 © Murray, A.
Det.: Greenslade, P.
Xenylla littoralis from Tasmania
2014.04.26 © Murray, A.
Det.: Greenslade, P.
Xenylla littoralis from Tasmania
Anal spines
2014.04.26 © Murray, A.
Det.: Greenslade, P.

Species longicauda Folsom, JW, 1898:53

Xenylla longicauda from Japan
After Folsom JW, 1898 Fig.10

             Syn.: Xenylla longicaudata(sic) Yosii, R, 1960:10 ISS


Species longispina Uzel, 1890

Species longistriata Lee, B-H, Park, N-Y et Park, K-H, 2005:268

Species lotharingiae Thibaud, 1963

            Syn.: Xenylla xavieri Simón & Selga, 1977 =?
            Syn.: Xenylla xavieri Acón & Simón, 1979 =?


Xenylla lotharingiae from Spain
After Jordana R & al, 2020 Fig.5D

Species louisiana da Gama, MM, 1976:323

Species malasica da Gama, 1969:38

Species malayana Salmon, 1951

Species manuelae Queiroz, GC et Mendonça, MC, 2016:37

Xenylla manuelae from Brazil
After Queiroz GC & de Mendonça MC, 2016 Fig.2B

Species manusiensis da Gama, MM, 1967:16

Species marina Lee, B-H, Park, N-Y et Park, K-H, 2005:271

Species martynovae Dunger, 1983

            Syn.: Xenylla erzinica Stebaeva, 1977


Species mongolica Martynova, 1975

Species mucronata Axelson, WM, 1903:5

            Syn.: Xenylla schillei Börner, 1903 =?
             Syn.: Xenylla nova-zelandia(sic) Salmon, 1941 =? ISS


Species murphyi da Gama, MM, 1969:41

Species myrmecophila Stebaeva SK et Potapov MB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:265

Species namia Park, 2016

Species neivai da Gama, 1966

Species nepalensis da Gama, 1988

Species nigeriana da Gama & Lasebikan, 1976

Species nirae da Gama & de Oliveira, 1994

Species obscura Imms, 1912

            Syn.: Xenylla terricola Stach, J, 1965:347


             Syn.: Xenylla obscurus(sic) Greenslade P in Barker & Greenslade PJM, 1982 ISS

Species occidentalis Womersley, 1933

Species octooculata Carpenter, 1928

            Syn.: Xenylla octo-oculata(sic) Carpenter, 1928 IOS


             Syn.: Xenylla octo-oculata(sic) Handschin, 1938 ISS
             Syn.: Xenylla octo-oculata(sic) Stach, J, 1949:202 ISS
             Syn.: Xenylla octo-oculata(sic) Salmon, JT, 1964:196 ISS



Species orientalis Handschin, 1932

Species osetica Stebaeva SK et Potapov MB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:276

Species pallescens (Scott, HG, 1960:186) da Gama, MM, 1974:328

Xenyllodes pallescens from New Mexico
After Scott HG, 1960 Fig.3

            Syn.: Hypogastrura albamaculata Scott, HG, 1960:55
Hypogastrura albamaculata from New Mexico
After Scott HG, 1960 Fig.1
Remark by Christiansen, K. 2007.06.11 (in litt.): Type specimens lost. Species dubia.

             Syn.: Hypogastrura albomaculata(sic) Poole, RW in Poole, RW et Gentili, P, 1997:47 ISS
            Syn.: Xenyllodes alpinus Scott, HG, 1960:184
Xenyllodes alpinus from New Mexico
After Scott HG, 1960 Fig.1.






Species paludis (Bacon, 1914, s.i.) Bacon, 1914

Species piceeta Stebaeva SK et Potapov MB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:295

Species portoricensis da Gama, MM, 1976:319

Species proxima Denis, 1931

Species pseudobrevicauda Ritter, 1911




             Syn.: Xenylla pseudobrevicaudata(sic) Yosii, 1959 ISS

Species pseudomaritima James, 1933

Species pyrenaica Cassagnau, P, 1959:73

Species raynalae Najt, Thibaud & Weiner, 1990

             Syn.: Xenylla raynale(sic) Queiroz, GC et Mendonça, MC, 2016:46 ISS
             Syn.: Thibaudylla? raynale(sic) Queiroz, GC et Mendonça, MC, 2016:46 ISS

Species reducta Prabhoo, 1971

Species rhodesiensis Womersley, 1926

Species saludoi Izarra, 1970

Species schillei Börner, 1903

             Syn.: Xenylla shillei(sic) Fiera, C, 2007:9 ISS

Species simberloffi da Gama, MM, 1974:334

Species similata Denis, 1948

            Syn.: Xenylla yucatana da Gama ?


