http://www.collembola.org/taxa/lepiinae.htm - Last updated on 2019.07.26 by Frans Janssens
Checklist of the Collembola: Lepidocyrtinae

Subfamilia Lepidocyrtinae Wahlgren E, 1906:67 key

Genus Number
of species
Acanthurella9
Acrocyrtus28
Bromacanthus12
Lepidiaphanus2
Lepidocyrtus (Allocyrtus)1
Lepidocyrtus (Ascocyrtus)9
Lepidocyrtus (Cinctocyrtus)11
Lepidocyrtus (Fractocyrtus)2
Lepidocyrtus (Lanocyrtus)30
Lepidocyrtus (Lepidocyrtus)165
Lepidocyrtus (Setogaster)15
Lepidocyrtus243
Lepidonella16
Metasinella (Metasinella)1
Metasinella (Sulcuncus)10
Metasinella11
Pseudocyrtus3
Pseudosinella353
Rambutsinella3
Rhynchocyrtus1
Sinelloides2
Trichorypha1
Vietsira1
Lepidocyrtinae685
Genera with number of species
Lepidocyrtinae from Sweden
2006.03.25 © Hall, K.
Lepidocyrtinae from Singapore
2014.01.22 © Bay, N.


Tribus Lepidocyrtini Gisin, 1963 key

Lepidocyrtini from Singapore
2018.12.12 © Weishou.


Genus Vietsira Yoshii, 1994

Species squamigera Yoshii, 1994, t.t.


Genus Lepidocyrtus Bourlet, C, 1839:391 key  key  key  key

Lepidocyrtus sp. from the UK
After Beck R, 1862 Plate X Fig A.
Lepidocyrtus sp. from the UK
Group of dorsal scales (150x)
After Beck R, 1862 Plate X Fig B.
Lepidocyrtus sp. from the UK
Scale surface markings (1300x)
After Beck R, 1862 Plate X Fig 1.
Lepidocyrtus sp. from the UK
'Podura scale' microscopy test object slide'
Made by Watson from London, 1860-1900
2010.11.07 © Walker, D.
Lepidocyrtus sp. from the UK
'Podura scale' microscopy test object slide'
Scale surface markings (1000x, phase)
2010.11.02 © Walker, D.




  Syn.: Podurida(sic) Lepidocyrthus(sic) Kolenati, FA, 1858:131 ISS
  Syn.: ... Lepidocystus(sic) Mari Mutt, JA, 1987:366 ISS
  Syn.: Entomobryidae Lepidocirtus(sic) Delgadillo-Reyes, J et Padilla-Ramírez, J, 2004:306 ISS
Lepidocyrtus sp. from the UK
Lepidocyrtus sp. riding Lasius niger
2005 © Valentine., B
Lepidocyrtus sp. from Sweden
2005.12.01 © Hall, K.
Lepidocyrtus sp. from Estonia
2006.10.01 © Tartes, U.
Lepidocyrtus sp. from the UK
2006.11.22 © Valentine., B
Lepidocyrtus sp. from the UK
2006.11.22 © Valentine., B
Lepidocyrtus sp. from Sweden
2006 © Hall, K.
Lepidocyrtus sp. from France
2006 © Jacquemin, G.
Lepidocyrtus sp. from Germany
2006.12.28 © Kohl, F.
Remark by ICZN. 2007:443: Placed on the Official List of Names in Zoology. Opinion 291.

Lepidocyrtus sp. from the UK
2007 © Barton, T.
Lepidocyrtus sp. from the UK
2007 © Cornwall, N.J.
Lepidocyrtus sp. from the UK
2007.02.26 © Botting, J.
Lepidocyrtus sp. from the USA
2007.03.03 © Murray, T.
Lepidocyrtus sp. from the USA
2007.03.17.25 © Murray; T.
Lepidocyrtus sp. from the USA
2007.05.20 © Murray, T.
Lepidocyrtus sp. from Czechia
2007.09.08 © Krásenský P.
Lepidocyrtus sp. from the UK
2007.09.29 © Valentine., B
Lepidocyrtus sp. from the UK
2007.11.10 © Valentine., B
Lepidocyrtus sp. from the UK
2007.11.10 © Valentine., B
Lepidocyrtus sp. from the UK
2008.01.01 © Barton, T.
Lepidocyrtus sp. from the UK
2008.01.01 © Barton, T.
Lepidocyrtus sp. from the UK
2008.01.11 © Farrell, S.
Lepidocyrtus sp. from the UK
2008.12.01 © Robertson, A.
Lepidocyrtus sp. from Czechia
2009.01.31 © Krásenský, P.
Lepidocyrtus sp. from the UK
2009.03.06 © Barton, T.
Lepidocyrtus sp. from the UK
2009.03.23 © Robertson, A.
Lepidocyrtus sp. from the UK
2009.10.25 © Barton, T.
Lepidocyrtus sp. from the UK
2009.11.02 © Barton, T.
Lepidocyrtus sp. from Canada
2011.10.02 © Hochhalter, R.
Lepidocyrtus sp. from the USA
2012.04.08 © Moon, S.
Lepidocyrtus sp. from Portugal
2012.04.20 © Coelho, J.
Lepidocyrtus sp. from Portugal
2012.10.13 © Murray, A.
Lepidocyrtus sp. from Australia
2013.02.18 © Steve & Alison.
Lepidocyrtus sp. from Singapore
2013.12.21 © Koh, J.
Lepidocyrtus sp. from New Zealand
With spermatophore
2014.03.04 © Murray, A.

  Syn.: Entomobryidae Lepidocytrus(sic) Katz, AD, Taylor, SJ, Soto-Adames, FN, Addison, A, Hoese, GB, Sutton, MR et Toulkeridis, T, 2016:113 ISS


Species sp.1 Zeppelini, D, Bellini, BC, Creao-Duarte, AJ et Hernández, MIM, 2009:1166

Species sp.2 Soto-Adames, FN et Anderson, EW, 2017:35

Species sp.3 Ernesto, MV, Liberal, CN, Ferreira, AS, Alves, ACF, Zeppelini, D, Martins, CF, Pereira-Colavite, A, Creao-Duarte, AJ, Vasconcellos, A, 2018:5

Species sp.4 Ernesto, MV, Liberal, CN, Ferreira, AS, Alves, ACF, Zeppelini, D, Martins, CF, Pereira-Colavite, A, Creao-Duarte, AJ, Vasconcellos, A, 2018:5

Species ambricus Handschin, 1926 -

Species exploratorius Carpenter, 1924

Species fimetarius Gisin, H, 1964:390

Lepidocyrtus fimetarius from Europe
Cephalic dorsal chaetotaxy
After Gisin H, 1964 Fig.1 right
Lepidocyrtus fimetarius from India
After Santeshwari, Raghuraman M & Singh J, 2015 p.574

Species inormatus Folsom, JW, 1932:68

Species parallelus Folsom, 1924

Species pilifer Schött, 1901

Subgenus Lanocyrtus Yoshii, R et Suhardjono, YR, 1989:58


Lanocyrtus sp. from Taiwan
Sensillum of 4th antennal segment
2006.10.30 © Chen, J.-J.
Lanocyrtus sp. from Taiwan
5th instar
2009.04.24 © Chen, J.-J.
Lanocyrtus sp. from Taiwan
2009.05.11 © Chen, J.-J.
 
Lanocyrtus sp. from Taiwan
Mucro
2009.05.11 © Chen, J.-J.
Lepidocyrtus (Lanocyrtus) cyaneus from the UK
Scaleless basal antennal segments
2012.01.10 © Jones, A.
Lepidocyrtus (Lanocyrtus) sp. from Canada
2013.01.18 © Kyron, T.F.

Species absens Zhang, F, Chatterjee, T et Chen, J-X, 2009:44

Lepidocyrtus (Lanocyrtus) absens from India
After Zhang F, Chatterjee T & Chen J-X, 2009 Fig.1

Species altmontanus Tshelnokov, 1978

Species beaucatcheri (Wray, 1946) DuRant & Fox, 1966

Species biphasis Mari Mutt, JA, 1986:10

Species caeruleicornis Bonet, 1930

Species cinereus Folsom, JW, 1924:9

Lepidocyrtus cyaneus var. cinereus from the USA
After Folsom, J.W., 1924 Fig.24.

            Syn.: Lepidocyrtus cyaneus f. pallidus Gisin, 1944 =?
             Syn.: Lepidocyrtus cyaneus f. pallida(sic) Maynard, EA, 1951:183 =? ISS
            Syn.: Lepidocyrtus pallidus Christiansen, K et Bellinger, P, 1998:1058 partim =?



Lepidocyrtus cyaneus f. cinereus from the USA
After Maynard EA, 1951 PlateXIX Fig.322
Lepidocyrtus cinereus from the USA
2007.02.22 © Bernard, E.C.
Lepidocyrtus cinereus from the USA
2016.09.19 © Rosenfeld, J.
Lepidocyrtus cinereus from the USA
2016.09.20 © Rosenfeld, J.