Species sincta Baijal, 1956

Species spinosissima Najt & Rubio, 1978

Species stachi da Gama, 1966

  Subspecies bismarckensis da Gama, MM, 1969:39

            Syn.: Xenylla stachi ssp. stachi da Gama, 1967, nec da Gama, 1966


  Subspecies wolffi da Gama, MM, 1967:15

Species stepposa Stebaeva, 1980

Species subacauda Stebaeva SK et Potapov MB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:303

Species subbellingeri da Gama, MM, 1976:320

Species subcavernarum da Gama, MM, 1969:20

Species subcorta (Salmon, JT, 1941:295) Greenslade, P, Stevens, MI, Torricelli, G & D'Haese, C, 2011:234

Species tadzhika Martynova, 1968

Species taihangensis Jia, J-L, Wang, Z-J et Skarzynski, D, 2020:179

Species thailandensis da Gama, 1986

Species thiensis Deharveng, L et Najt, J in Tillier, S, 1988:17

Species trisubloba Stebaeva SK et Potapov MB in Babenko, AB, Chernova, NM, Potapov, MB et Stebaeva, SK, 1994:284

Species tullbergi Börner, 1903, B

            Syn.: Xenylla brevicauda Börner, 1901 sensu Börner, 1903
            Syn.: Xenylla Birói(sic) Stach, 1926 IOS
             Syn.: Xenylla Birói(sic) Stach, J, 1930:271 ISS


Xenylla tullbergi from Spain
After Jordana R, 1981 Lam.X Fig.51



Species uniseta da Gama, 1963

Species vanharteni Weiner, WM, Najt, J et Pasnik, G, 2012:555

Xenylla vanharteni from Yemen
After Weiner WM, Najt J & Pasnik G, 2012 Fig.1A

Species victoriana da Gama, MM, 1980:126

Species vilhenaorum da Gama, 1966

Species villiersi Thibaud, 1963

Species wandae Queiroz, GC et Mendonça, MC, 2016:42

Species weinerae Jia, J-L et Skarzynski D, 2019:580

Species welchi Folsom, JW, 1916:497, B

            Syn.: Xenylla Subwelchi(sic) Denis, J-R, 1924:216 IOS
            Syn.: Xenylla Subwelchi(sic) Denis, J-R, 1924:217 IOS
Xenylla Subwelchi(sic) from Monaco
After Denis J-R, 1924 Fig.17

             Syn.: Xenylla Subwelchi(sic) Denis, J-R, 1933:225 ISS
            Syn.: Xenylla sensilis Folsom, JW, 1932:54
            Syn.: Xenylla indus Yosii & Ashraf, 1964
            Syn.: Achorutes magnoliana Wray DL in Bernard, EC, 2015:323



             Syn.: Xenylla Welchi(sic) Denis, J-R, 1924:218 ISS
             Syn.: Xenylla Welchi(sic) Denis, J-R, 1933:225 ISS
             Syn.: Xenylla Welchi(sic) Marlier, G in Pasteels, M, 1948:59,B ISS



Species wilsoni da Gama, MM, 1974:326

Species womersleyi da Gama, 1974

Species xavieri da Gama, 1959

            Syn.: Xenylla lotharingiae da Gama, 1964
            Syn.: Xenylla lotharingiae Thibaud, J-M, Schulz, H-J et da Gama Assalino, MM, 2004:250
            Syn.: Xenylla brevicauda Jordana, 1981
            Syn.: Xenylla brevicauda Arbea & Jordana, 1985


Species yosiiana da Gama, 1971

Species yucatana Mills, HB in Pearse, AS, 1938:183

            Syn.: Xenylla mucronata Yosii, 1960 nec Axelson, 1903
            Syn.: Xenylla aelleni da Gama, 1964


             Syn.: Xenylla yukatana(sic) Babenko, A, Efeykin, B et Bizin, M, 2020:16 ISS

Species zairensis Martynova, 1979

Species zavattari (Tarsia in Curia, 1939) Stach, J, 1949:212, s.i.

             Syn.: Xenylla zovattari(sic) Stach, J, 1949:212,si ISS
             Syn.: Xenylla zavattarii(sic) Salmon, JT, 1964:198 ISS

Species-group maritima Skarzynski D, Piwnik A & Porco D, 2018

Species asiatica Martynova, 1975

Species brevisimilis Stach, J, 1949:232, B

             Syn.: Xenylla brevissimilis(sic) Özata, MA, Sevgili, H et Kaprus, IJ, 2017:239 ISS

Species maritima Tullberg, 1869, t.t., B

            Syn.: Podura maritima Gmelin, 1793             Syn.: Xenylla nitida Tullberg, 1871=?
            Syn.: Xenylla nitida Ågren, 1903 sensu Linnaniemi, 1912=?
            Syn.: Xenylla nitida Börner, 1903=?
            Syn.: Xenylla nitida Gisin, 1944=?
            Syn.: Xenylla nitida Stach, 1949=?
            Syn.: Xenylla humicola Moniez, 1889
            Syn.: Xenylla brevicauda Linnaniemi, 1912
            Syn.: Xenylla corticalis Börner, 1901, sensu Gisin, 1944
            Syn.: Xenylla pallida Ågren, 1903
            Syn.: Xenylla proxima Denis, 1931, sensu Gisin, 1944
            Syn.: Xenylla pseudomaritima James, 1933, sensu Gisin, 1944
            Syn.: Odontella minutadentata Salmon, JT, 1941:297
Odontella minutadentata from New Zealand
After Salmon, J.T., 1941 Pl.43 Fig.61.

             Syn.: Zealandella minutadenta(sic) Greenslade, P in Houston, WWK, 1994:40 ISS
            Syn.: Xenylla nova-zealandia(sic) Salmon, JT, 1941:287 IOS
             Syn.: Xenylla nova-zealandia(sic) Salmon, 1949 ISS
             Syn.: Xenylla nova-zealandia(sic) Stach, J, 1949:207 ISS
             Syn.: Xenylla nova-zealandia(sic) Salmon, JT, 1964:111 ISS
             Syn.: Xenylla nova-zealandia(sic) Salmon, JT, 1964:196 ISS
             Syn.: Xenylla nova-zealandia(sic) Skarzynski, D, Piwnik, A et Porco, D, 2018:31 ISS
             Syn.: Xenylla nova-zealandia(sic) Skarzynski, D, Piwnik, A et Porco, D, 2018:32 ISS
             Syn.: Xenylla nova-zealandia(sic) Skarzynski, D, Piwnik, A et Porco, D, 2018:33 ISS
             Syn.: Xenylla nova-zealandia(sic) Skarzynski, D, Piwnik, A et Porco, D, 2018:34 ISS
             Syn.: Xenylla nova-zealandia(sic) Skarzynski, D, Piwnik, A et Porco, D, 2018:38 ISS
             Syn.: Xenylla nova-zealandia(sic) Skarzynski, D, Piwnik, A et Porco, D, 2018:39 ISS
            Syn.: Xenylla maritima ssp. meridionalis Stach, J, 1949:231
            Syn.: Xenylla schillei Delamare Deboutteville & Cassagnau, 1951
            Syn.: Xenylla brevisimilis ssp. brevisimilis da Gama, 1969



Lipura(lapsus) maritima from the UK
After Lubbock J, 1873 Pl.47 Fig.1
Xenylla maritima from the UK
2006 © Valentine., B
Xenylla maritima from the UK
2006.10.01 © Valentine., B
Xenylla maritima from the UK
2007.07.01 © Cornwall, N.J.
Xenylla maritima from the UK
2007.07.29 © Valentine., B
Xenylla maritima from the UK
2007.10.01 © Valentine., B
Xenylla maritima from the UK
2007.10.02 © Valentine., B
Xenylla maritima from the UK
2007.10.03 © Valentine., B
Xenylla maritima from the UK
2008.08.11 © Valentine., B
Xenylla maritima from the UK
Five single eyes
2008.08.15 © Valentine., B
Xenylla maritima from the UK
2009.08.29 © Valentine., B
Xenylla maritima from the UK
2009.09.17 © Valentine., B
Xenylla maritima from the UK
2015.08.25 © Valentine., B
Xenylla maritima from Holland
2015.11.27 © Wieringa, P.

             Syn.: Xenylla maritime(sic) Yadav, RS, 2017:1339 ISS
Xenylla maritima from Holland
2019.02.07 © Kamsteeg, G.
Xenylla maritima from Holland
2019.02.07 © Kamsteeg, G.
Xenylla maritima from Holland
Furca with narrow pointed mucrones fused with dentes
2019.02.07 © Kamsteeg, G.

             Syn.: Xenylla maritime(sic) Budha, PB et Shrestha, P, 2021:26 ISS




Species mediterranea da Gama, 1964

            Syn.: Xenylla brevicauda f. atypica Denis, 1927
            Syn.: Xenylla brevicauda f. atypica Denis, 1931
            Syn.: Xenylla brevicauda f. atypica Denis, J-R, 1933:320
            Syn.: Xenylla grisea f.. bidentata Denis, 1931


Species nitida Tullberg, 1871

            Syn.: Xenylla pseudomaritima James, 1933 =?
            Syn.: Xenylla brevisimilis Skarzynski, D, Piwnik, A et Porco, D, 2018:31
            Syn.: Xenylla maritima Babenko & al, 1994 partim
            Syn.: Xenylla maritima Thibaud al, 2004 partim