Species dubius Christiansen, K et Bellinger, P, 1998:1045

Species felipei Wang, F, Chen, J-X et Christiansen, K, 2003:825

             Syn.: Lepidocyrtus (Lanocyrtus) felpei(sic) Wang, F, Chen, J-X et Christiansen, K, 2003:823 ISS

Species fernandi Christiansen, K et Bellinger, P, 1998:1047

Species fimicolus Mari Mutt, JA, 1988:200

            Syn.: Lepidocyrtus pallidus Christiansen, K et Bellinger, P, 1998:1058 partim


Lepidocyrtus fimicolus from the USA
2012.01.19 © Kouri, J.
Det. Soto-Adames, F.N., 2012.08.18
Lepidocyrtus fimicolus? from the USA
2012.02.05 © Allen, K.

Species finus Christiansen, K et Bellinger, P, 1980:943

Species floridensis Snider, RJ, 1967:14

Lepidocyrtus floridensis from the USA
2014.12.17 © Snyder, S.L.

Species gumgum Yoshii, R, 1982:35

Lepidocyrtus (Lepidocyrtus) gumgum from Malaysia
After Yoshii R, 1982 Fig.20A

Species helenae Snider, RJ, 1967:12

            Syn.: Lepidocyrtus aenescens Guthrie, JE, 1903:89
            Syn.: Lepidocyrtus cyaneus var. aenescens Mills, HB, 1934:73
            Syn.: Lepidocyrtus aenescens Maynard, EA, 1951:183
            Syn.: Lepidocyrtus cyaneus var. aenescens Maynard, EA, 1951:183
            Syn.: Lepidocyrtus cyaneus f. assimilis Maynard, EA, 1951:183
Lepidocyrtus cyaneus f. assimilis from the USA
After Maynard EA, 1951 PlateXIX Fig.323

            Syn.: Lepidocyrtus cyaneus var. assimilis Christiansen, K et Bellinger, P, 1980:946 =?
            Syn.: Lepidocyrtus cyaneus var. assimilis Christiansen, K et Bellinger, P, 1998:1052 =?


Species pseudofimetarius Gruia, M, 1983:196

Species simsim Yoshii, R, 1982:37

Lepidocyrtus (Lepidocyrtus) simsim from Malaysia
After Yoshii R, 1982 Fig.21A

Species vestitus Handschin, 1930

             Syn.: Lepidocyrtus (Lanocyrtus) vistitus(sic) Wang, F, Chen, J-X et Christiansen, K, 2003:826 ISS

Species-group lusitanicus Mateos E, 2008:?
Lepidocyrtus lusitanicus-group
Mesothorax not protruded
After Mateos E & al, 2018 Fig.1A



Species bilobatus Mateos, E, 2008:39

            Syn.: Lepidocyrtus bilobatus f. A Mateos, E, 2008:45
Lepidocyrtus bilobatus form A from Spain
After Mateos E, 2008 Fig.2
Lepidocyrtus bilobatus form A
from Spain
After Mateos E, 2008 Fig.30

            Syn.: Lepidocyrtus bilobatus f. B Mateos, E, 2008:45
Lepidocyrtus bilobatus form B from Spain
After Mateos E, 2008 Fig.3
Lepidocyrtus bilobatus form B
from Spain
After Mateos E, 2008 Fig.32



Species lusitanicus da Gama, 1964

            Syn.: Lepidocyrtus lusitanicus f. A Mateos, E, 2008:47
Lepidocyrtus lusitanicus form A
from Spain
After Mateos E, 2008 Fig.25
Lepidocyrtus lusitanicus form A
from Spain
After Mateos E, 2008 Fig.34




Lepidocyrtus lusitanicus ssp. lusitanicus
from Spain
After Simón-Benito, J.C., 2007 Fig.1.
Lepidocyrtus lusitanicus ssp. lusitanicus
from Spain
After Mateos E, 2008 Fig.21
Lepidocyrtus lusitanicus ssp. lusitanicus
from Spain
After Mateos E, 2008 Fig.30

  Subspecies coloratus da Gama, 1964

Lepidocyrtus lusitanicus ssp. coloratus
from Spain
After Simón-Benito, J.C., 2007 Fig.7.




             Syn.: Lepidocyrtus lusitanicus ssp. colorata(sic) Mateos, E, 2008:49 ISS


Lepidocyrtus lusitanicus ssp. coloratus
from Spain
After Mateos E, 2008 Fig.22
Lepidocyrtus lusitanicus ssp. coloratus
from Spain
After Mateos E, 2008 Fig.31

  Subspecies nigrus Simón-Benito, JC, 2007:322

Lepidocyrtus lusitanicus ssp. nigrus
from Spain
After Simón-Benito, J.C., 2007 Fig.12.
Lepidocyrtus lusitanicus ssp. nigrus
from Spain
After Mateos E, 2008 Fig.23
Lepidocyrtus lusitanicus ssp. nigrus
from Spain
After Mateos E, 2008 Fig.32

  Subspecies piezoensis Simón-Benito, JC, 2007:322

Lepidocyrtus lusitanicus ssp. piezoensis
from Spain
After Simón-Benito, J.C., 2007 Fig.16.
Lepidocyrtus lusitanicus ssp. piezoensis
from Spain
After Mateos E, 2008 Fig.24
Lepidocyrtus lusitanicus ssp. piezoensis
from Spain
After Mateos E, 2008 Fig.33
Lepidocyrtus lusitanicus ssp. piezoensis from the Azores
2010.09.22 © E.D.E.N.

Species selvaticus Arbea, JI et Ariza, E, 2007:87

Lepidocyrtus selvaticus from Spain
After Arbea JI & Ariza E, 2007 Fig.1
Lepidocyrtus selvaticus from Spain
After Mateos E, 2008 Fig.28
Lepidocyrtus selvaticus
from Spain
After Mateos E, 2008 Fig.30

Species-group lanuginosus Mateos E, 2012:79
            Syn.: Lepidocyrtus lanuginosus-cyaneus-group Gisin, 1964



Lepidocyrtus (Lanocyrtus) cyaneus from Belgium
Interocular macrosetae (*)
2017.09.12 © Huskens, M.-L.
Lepidocyrtus lanuginosus-group
Mesothorax not protruded
After Mateos E & al, 2018 Fig.1A

Species bicoloris Mateos, E, 2012:79

Lepidocyrtus bicoloris from Spain
After Mateos E, 2012 Fig.3
            Syn.: Lepidocyrtus violaceus Lubbock, J, 1873:154 =?
Lepidocyrtus violaceus from the UK
After Lubbock J, 1873 Pl.28
Lepidocyrtus violaceus from the UK
Body scale with short ribs
After Beck J in Lubbock J, 1873 Pl.75




Lepidocyrtus bicoloris? from Norway
2007.08.04 © Fauskanger, I.

Species cyaneus Tullberg, 1871, B

            Syn.: Podura tota argenteo-aurata(sic) Olafsen & Povelsen, 1772 =?
            Syn.: Podura pusilla Mohr, 1786 =?
            Syn.: Podura pusilla Börner, C, 1906:155 =?
            Syn.: Podura pusilla Bödvarsson, H, 1957:32 =?
            Syn.: Lepidocyrtus metallicus Folsom, JW, 1919:15A
            Syn.: Lepidocyrtus metallicus Folsom, JW, 1924:10
            Syn.: Lepidocyrtus metallicus Mills, HB, 1934:72
            Syn.: Lepidocyrtus metallicus Maynard, EA, 1951:182
            Syn.: Lepidocyrtus purpureus Lubbock, J, 1873:155
Lepidocyrtus purpureus from the UK
After Lubbock J, 1873 Pl.30
Lepidocyrtus purpureus from the USA
Adult and juvenile instar form
After Jackson, A.D., 1909 Pl.29 fig.1 & 6.

            Syn.: Lepidocyrtus violaceus Folsom, JW, 1919:14A
            Syn.: Lepidocyrtus violaceus Handschin, E in Dahl, F, 1929:86
            Syn.: Lepidocyrtus violaceus Kloet, GS et Hincks, WD, 1945:8
            Syn.: Lepidocyrtus violaceus Palissa, A in Brohmer, P, Ehrmann, P et Ulmer, G, 1964:221
            Syn.: Lepidocyrtus assimilis Reuter, 1890
Lepidocyrtus cyaneus var. assimilis from Finnland
After Schött H, 1893 Taf.III Fig.10

            Syn.: Lepidocyrtus pallidus Reuter, 1890
            Syn.: Lepidocyrtus elegantulus Meinert, 1896
            Syn.: Lepidocyrtus vicarius Börner, C, 1906:174
            Syn.: Lepidocyrtus anglicanus Jackson, 1926
            Syn.: Lepidocyrtus serbicus Denis, 1936 =?
            Syn.: Lepidocyrtus cyaneus f. principalis Gisin, 1944