Species pomorskii Skarzynski, D, Piwnik, A et Porco, D, 2018:40

            Syn.: Xenylla maritima Stach, J, 1949:227 partim
            Syn.: Xenylla maritima Babenko & al, 1994 partim
            Syn.: Xenylla maritima Tibaud & al, 2004 partim
            Syn.: Xenylla sp.2 Skarzynski, D, Piwnik, A et Porco, D, 2018:34


Species szeptyckii Skarzynski, D, Piwnik, A et Porco, D, 2018:39

            Syn.: Xenylla maritima Stach, J, 1949:227 partim
            Syn.: Xenylla maritima Babenko & al, 1994 partim
            Syn.: Xenylla maritima Tibaud & al, 2004 partim
            Syn.: Xenylla nitida Skarzynski, D, Piwnik, A et Porco, D, 2018:40
            Syn.: Xenylla sp.1 Skarzynski, D, Piwnik, A et Porco, D, 2018:34


Xenylla szeptyckii from Iran
After Yahyapour E & al, 2021 Fig.3


Genus Cosberella Wray, DL, 1963:103 key

F
  Syn.: Hypogastruridae Proxenyllodes Christiansen, K et Bellinger, P, 1980:51
  Syn.: Hypogastruridae Proxenyllodes Christiansen, K et Bellinger, P, 1998:50
  Syn.: Hypogastrura Mucrella Fjellberg, 1985
  Syn.: Hypogastrura Mucronella(sic) Skidmore, RE, 1995:47 ISS

Species acuminata (Cassagnau, P, 1952:306) Bernard, 2006

            Syn.: Proxenyllodes folsomi Denis, 1926
            Syn.: Hypogastrura nodosa Steiner, 1958,si
            Syn.: Hypogastrura minuta Cassagnau, 1965 =?
            Syn.: Hypogastrura ornata Palissa, 1968
            Syn.: Hypogastrura (Ceratophysella) acuminata f. hipergranulata Jordana, R, 1980:16


             Syn.: Hypogastrura accuminata(sic) Jordana, R, 1980:9 ISS

Species arborea (Fjellberg, 1992) Bernard, 2006

Species conatoa Wray, DL, 1963:103, t.t.

Species denali (Fjellberg, 1985) Bernard, 2006

Species hibernica (Fjellberg, A, 1987:289) Bernard, 2006


Cosberella lamaralexanderi
2006 © Bernard, E.
Species lamaralexanderi Bernard, 2006

Species mendozarum Palacios-Vargas, JG et Castaño-Meneses, G, 2019:16

Species navicularis (Schött, H, 1893:83) Bernard, 2006

Species yoshiana (Babenko, 2000) Bernard, 2006


Genus Jacutogastrura Martynova, 1981

F
Species silvatica Martynova, 1981, t.t.


Genus Neobeckerella Wray, 1952

F
Species allusa Wray, 1952, t.t.


Genus Octoacanthella Martynova, 1961

F
Species aethiopica Martynova, 1961, t.t.


Genus Ongulogastrura Thibaud & Massoud, 1983

F
Species longisensilla Thibaud & Massoud, 1983, t.t.


Genus Ecuadogastrura Palacios-Vargas, JG et Thibaud, J-M, 2001:27

Species trinkleini Palacios-Vargas, JG et Thibaud, J-M, 2001:28, t.t.


Genus Stenogastrura Christiansen & Bellinger, 1980

F
Species hiemalis Christiansen & Bellinger, 1980:207, t.t.


Genus Gomphiocephalus Carpenter, 1908

M
  Syn.: Hypogastruridae Comphiocephalus(sic) Salmon, JT, 1964:199 ISS
Remark by Janssens, F. 2012.11.15: According to Greenslade & al. (2011), Hypogastruridae is not monophyletic and Gomphiocephalus is the most derived "hypogastrurid" genus, and sister group of (Brachystomellidae + Neanuridae). Therefore, to partially resolve the paraphyly of the Hypogastruridae, we tentatively propose to place Gomphiocephalus in a monogenic family Gomphiocephalidae most basic in Neanuroidea. Note: this is not yet reflected in the current taxonomic hierarchy.


Species hodgsoni Carpenter, 1908, t.t.

Gomphiocephalus hodgsoni from Antarctica
After Carpenter GH, 1921 Pl.I Fig.1
Gomphiocephalus hodgsoni from Antarctica
After Carpenter GH, 1921 Pl.I Fig.2
Gomphiocephalus hodgsoni from Antarctica
Eyepatch with ocelli A-H
Modified after Meyer-Rochow VB & al, 2005 Fig.3
Gomphiocephalus hodgsoni from Antarctica
2008.05.13 © O'Brien., B
Gomphiocephalus hodgsoni from Antarctica
2010 © Stevens, M.
Gomphiocephalus hodgsoni from Antarctica
After Greenslade P & al, 2011 Fig.9
             Syn.: Gomphiocephalus hodysoni(sic) Fiera, C, 2013:71 ISS

References