Lepidocyrtus cyaneus from Europe
Scale apically rounded
After Kolbe HJ, 1893 Fig.42
Lepidocyrtus cyaneus from Japan
After Yosii R, 1942 Abb.15.1
Lepidocyrtus cyaneus from the USA
After Maynard EA, 1951 PlateI Fig.13
Lepidocyrtus cyaneus from the USA
After Maynard EA, 1951 PlateXIX Fig.319
Lepidocyrtus cyaneus from Europe
Cephalic dorsal chaetotaxy
After Gisin H, 1964 Fig.1 midleft
Lepidocyrtus cyaneus from the UK
2001 © Hopkin, S.P.
Lepidocyrtus cyaneus from Japan
After Suetsugu N, Satoh F & Koike T, 2010 Photo46
Lepidocyrtus cyaneus from the UK
After Mateos E, 2012 Fig.4
Lepidocyrtus cyaneus from the UK
2012.12.12 © Jones, A.
Lepidocyrtus cyaneus from Holland
2012.10.02 © van Duinen, J.
Lepidocyrtus (Lanocyrtus) cyaneus from Belgium
Interocular macrosetae (*)
2017.09.12 © Huskens, M.-L.
Lepidocyrtus (Lanocyrtus) cyaneus from Belgium
Interocular macrosetae (*)
2018.01.12 © Huskens, M.-L.
Lepidocyrtus (Lanocyrtus) cyaneus juv. from Belgium
With transparant scale cover
2018.01.12 © Huskens, M.-L.
Lepidocyrtus (Lanocyrtus) cyaneus juv. from Belgium
Grooming
2018.01.12 © Huskens, M.-L.
Lepidocyrtus (Lanocyrtus) cyaneus from France
Interocular macrosetae (*)
2018.02.10 © Picard, J.
Lepidocyrtus (Lanocyrtus) cyaneus from the UK
Descaled specimen
2018.10.31 © Marquis, A.

Species lanuginosus (Linnaeus in Gmelin, 1788) Lubbock, J, 1870:300, t.t., B

            Syn.: Podura pusilla von Linné, C, 1767:1014
            Syn.: Paidium æneum(sic) Koch, 1841 IOS
            Syn.: Cyphodeirus(sic) agilis Nicolet, H, 1842:66
            Syn.: Paidium flaveolum Koch, 1841
            Syn.: Lepidocyrtus argentatus Bourlet, 1842
            Syn.: Lepidocyrtus uvularis Bourlet, 1842
            Syn.: Cyphodeirus(sic) Æneus(sic) Nicolet, H, 1842:66 IOS
             Syn.: Cyphoderus æneus(sic) Nicolet, H, 1847:383 ISS
             Syn.: Lepidocyrtus æneus(sic) Lubbock, J, 1870:300 ISS
             Syn.: Lepidocyrtus æneus(sic) Lubbock, J, 1873:154 ISS
Lepidocyrtus aeneus from the UK
After Lubbock J, 1873 Pl.29

            Syn.: Cyphodeirus(sic) gibbulus Nicolet, H, 1842:64
Lepidocyrtus gibbulus from the UK
After Lubbock J, 1870 Pl.XXII Fig.22
Lepidocyrtus gibbulus from the UK
After Lubbock J, 1873 Pl.27

            Syn.: Cyphodeirus(sic) parvulus Nicolet, H, 1842:67
            Syn.: Lepidocyrtus rivularis Bourlet, 1843
            Syn.: Isotoma pulex Gervais in Walckenaër, 1844 =?
Isotome puce from France
Isotoma pulex
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.10

            Syn.: Lepidocyrtus fucatus Uzel, 1891
            Syn.: Lepidocyrtus montanus Carl, 1901
            Syn.: Lepidocyrtus lanuginosus f. ruber Schött, 1902
            Syn.: Lepidocyrtus vespensis Womersley, 1930



Lepidocyrtus lanuginosus from Europe
Cephalic dorsal chaetotaxy
After Gisin H, 1964 Fig.1 midleft

             Syn.: Lepidocyrtus lanunginosus(sic) Szeptycki & Weiner in Razowski, 1990 ISS
             Syn.: Lepidocyrtus languinosa(sic) Poole in Poole & Gentili, 1997 ISS
             Syn.: Lepidocyrtus lanuginosum(sic) Jaksic, P, 2005:194 ISS
Lepidocyrtus lanuginosus from the UK
After Mateos E, 2012 Fig.1
Lepidocyrtus lanuginosus from the UK
Head pigmentation
After Mateos E, 2012 Fig.2
Lepidocyrtus (Lanocyrtus) lanuginosus from Belgium
2017.02.17 © Huskens, M.-L.
Lepidocyrtus (Lanocyrtus) lanuginosus from Belgium
Distinctly bluish antennae
2017.06.05 © Huskens, M.-L.
Lepidocyrtus (Lanocyrtus) lanuginosus from Belgium
Interocular macrosetae (*)
2018.02.03 © Huskens, M.-L.


Species-group pallidus-serbicus Winkler & Traser, 2012:?
Lepidocyrtus pallidus-serbicus-group
Mesothorax not protruded
After Mateos E & al, 2018 Fig.1A

Species-subgroup pallidus Gisin, 1965




Species pallidus Reuter, 1890, B

            Syn.: Lepidocyrtus cyaneus Folsom, 1932
            Syn.: Lepidocyrtus serbicus Gisin, H, 1965:522
            Syn.: Lepidocyrtus cyaneus Heungens, A, 1967:243,B
            Syn.: Lepidocyrtus ruber Fjellberg, 1980 partim


Lepidocyrtus pallidus from the USA
After Christiansen K & Bellinger P, 1998 Fig.776A
Lepidocyrtus pallidus? from the USA
2016.02.23 © Snyder, S.L.
Lepidocyrtus pallidus? from the USA
Antennal segments apically ringed
2016.02.23 © Snyder, S.L.

Species weidneri Hüther, W, 1971:159

            Syn.: Lepidocyrtus ruber Fjellberg, 1980 partim



Species-subgroup serbicus Winkler D, 2017:538 key
Species isabelleae Winkler, D, 2017:530

Lepidocyrtus isabelleae from Hungary
After Winkler D, 2017 Fig.1
Lepidocyrtus isabelleae from Hungary
Dorsal chaetotaxy
After Winkler D, 2017 Fig.6

Species pseudosinelloides Gisin, 1967

Species serbicus Denis, 1933

Species tomosvaryi Winkler & Traser, 2012


Species-subgroup arrabonicus Winkler D, 2017:539
Species arrabonicus Traser, G, 2000:65


Subgenus Lepidocyrtus Bourlet C, 1839:391, n.c., s.s.
  Syn.: Thysanoura Podura Say, 1821 partim
  Syn.: ... Paidium Koch, 1840 =?
  Syn.: Entomobryidæ(sic) Strongylonotus MacGillivray, AD, 1894:107


  Syn.: Lepidocyrtus curvicollis-violaceus-group Gisin, 1964
Lepidocyrtus (Lepidocyrtus) sp. from Norway
Ant.1 & ant.2 with scales
2009.12.05 © Haraldseide, H.
Lepidocyrtus (Lepidocyrtus) lignorum from the UK
Ant.1 & ant.2 with scales
2010.01.31 © Robertson, A.
Lepidocyrtus curvicollis & paradoxus from France
2011.09.10 © Dehalleux, A.



Species achromatis Maynard, EA, 1951:174

Species agraensis Baijal & Sinha, 1971

Lepidocyrtus agraensis from India
After Santeshwari, Raghuraman M & Singh J, 2015 p.574

Species aho Christiansen, K et Bellinger, P, 1992:278

Species albicaudatus Hammer, 1953

Species albidus Stach, 1920

Species annulicornis Carpenter GH, 1916:40, nec Börner

Species apo Christiansen, K et Bellinger, P, 1992:279

Species arctomontanus Tshelnokov, 1990

Species assymetrica (Salmon, JT, 1937:355)

Entomobrya assymetrica from New Zealand
After Salmon JT, 1937 Pl.50 Fig.a
Pseudosinella assymetrica from New Zealand
After Salmon JT, 1941 Pl.66 Fig.431

Species atratus Mari Mutt, JA, 1983:54

Species atropurpureus Packard, 1888

Species aurantiacus Maynard, EA, 1951:176

Lepidocyrtus aurantiacus from the USA
After Maynard EA, 1951 PlateI Fig.12

  Subspecies rubidus Maynard, EA, 1951:177

Lepidocyrtus aurantiacus f. rubidus from the USA
After Maynard EA, 1951 PlateXIX Fig.308
Lepidocyrtus aurantiacus f. rubidus from the USA
After Maynard EA, 1951 PlateXIX Fig.309

Species aureus Womersley, 1934

Species balteatus Mari Mutt, JA, 1983:53

Species bidenticulatus Ritter, 1911

Species caeruleacrurus (Salmon, JT, 1941:370)

Mesira caeruleacrura from New Zealand
After Salmon JT, 1941 Pl.61 Fig.355

Species caeruleus Ritter, 1911

Species ceratoxenus Börner, 1907

Species coerulea Denis, 1941

Species coeruleocinctus Handschin, 1930

Species concolourus Nguyen, 2001

Species congolensis Marlier, 1945

Species crassicornus (Koch & Berendt, 1854 -) -

Species dealbatus Barra, J-A, 2006:71

Species denisi Marlier, 1945

Species dentilus Winter, 1963, n.n.

Species diminutus Mari Mutt, JA, 1986:16

Species dispar Mari Mutt, JA, 1986:11

Species distinctus Mari Mutt, JA, 1986:12

Species eeu Christiansen, K et Bellinger, P, 1992:281

             Syn.: Lepidocyrtus eou(sic) Christiansen, K et Bellinger, P, 1994:342 ISS

Species elongatus (Salmon, JT, 1944:156)

Species epiphytus Winter, 1963, n.n.

Species exaggeratus Denis, 1931

Species extensus Wahlgren, 1908

Species ferrugineus Schött, H, 1893:13

Species fimbriatus Salmon, JT, 1944:159

Species finensis Maynard, EA, 1951:175

Lepidocyrtus finensis from the USA
2006 © Grinnell College.

Species fryeri Carpenter, GH, 1916:40

            Syn.: Lepidocyrtus Fryeri(sic) Carpenter, GH, 1916:40 IOS

Species fuscatus Wahlgren, 1908

Species gardineri Carpenter, GH, 1916:41

            Syn.: Lepidocyrtus Gardineri(sic) Carpenter, GH, 1916:41 IOS

Species geayi Denis, J-R, 1924:258

            Syn.: Lepidocyrtus Geayi(sic) Denis, J-R, 1924:258 IOS
             Syn.: Lepidocyrtus gaeyi(sic) Bernard, EC, Soto-Adames, FN et Wynne, JJ, 2015:256 ISS



Species geayides Denis, 1931

Species griseolus Mari Mutt, JA, 1986:15

Species guthriei Maynard, EA, 1951:181

Lepidocyrtus guthriei from the USA
After Maynard EA, 1951 PlateI Fig.9
            Syn.: Lepidocyrtus purpureus Guthrie, JE, 1903:87



Species hakea Christiansen, K et Bellinger, P, 1992:284

Species heterophthalmus Carpenter, 1904

             Syn.: Lepidocyrtus heteropthalmus(sic) Christiansen & Bellinger, 1991 ISS

Species himalayanus Yosii, 1971

Species hirtus Christiansen, K et Bellinger, P, 1980:947

Species hukulii Christiansen, K et Bellinger, P, 1992:284

Species ianthinus Mari Mutt, JA, 1986:10

Species immaculatus Folsom, JW, 1932:68

Species imperialis Carpenter, GH, 1916:41

Species inornatus Folsom, 1932

Species iricolor (Say, 1821) Jackson, AD, 1909:531

Species kauriensis Salmon, JT, 1941:375

Lepidocyrtus kauriensis from New Zealand
After Salmon JT, 1941 Pl.62 Fig.373

Species kolymensis Tshelnokov, 1990

Species lamarcki Delamare Deboutteville, 1952

Species lepargus Mari Mutt, JA, 1986:14

Species lindensis Salmon, JT, 1941:371

Species longithorax Delamare Deboutteville, 1948

Species luteus Jackson, AD, 1909:533

Lepidocyrtus luteus from the USA
After Jackson, A.D., 1909 Pl.31 fig.1.

Species maldonadoi Mari Mutt, JA, 1986:13

Species marimutti Thibaud, J-M et Najt, J, 1987:934

Species maximus Schött, H, 1893:11

Species medioides Womersley, 1928

Species mele Christiansen, K et Bellinger, P, 1992:295

Species millsi Folsom in Snider, 1967:3

Species moorei Salmon, JT, 1941:373

Lepidocyrtus moori(sic) from New Zealand
After Salmon JT, 1941 Pl.62 Fig.366
Lepidocyrtus moorei from New Zealand
2013.10.19 © Kerr, S.

Species mucroserratus Salmon, 1954

Species mutabilis Denis, 1931

Lepidocyrtus mutabilis from Guatemala
After Ellis WN, 1967 Fig.4a

Species neofasciatus Wray, 1948

Lepidocyrtus neofasciatus from the USA
2010.07.10 © Moody, E.

Species nigrofasciatus Womersley, 1934

Species nilatus Mari Mutt, JA, 1987:366

Lepidocyrtus nilatus from Colombia
After Mari Mutt JA, 1987 Fig.18

Species obscuricornis Carpenter, GH, 1916:39

Species obscurus Stach, J, 1960:540

Species obtusus Wahlgren, E, 1906:14

Species olena Christiansen, K et Bellinger, P, 1992:299

Lepidocyrtus olena from Rapa Nui
After Bernard EC & al, 2015 Fig.7E

Species orientalis Handschin, E, 1929:241

Lepidocyrtus orientalis from South-India
After Handschin E, 1929 Fig.17
Lepidocyrtus orientalis from India
After Denis JR, 1936 Fig.31
Lepidocyrtus orientalis from India
After Yadav RS et al, 2018 Fig.1

Species orientalis (Tshelnokov, 1978) Wang, F, Chen, J-X & Christiansen, K, 2003:837

Species ornatus Yosii, 1966

Species paracaprilesi Mari Mutt, JA, 1988:197

Species pearsei Mills, HB in Pearse, AS, 1938:188

Species perturbans Denis, 1948

Species pistiae Delamare Deboutteville & Paulian, 1945

Species plumosus Carpenter, 1934

Species poko Christiansen, K et Bellinger, P, 1992:301

Species praecisus Schött, H, 1917:37

            Syn.: Lepidocyrtus præcisus(sic) Schött, H, 1917:37 IOS

Species pseudopictus Womersley, 1934

Species pulchellus Denis, 1926

Remark by Mateos, E. 2008:48: Poorly documented and status is questionable. Known only from the type locality, Campigna, Italy; described on the basis of only two specimens, one incomplete, and, curiously, Denis describes two different times this species as a new species (in, 1926 and, 1927).

Species pyriformus (Koch & Berendt, 1854 -) -

Species ramosi Mari Mutt, JA, 1986:13

Species rataensis Salmon, JT, 1941:375

Lepidocyrtus rataensis from New Zealand
After Salmon JT, 1941 Pl.63 Fig.378

Species robustus Imms, 1912

Species schmidti Handschin, 1927

Species sejmczanicus Tshelnokov, 1990

Species sibiricus Rusek, 1985

Species silvestris Carpenter, GH, 1916:39

Species simularis Brown, JM, 1926:211

Species sp. Palacios-Vargas, 1981:90

Species sp. da Gama, MM, 1984:59

Species sp. Maes & Palacios-Vargas, 1988:5

Species sp. Blackith & Disney, 1988

Species sp. Palacios-Vargas, 1990:28

Species sp. Vázquez & Palacios-Vargas, 1990:17

Species sp. Palacios-Vargas & Gómez, 1991

Species sp. Terrón-Sierra & Palacios-Vargas, 1991

Species sp. Palacios-Vargas & Gómez-Anaya, 1995:28

Species sp. Zeppelini & Castaño, 1995:10

Species sp. Thibaud, J-M et Weiner, WM in Najt, J et Matile, L, 1997:88

Species sp. Loranger, G, Ponge, J-F, Blanchart, E et Lavelle, P, 1998:24

Species sp. Loranger, G, Ponge, J-F, Blanchart, E et Lavelle, P, 1998:24

Species sp. Loranger, G, Ponge, J-F, Blanchart, E et Lavelle, P, 1998:24

Species sp. Christiansen & Bellinger, 1991:209

Species sp. Christiansen & Bellinger, 1991:208

Species sp. Palacios-Vargas & Gómez-Anaya, 1995:27

Species sp. Christiansen & Bellinger, 1991:208

Species sp. Christiansen & Bellinger, 1991:208

Species sp. Christiansen & Bellinger, 1991:209

Species sp. Christiansen & Bellinger, 1991:209

Species sp. Dunger, 1977:182

Species sp. Dunger, 1977:182

Species sp. Christiansen & Bellinger, 1991:209

Species sp. Christiansen & Bellinger, 1991:209

Species sp. Christiansen & Bellinger, 1991:209

Species sp. Christiansen & Bellinger, 1994:342

Species sp. Christiansen & Bellinger, 1991:208

Species sp. Christiansen & Bellinger, 1991:208

Species sp. Christiansen & Bellinger, 1991:207, s.i.

Species sp. Christiansen & Bellinger, 1991:208

Species sp. Christiansen & Bellinger, 1994:342

Species stramineus Carpenter, GH, 1916:40

Species submontanus Salmon, JT, 1941:372

Lepidocyrtus submontanus from New Zealand
After Salmon JT, 1941 Pl.62 Fig.361
Lepidocyrtus submontanus from New Zealand
2014.02.27 © Murray, A.

Species summersi (MacGillivray, AD, 1894:107) Folsom, JW, 1927:9

            Syn.: Strongylonotus Summersii(sic) MacGillivray, AD, 1894:107 IOS
             Syn.: Lepidocyrtus summersii(sic) Mari Mutt, JA, 1979:541 ISS
             Syn.: Lepidocyrtus summersii(sic) Mari Mutt, JA, 1983:53 ISS



Species szeptyckii Rusek, 1985

Species tropicus Winter, 1963, n.n.

Species tundriensis Tshelnokov & Bondarenko, 1978

Species uku Christiansen, K et Bellinger, P, 1992:303

Species unafascius Salmon, JT, 1941:374

            Syn.: Lepidocyrtus unafascia(sic) Salmon, JT, 1941:374 IOS
Lepidocyrtus unafascia(sic) from New Zealand
After Salmon JT, 1941 Pl.62 Fig.372



Species unicolor Womersley, 1934

Species unifasciatus James, 1933

Species usitatus Folsom, JW, 1927:8

Species vespensis Womersley, 1930

Remark by Mateos, E. 2008:48: Poorly documented and status is questionable. Described as a new species from Britain; the description is very brief and does not indicate any chaetotaxic character making comparisons with other species impossible.

Species vexans Denis, 1933

Species vicarius Börner, C, 1906:174

Species vireticulus Mari Mutt, JA, 1986:15


Species-group lignorum Mateos E, 2011:37
            Syn.: Lepidocyrtus lignorum-violaceus-subgroup Gisin, 1964


Lepidocyrtus lignorum-group
Mesothorax protruded
After Mateos E & al, 2018 Fig.1B

Species barbulus Mateos, E, 2011:32

Lepidocyrtus barbulus from Greece
Unpigmented form
After Mateos E, 2011 Fig.18
Lepidocyrtus barbulus from Greece
Pigmented form
After Mateos E, 2011 Fig.19

            Syn.: Lepidocyrtus lignorum Ellis, WN, 1976:312 partim


Species chorus Mateos, E et Lukic, M, 2019:223

            Syn.: Lepidocyrtus (lignorum-group) sp. Mateos, E, Escuer, P, Busmachiu, G, Riutort, M et Alvarez-Presas, M, 2018:640
Lepidocyrtus chorus from Croatia
After Mateos E & Lukic M, 2019 Fig.3
Lepidocyrtus chorus from Croatia
After Mateos E & Lukic M, 2019 Fig.5
Lepidocyrtus chorus from Croatia
After Mateos E & Lukic M, 2019 Fig.6
Lepidocyrtus chorus from Croatia
Dancing behaviour
2019.01.24 © Mateos, E. & Lukic, M.



Species instratus Handschin, 1924

Lepidocyrtus instratus from Europe
Cephalic dorsal chaetotaxy
After Gisin H, 1964 Fig.1 left

Species intermedius Mateos E, Escuer P et Alvares-Presas M in Mateos, E, Escuer, P, Busmachiu, G, Riutort, M et Alvarez-Presas, M, 2018:644

Lepidocyrtus intermedius from Spain
Habitus
After Mateos E & al, 2018 Fig.3
Lepidocyrtus intermedius from Spain
Dorsal chaetotaxy
After Mateos E & al, 2018 Fig.8

Species juliae Mateos, E, 2011:28

Lepidocyrtus juliae from Greece
After Mateos E, 2011 Fig.1

            Syn.: Lepidocyrtus lignorum f. ? Ellis, WN, 1976:314
            Syn.: Lepidocyrtus lignorum f. triangulata Gruia, Poliakov & Broza, 2000


Species lignorum (Fabricius O, 1775) Gervais, 1844, B

            Syn.: Lepidocyrtus argentatus Bourlet, 1843
            Syn.: Lepidocyrtus albicans Reuter, 1895
            Syn.: Lepidocyrtus lignorum var. da Gama, MM, 1973:60


             Syn.: Podura Lignorum(sic) Fabricius, JC, 1796:138 ISS
Lepidocyrtus lignorum from the UK
After Lubbock J, 1873 Pl.26

             Syn.: Lepidocyrtus lignorus(sic) Tamura & Chiba, 1977 ISS
Lepidocyrtus lignorum from Sweden
2005 © Hall, K.


             Syn.: Lepidocyrtus lignorus(sic) Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:894 ISS
Lepidocyrtus lignorum from the UK
Descaled specimen
2007 © Cornwall, N.J.
Lepidocyrtus lignorum from the Netherlands
2007 © Hautus, M.A.
Lepidocyrtus lignorum from the UK
2007.12.15 © Kilford., B
Lepidocyrtus lignorum from the UK
2007.12.23 © Kilford., B
Lepidocyrtus lignorum from the UK
2008.03.25 © Robertson, A.
Lepidocyrtus lignorum from the UK
2008.03.25 © Robertson, A.
Lepidocyrtus lignorum from the UK
2008.04.13 © Kilford., B
Lepidocyrtus lignorum from the UK
2008.12.14 © Kilford., B
Lepidocyrtus lignorum from the UK
2009.02.12 © Jeffery, J.
Lepidocyrtus lignorum from Norway
2009.11.27 © Haraldseide, H.
Lepidocyrtus lignorum from the UK
Dorsal body scales
2010.01.31 © Robertson, A.
Lepidocyrtus lignorum from the UK
Distinctly round eyepatch
2010.01.31 © Robertson, A.


             Syn.: Lepidocyrtus ligonorum(sic) Mateos, E, 2011:25 ISS
Lepidocyrtus lignorum from Holland
2011.02.28 © van Duinen, J.
Lepidocyrtus lignorum from the UK
2012.02.19 © Barton, T.
Lepidocyrtus lignorum from the UK
Descaled
2012.08.25 © Murray, A.
Lepidocyrtus lignorum from the UK
2012.08.25 © Murray, A.
Lepidocyrtus lignorum from Holland
2012.10.28 © van Duinen, J.
"Lepidocyrtus lignorum"(sic) from India
After Santeshwari, Raghuraman M & Singh J, 2015 p.575
See note by Janssens F, 2016.11.02
Remark by Janssens, F. 2016.11.02: Lepidocyrtus lignorum is to be excluded for the specimen of Santeshwari et al., 2015:574 from India given distincly dark dorsal abdominal pigmentation is present.
Lepidocyrtus lignorum from Germany
Eyepatch roundish
2015.01.11 © Mattew, M.
Lepidocyrtus lignorum from Germany
Descaled revealing intrinsic yellow body colour
2015.01.12 © Mattew, M.
Lepidocyrtus lignorum from Holland
Eyepatch roundish
2015.02.13 © van Duinen, J.
Lepidocyrtus lignorum from Holland
2015.02.24 © Baas, A.H.
Lepidocyrtus lignorum from the UK
2015.12.28 © David W.
Lepidocyrtus lignorum from Belgium
2017.10.18 © Huskens, M.-L.
Lepidocyrtus lignorum from France
Ant.1+2 scaled
2017.11.04 © Picard, J.
Lepidocyrtus lignorum from Belgium
Interocular macrosetae absent
Group of 5 interantennal macrosetae present
2018.03.13 © Huskens, M.-L.
Lepidocyrtus lignorum from the UK
2018.12.25 © Barton, T.
Lepidocyrtus lignorum from Japan
2019.02.05 © Hara, Y.
Lepidocyrtus lignorum from Japan
2019.02.05 © Hara, Y.
Lepidocyrtus lignorum from Japan
Ant.1+2 iridiscent
2019.02.05 © Hara, Y.
Lepidocyrtus lignorum from Japan
Group of interantennal macrosetae
2019.02.05 © Hara, Y.
Lepidocyrtus lignorum from the UK
Cephalic interocular pattern
Descaled
2019.03.03 © Cann, A.J.



Species peisonis Traser, G et Christian, E, 1992:119

Lepidocyrtus peisonis from Hungary
After Winkler D & Mateos E, 2018 Fig.1
Lepidocyrtus peisonis from Hungary
Dorsal chaetotaxy
Abd.3 with lateral tuft of filliform chaetae
After Winkler D & Mateos E, 2018 Fig.7

Species ruber Schött, 1902

            Syn.: Lepidocyrtus lanuginosus Börner, nec Tullberg
            Syn.: Lepidocyrtus discolor Latzel, R, 1917:245
            Syn.: Lepidocyrtus discolor f. principalis Latzel, R, 1917:245
            Syn.: Lepidocyrtus discolor var. bicinctus Latzel, R, 1917:245
             Syn.: Lepidocyrtus discolor-bicinctus(sic) Handschin, E in Dahl, F, 1929:88,si ISS
            Syn.: Lepidocyrtus discolor var. palliatus Latzel, R, 1917:245
             Syn.: Lepidocyrtus discolor-palliatus(sic) Handschin, E in Dahl, F, 1929:88,si ISS
            Syn.: Lepidocyrtus instratus Handschin, 1924
            Syn.: Lepidocyrtus pulchellus Denis, 1927


Lepidocyrtus ruber from the UK
Empodium with characteristic apex
2005 © Hopkin, S.P.
Lepidocyrtus "ruber" from Scandinavica
Empodium with characteristic apex (*)
After Fjellberg A, 2007 Fig.83H
Lepidocyrtus ruber from Europe
2008.12.08 © Mateos, E.
Lepidocyrtus ruber from France
2013.03.05 © Delhem, R.
Lepidocyrtus ruber from France
Empodium with characteristic apex
2013.03.05 © Delhem, R.
Lepidocyrtus ruber from Holland
2013.10.14 © van Bezouw, R.
Lepidocyrtus ruber from the UK
2013.10.15 © Shaw, P.

Species tellecheae Arbea, JI et Jordana, R, 1990:28

             Syn.: Lepidocyrtus tellechae(sic) Simón-Benito, JC, 2007:323 ISS
             Syn.: Lepidocyrtus tellechae(sic) Mateos, E, 2008:48 ISS
Lepidocyrtus tellecheae from Europe
2008.12.06 © Mateos, E.



Species traseri Winkler, 2016

Species uzeli Rusek, 1985

Species vexillosus Loksa & Bogojevic, 1967

Species violaceus (Fourcroy, 1785) Parona, 1882, B

            Syn.: Lepidocyrtus metallicus Packard, AS, 1873:36
            Syn.: Lepidocyrtus assimilis Reuter, 1890
            Syn.: Lepidocyrtus cyaneus var. assimilis Reuter, 1892
            Syn.: Lepidocyrtus assimilis Reuter, 1892
            Syn.: Lepidocyrtus assimilis Gisin, H, 1965:520
            Syn.: Lepidocyrtus cyaneus var. assimilis Snider, RJ, 1967:11
            Syn.: Lepidocyrtus assimilis Christiansen, K et Bellinger, P, 1980:936
            Syn.: Lepidocyrtus assimilis Christiansen, K et Bellinger, P, 1998:1038
            Syn.: Lepidocyrtus cyaneus Lameere, 1895
             Syn.: Lepidocyrtus cyanus(sic) van den Bruel, WE in de Jonghe d'Ardoye, E, 1937:190,B ISS
             Syn.: Lepidocyrtus cyanus(sic) van den Bruel, WE in de Jonghe d'Ardoye, E, 1937:321,B ISS
            Syn.: Lepidocyrtus pusillus Jackson, AD, 1909:529
Lepidocyrtus pusillus from the USA
After Jackson, A.D., 1909 Pl.28 fig.1

            Syn.: Lepidocyrtus pusillus Maynard, EA, 1951:180
Lepidocyrtus pusillus from the USA
After Maynard EA, 1951 PlateI Fig.10
Lepidocyrtus pusillus from the USA
After Maynard EA, 1951 PlateXIX Fig.310

             Syn.: Podura pusillus(sic) Maynard, EA, 1951:180 ISS
            Syn.: Lepidocyrtus cyaneus ssp. cinereus Greenslade & Wise, 1986
            Syn.: Lepidosira terraereginae Greenslade & Wise, 1986
            Syn.: Lepidocyrtus sp. Greenslade, 1990
            Syn.: Lepidocyrtus sp. Greenslade & van Klinken, 1994
            Syn.: Lepidocyrtus cyaneus Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:22,B



            Syn.: Podura violacea Geoffroy, 1762, nr ICZN
Lepidocyrtus violaceus from the USA
After Maynard EA, 1951 PlateI Fig.11

             Syn.: Isotoma violaceus(sic) Maynard, EA, 1951:177 ISS
Lepidocyrtus violaceus from Europe
Cephalic dorsal chaetotaxy
After Gisin H, 1964 Fig.1 left


             Syn.: Lepidocyrtus (Lepidocyrtus) violaceous(sic) Snider, RJ, 1967:11 ISS
             Syn.: Lepidocyrtus violaceous(sic) Snider, RJ, 1967:209 ISS
Lepidocyrtus violaceus from the UK
Scales (s) on 2 most basal antennal segments
2005 © Hopkin, S.P.
Lepidocyrtus violaceus from Europe
2008.12.08 © Mateos, E.
Lepidocyrtus violaceus from Europe
2008.12.08 © Mateos, E.
Lepidocyrtus violaceus from Germany
2009.12.11 © Marquard, F.

             Syn.: Lepidocyrtus violacaeus(sic) Greenslade, P et Convey, P, 2012:411 ISS
             Syn.: Lepidocyrtus violaceous(sic) Santeshwari, X, Raghuraman, M et Singh, J, 2015:572 ISS
Lepidocyrtus violaceous(sic) from India
After Santeshwari, Raghuraman M & Singh J, 2015 p.574
Lepidocyrtus violaceus from Serbia
2017.11.21 © Sántha, K.
Lepidocyrtus (Lepidocyrtus) violaceus from Belgium
Grooming
2017.10.09 © Huskens, M.-L.
Lepidocyrtus (Lepidocyrtus) violaceus from Belgium
2018.10.21 © Huskens, M.-L.
Lepidocyrtus (Lepidocyrtus) violaceus? from Canada
2019.02.20 © Yungwirth, E.


Species-group curvicollis Mateos E & Petersen H, 2012:60
            Syn.: Lepidocyrtus curvicollis-paradoxus-subgroup Gisin, 1964



             Syn.: Lepidocyrtus curvocillis(sic)-group Mateos, E, 2012:81 ISS
Lepidocyrtus curvicollis-group
Mesothorax distinctly protruded
After Mateos E & al, 2018 Fig.1C



Species apicalis Mateos, E et Petersen, H, 2012:53

Lepidocyrtus apicalis from Sardinia
After Mateos E & Petersen H, 2012 Fig.1

Species curvicollis Bourlet, C, 1839:392, t.t., B

            Syn.: Podura plumbea Pritchard, A, 1832:149
Podura plumbea from the UK
Scales with waved and curved lines. 350x
After Pritchard A, 1832 Pl.12 Fig.9,10
Podura plumbea from the UK
Scale with matrix pattern. 350x
After Pritchard A, 1832 Pl.12 Fig.13
Podura plumbea Pritchard, 1832
Test slide of dry mounted scales by Pritchard
2010.12.29 © Lynk, H.
Podura plumbea Pritchard, 1832
Scale with waved pattern
2010.12.29 © Lynk, H.
Podura plumbea Pritchard, 1832
Scale with matrix pattern
2010.12.29 © Lynk, H.
Podura plumbea from the UK
Scales. 350x
After Pritchard A, 1832 Pl.12 Fig.5,8

Podura scales, 1840
Lepidocyrtus curvicollis from the UK
2011.08.dd © Walker, D.


            Syn.: Podura plumbea Quekett, J, 1848:435
Podura plumbea from the UK
Scales with wedge-shaped dots. 500x. 1250x
After Quekett J, 1848 Fig.4-5


            Syn.: Podura plumbea McIntire, SJ, 1867:54
Lepidocyrtus curvicollis? from the UK
After McIntire SJ, 1867 Fig.39
After McIntire SJ, 1868
Lepidocyrtus curvicollis? from the UK
Frontal aspect
After McIntire SJ, 1867 Fig.40
After McIntire SJ, 1868
Lepidocyrtus curvicollis? from the UK
Scale
After McIntire SJ, 1867 Fig.42
After McIntire SJ, 1868
Lepidocyrtus curvicollis? from the UK
Test Podura scale
After McIntire SJ, 1867 Fig.48
After McIntire SJ, 1868


            Syn.: Macrotoma nigra McIntire, SJ, 1867:55
            Syn.: Cyphodeirus(sic) Æneus(sic) Nicolet, H, 1842:66 IOS
             Syn.: Cyphodeirus(sic) ceneus(sic) Bourlet, C, 1845:67 ISS
            Syn.: Cyphodeirus(sic) agilis Nicolet, H, 1842:66
            Syn.: Cyphodeirus(sic) capucinus Nicolet, H, 1842:382
            Syn.: Cyphodeirus(sic) gibbulus Nicolet, H, 1842:64
            Syn.: Cyphodeirus(sic) lignorum Nicolet, H, 1842:65
            Syn.: Cyphodeirus(sic) parvulus Nicolet, H, 1842:67
            Syn.: Cyphodeirus(sic) pusillus Nicolet, H, 1842:65
            Syn.: Lepidocyrtus neglectus Moniez, 1890
            Syn.: Lepidocyrtus Börneri(sic) Willem, V, 1902:281,B IOS
             Syn.: Lepidocyrtus börneri(sic) Bonet, F, 1931:383,B ISS
            Syn.: Lepidocyrtus luteus Latzel, R, 1917:244 nec Jackson, 1909
            Syn.: Lepidocyrtus curvicollis ssp. fuscicans Latzel, 1922
            Syn.: Lepidocyrtus curvicollis ssp. oribates Latzel, 1922
            Syn.: Lepidocyrtus curvicollis ssp. semiviolaceus Latzel, 1922
            Syn.: Lepidocyrtus latzeli Salmon, JT, 1964:522



Lepidocyrtus curvicollis from France
After Bourlet C, 1839 Fig.6,7
Cyphoderus curvicollis from Switzerland
After Nicolet H, 1847 Pl.6 Fig.13
Lepidocyrtus curvicollis from Czechia
Descaled specimen
2007.07.05 © Krásenský, P.

Cyphoderus curvicollis from the UK
After Lubbock, J., 1873 Pl.25.
Lepidocyrtus curvicollis from the UK
After Beck J in Lubbock J, 1873 Pl.73
Lepidocyrtus curvicollis from the UK
After Beck J in Lubbock J, 1873 Pl.74
Lepidocyrtus curvicollis from Europe
Cephalic dorsal chaetotaxy
After Gisin H, 1964 Fig.1 left
Lepidocyrtus curvicollis from the UK
TEM of microspines on outer scale surface
After Baker RM & McCrae JM, 1966 Fig.3,4
Lepidocyrtus curvicollis from the UK
Imagemodel of microspine in light microscopy
Baker RM & McCrae JM, 1966 Fig.3,4
2010.12.28 © Modified by Janssens F

Lepidocyrtus curvicollis from Malta
After Stach J, 1967 Pl.XXXIII Fig.4

             Syn.: Lepidocyrtus curvicolis(sic) Thibaud, J-M et Massoud, Z, 1977:6,B ISS
Cyphoderus curvicollis from the UK
2004 © Hopkin, S.P.
Lepidocyrtus curvicollis from the UK
2005 © Hopkin, S.P.


             Syn.: Lepidocyrtus curvicolis(sic) Fiera, C, 2007:28 ISS

Lepidocyrtus curvicollis from France
2007.03.12 © Lebeaux, P.
Lepidocyrtus curvicollis from France
2007.03.12 © Lebeaux, P.
Lepidocyrtus curvicollis from France
2007.11.27 © Lebeaux, P.
Lepidocyrtus curvicollis from the UK
2009.02.15 © Goldenorfe, P.
Lepidocyrtus curvicollis from Germany
2009.02.16 © Schulz, H.-J.
Lepidocyrtus curvicollis from the UK
2009.06.12 © Robertson, A.
Lepidocyrtus curvicollis from the UK
2009.10.07 © Robertson, A.
Lepidocyrtus curvicollis from the UK
Mesothoracic anterior angle about 90 degrees
2011.07.05 © Shaw, P.
Lepidocyrtus curvicollis from France
2011.09.10 © Dehalleux, A.
Lepidocyrtus curvicollis from the UK
2011.12.02 © Nurcombe, E.
Lepidocyrtus curvicollis from Ireland
2012.02.07 © Cluskey, D.
Lepidocyrtus curvicollis from Canada
Antenna
2012.02.19 © Hochhalter, R.
Lepidocyrtus curvicollis from Canada
Lateral mesothoracic scales
2012.02.19 © Hochhalter, R.
Lepidocyrtus curvicollis from the UK
2012.11.21 © Phillips, E.
Lepidocyrtus curvicollis from the UK
2013.02.06 © Murray, A.
Lepidocyrtus curvicollis juv. from the UK
2013.04.16 © Murray, A.
Lepidocyrtus curvicollis from France
2013.06.15 © Sandra.
Lepidocyrtus curvicollis from Russia
2013.10.05 © Haritonoff.
Lepidocyrtus curvicollis from India
After Santeshwari, Raghuraman M & Singh J, 2015 p.575
Lepidocyrtus curvicollis from the UK
Dorsally strongly curved mesothorax
2015.01.31 © Murray, A.
Lepidocyrtus curvicollis from the UK
Iridiscent appically rounded body scales
2015.01.31 © Murray, A.
Lepidocyrtus curvicollis from Holland
2015.09.16 © van Duinen, J.
Lepidocyrtus curvicollis from the USA
2015.11.17 © Leitkam, S.
Lepidocyrtus curvicollis from France
2016.02.21 © Garcelon, P.
Lepidocyrtus curvicollis from Belgium
2016.09.17 © Huskens, M.-L.

             Syn.: Lepidocyrtus curvicolis(sic) Yadav, RS, 2017:1725 ISS
             Syn.: Lepidocyrtus curvicolis(sic) Yadav, RS, 2017:1726 ISS
             Syn.: Lepidocyrtus curvicolis(sic) Yadav, RS, 2017:11057 ISS
             Syn.: Lepidocyrtus curvicolis(sic) Yadav, RS, 2017:11058 ISS
Lepidocyrtus curvicolis(sic)from India
After Yadav RS, 2017 Img.5

             Syn.: Lepidocyrtus curvicolis(sic) Yadav, RS, 2017:1339 ISS
Lepidocyrtus curvicollis from Holland
2017.02.28 © van Leeuwen, J.
Lepidocyrtus curvicollis from Holland
2017.09.28 © van Duinen, J.
Lepidocyrtus curvicollis juv. from Holland
2017.09.28 © van Duinen, J.

             Syn.: Lepidocyrtus curvicolis(sic) Yadav, RS, Prasad, S et Kumar, D, 2018:17 ISS
             Syn.: Lepidocyrtus curvicolis(sic) Yadav, RS, Prasad, S et Kumar, D, 2018:18 ISS
             Syn.: Lepidocyrtus curvicolis(sic) Yadav, RS, Prasad, S et Kumar, D, 2018:19 ISS
             Syn.: Lepidocyrtus curvicolis(sic) Yadav, RS, Prasad, S et Kumar, D, 2018:20 ISS
Lepidocyrtus curvicollis from France
Right angled mesothorax
2018.11.22 © Dejoux, P.

  Subspecies cyaneipes Schille, 1908





Species flexicollis Gisin, H, 1965:523

Lepidocyrtus flexicollis from Spain
After Mateos E, 2008 Fig.1
Lepidocyrtus flexicollis from Europe
2008.12.08 © Mateos, E.

             Syn.: Lepidocyrtus fexicollis(sic) Mateos, E, 2008:41 ISS
Lepidocyrtus flexicollis from La Palma
2012.12.04 © Barton, T.
Lepidocyrtus flexicollis from La Palma
2012.12.04 © Barton, T.
Lepidocyrtus flexicollis from La Palma
2012.12.06 © Barton, T.
Lepidocyrtus flexicollis from Spain
After Arbea JI, 2013 Fig.14



Species mariani Traser & Dányi, 2008

Lepidocyrtus mariani? from Hungary
2008.09.26 © Keresztes, G.
Lepidocyrtus mariani? from Hungary
2008.09.26 © Keresztes, G.
Lepidocyrtus mariani? from Hungary
2008.09.28 © Keresztes, G.
Lepidocyrtus mariani from Hungary
After Winkler D & Mateos E, 2018 Fig.24

Species montseniensis Mateos, 1985

             Syn.: Lepidocyrtus monseniensis(sic) Mateos, E, 2008:35 ISS
Lepidocyrtus montseniensis from Europe
2008.12.06 © Mateos, E.



Species nigrescens Szeptycki, 1967

             Syn.: Lepidocyrtus nigresces(sic) Mateos, E, 2008:36 ISS
Lepidocyrtus nigrescens from Europe
2008.12.08 © Mateos, E.
Lepidocyrtus nr nigrescens from the UK
2016.01.07 © Evans, K.
Lepidocyrtus nr nigrescens from the UK
2016.01.07 © Evans, K.
Lepidocyrtus nr nigrescens from the UK
2016.04.16 © Shepherd, M.
Lepidocyrtus nr nigrescens from the UK
2016.04.16 © Shepherd, M.



Species paradoxus Uzel, 1890, B

            Syn.: Paidium cucullatum Koch, 1840
            Syn.: Cyphodeirus(sic) capucinus Nicolet, H, 1842:64
            Syn.: Lepidocyrtus cephalopurpureus Harvey, 1894
            Syn.: Lepidocyrtus insignis Reuter, 1895
            Syn.: Lepidocyrtus alleganyensis Maynard, EA, 1951:180 =?
Lepidocyrtus alleganyensis from the USA
After Maynard EA, 1951 PlateXIX Fig.312
Lepidocyrtus alleganyensis from the USA
After Maynard EA, 1951 PlateXIX Fig.313

            Syn.: Lepidocyrtus alleganyensis Christiansen, K et Bellinger, P, 1980:956=?
            Syn.: Lepidocyrtus alleganyensis Christiansen, K et Bellinger, P, 1998:1038=?
            Syn.: Lepidocyrtus christianseni Goto, 1953



Lepidocyrtus paradoxus from Europe
Cephalic dorsal chaetotaxy
After Gisin H, 1964 Fig.1 midright
Lepidocyrtus paradoxus from Malta
After Stach J, 1967 Pl.XXXIII Fig.5

             Syn.: Lepidocyrtus paradaoxus(sic) Jacquemart, S, 1976:139 ISS
             Syn.: Lepidocyrtus paradoxas(sic) Skidmore, RE, 1995:59 ISS
Lepidocyrtus paradoxus from the Netherlands
After Berg, M. & Heijerman, T., 2002 Fig.1.
Lepidocyrtus paradoxus from the Netherlands
After Berg, M. & Heijerman, T., 2002 Fig.2.
Lepidocyrtus paradoxus from Germany
2006 © Kohl, F.
Lepidocyrtus paradoxus from the USA
2006 © DiTerlizzi, T.
Det. Soto-Adames, F., 2007.10.22
Lepidocyrtus paradoxus from the USA
2006 © McClarin, J.
Lepidocyrtus paradoxus from Hungary
2006.08.26 © Keresztes, G.
Lepidocyrtus paradoxus from the USA
2007.03.14 © Murray, T.
Det. Soto-Adames, F., 2007.10.22
Lepidocyrtus paradoxus from the USA
2007.07.22 © Cresswell, S.
Det. Soto-Adames, F., 2007.10.22
Lepidocyrtus paradoxus from the USA
2007.10.19 © McClarin, J.
Det. Soto-Adames, F., 2007.10.22
Lepidocyrtus paradoxus from Czechia
2008.05.21 © Deml, M.
Lepidocyrtus paradoxus from Germany
2008.08.15 © Petzoldt, S.
Lepidocyrtus paradoxus from Germany
2008.08.15 © Petzoldt, S.
Lepidocyrtus paradoxus from Europe
2008.12.08 © Mateos, E.
Lepidocyrtus paradoxus from the USA
2009.07.22 © Loser, I.
Lepidocyrtus paradoxus from Ukraine
2009.07.22 © Loboda., B
Lepidocyrtus paradoxus from the USA
2010.08.18 © Gerber, G.K.
Lepidocyrtus paradoxus from Hungary
2011.08.13 © Rahmé, N.
Lepidocyrtus paradoxus from France
2011.09.10 © Dehalleux, A.
Lepidocyrtus paradoxus from the USA
2011.10.09 © Bradford, A.
Lepidocyrtus paradoxus (bar=1µm)
Hexagonal epicuticular ultrastructure
After Nickerl J & al, 2012 Fig.3C
Lepidocyrtus paradoxus from the USA
2012.10.13 © McVey, S.
Lepidocyrtus paradoxus from the UK
2013.08.02 © Gilligan, N.
Lepidocyrtus paradoxus from the UK
2013.08.02 © Gilligan, N.
Lepidocyrtus paradoxus from France
Feeding on dead Aphidae
2013.08.06 © Quintin, C.
"Lepidocyrtus paradoxus"(sic) from India
After Santeshwari, Raghuraman M & Singh J, 2015 p.574
See note by Janssens F, 2016.11.02
Remark by Janssens, F. 2016.11.02: Lepidocyrtus paradoxus is to be excluded for the specimen of Santeshwari et al., 2015:574 from India given the intrinsic body colour is not black.
Lepidocyrtus paradoxus from the USA
Intrinsic body colour blackish
2015.07.17 © Blocky.
Lepidocyrtus paradoxus from the USA
2015.11.20 © Schrayer, G.
Lepidocyrtus paradoxus from Canada
2016.02.21 © Murray, A.
Lepidocyrtus paradoxus from the USA
Intrinsic body colour blackish
2016.07.03 © Hunt, A.
Lepidocyrtus paradoxus from Belgium
2017.08.28 © Huskens, M.-L.
Lepidocyrtus paradoxus from Belgium
2017.08.29 © Huskens, M.-L.
Lepidocyrtus paradoxus from the USA
2017.09.23 © Dracunculus.
Lepidocyrtus paradoxus from the USA
2017.09.23 © Dracunculus.

Subgenus Setogaster Salmon, 1951
  Syn.: ... Trichogaster Handschin, 1932, nec Schneider, 1801
  Syn.: ... Pseudocyrtus Salmon, 1956,is





Species amazonicus Cipola NG et Bellini BC in Cipola, NG, de Morais, JW et Bellini, BC, 2018:5

Lepidocyrtus (Setogaster) amazonicus from Brazil
After Cipola NG, de Morais JW & Bellini BC, 2018 Fig.3A

Species bispinosus (Handschin, 1932) Yayuk, RS (sic), 1989:11, t.t.

Species caprilesi Wray, DL, 1953:145

Species coerulea (Salmon, 1951) Yayuk, RS (sic), 1989:11

            Syn.: Lepidocyrtus (Setogaster) sp.3 Yayuk, RS(sic), 1989:11


             Syn.: Trichogaster caerulea(sic) Yayuk, RS(sic), 1989:11 ISS

Species fasciata (Salmon, 1951) Yayuk, RS (sic), 1989:11

            Syn.: Lepidocyrtus (Setogaster) sp.2 Yayuk, RS(sic), 1989:11


Species frantzeni (Winter C, 1963:503) n.n.

Remark by Winter, C. 1963:395: 37 von mir neu aufgestellte Arten bzw. Formen sind in der vorliegende Arbeit mit n.sp. bzw. n.f. (ohne Autorennamen) bezeichnet und müssen bis zu ihrer Beschreibung (voraussichtlich im kommenden Jahr) als Nomina nuda betrachtet werden.
Species indicus Handschin, E, 1929:242

Lepidocyrtus indicus from South-India
After Handschin E, 1929 Fig.20

Species kulluensis Baquero E et Jordana R in Baquero, E, Mandal, G et Jordana, R, 2015:20

Cinctocyrtus kulluensis from India
After Baquero E & al, 2015 Fig.12

Species manipuri Salmon, 1969

Species merapicus (Yoshii, R & Suhardjono, YR, 1989:57) Yoshii, R & Suhardjno, YR, 1992:32

Ascocyrtus (Merapicyrtus) merapicus from Indonesia
After Yoshii R & Suhardjono YR, 1992 Fig.4A

Species multisensillatus Cipola NG et Bellini BC in Cipola, NG, de Morais, JW et Bellini, BC, 2018:12

Lepidocyrtus (Setogaster) multisensillatus from Brazil
After Cipola NG, de Morais JW & Bellini BC, 2018 Fig.3B

Species nigrosetosus Folsom, JW, 1927:9

            Syn.: Lepidocyrtus geayides Mari Mutt, 1986 =?
            Syn.: Lepidocyrtus leleupi Jacquemart, S, 1976:145


Lepidocyrtus nigrosetosus from the Galapagos
After Katz AD & al, 2016 Fig.7A

Species pallida (Salmon, 1951) Yayuk, RS (sic), 1989:11

            Syn.: Lepidocyrtus (Setogaster) sp.1 Yayuk, RS(sic), 1989:11


Species projectus Salmon, 1956

Species sotoi Bellini BC et Godeiro NN in Bellini, BC, Cipola, NG et Godeiro, NN, 2015:228

Lepidocyrtus sotoi from Brazil
After Bellini BC & Godeiro NN in Bellini BC & al, 2015 Fig.1

Subgenus Allocyrtus Yoshii, R et Suhardjono, YR, 1989:44
Species lepidornatus (Handschin, 1930) Yoshii, R, 1982:22, t.t.

Lepidocyrtus (Acrocyrtus) lepidornatus from Malaysia
After Yoshii R, 1982 Fig.12A
Lepidocyrtus (Allocyrtus) lepidornatus from the Himalayas
After Baquero E & al, 2014 Fig.11


Subgenus Cinctocyrtus Yoshii, R et Suhardjono, YR, 1989:55
Species cinctus Schäffer, C, 1898:421, t.t.

            Syn.: Ascocyrtus tongensis Yosii, R, 1964:12


Lepidocyrtus (Ascocyrtus) cinctus from Malaysia
After Yoshii R, 1982 Fig.15A
Ascocyrtus (Cinctocyrtus) cinctus from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.18A

Species filamentosus Yoshii, R, 1982:33

Lepidocyrtus (Ascocyrtus) filamentosus from Malaysia
After Yoshii R, 1982 Fig.19A
Ascocyrtus (Cinctocyrtus) filamentosus from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.18D

Species hankowi Denis, 1929

Ascocyrtus (Cinctocyrtus) hankowi from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.18C

Species koreanus Kim, J-T et Lee, B-H, 1995:450

Species medius Schäffer, C, 1898:420

Lepidocyrtus medius from India
After Mandal GP & Hazra AK, 2009 Fig.27a

Species minimus Park K-H et Rojanavongse V in Kim, JT, Park, K-H, Rojanavongse, V et Lee, B-H, 1999:220

Species reductus Park K-H et Rojanavongse V in Kim, JT, Park, K-H, Rojanavongse, V et Lee, B-H, 1999:220

Species sandakanicus Yoshii, R, 1982:32

            Syn.: Trichogaster fasciata Salmon, 1951


Lepidocyrtus (Ascocyrtus) sandakanicus from Malaysia
After Yoshii R, 1982 Fig.18A
Ascocyrtus (Cinctocyrtus) sandakanicus from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.18B
Remark by Jordana, R. 2008.12.10: in litt.: Species combination to be revised. This species clearly has 6 + 6 single eyes as in Pseudosinella.

             Syn.: Cinctocyrtus sandakanikus(sic) Baquero, E, Mandal, G et Jordana, R, 2015:25 ISS


Species sepilokensis Yoshii, R, 1982:31

Lepidocyrtus (Ascocyrtus) sepilokensis from Malaysia
After Yoshii R, 1982 Fig.17A
Remark by Jordana, R. 2008.12.10: in litt.: Species combination to be revised. This species clearly has 6 + 6 single eyes as in Pseudosinella.


Species simplex Park K-H et Rojanavongse V in Kim, JT, Park, K-H, Rojanavongse, V et Lee, B-H, 1999:219

Species sublobatus Yosii, 1961


Subgenus Ascocyrtus Yosii, 1963   Syn.: Lepidocyrtus Discocyrtus Yosii, R, 1959:34 nec Holmberg, 1876
  Syn.: Ascocyrtus Dahlcyrtus Yoshii, R et Suhardjono, YR, 1989:55


Species aseanus Yoshii, R, 1982:26

            Syn.: Lepidocyrtus (Discocyrtus) suborientalis Yosii, 1959, nec Denis, 1948


Lepidocyrtus (Ascocyrtus) aseanus from Malaysia
After Yoshii R, 1982 Fig.14A

Species dahli Schäffer, C, 1898:419

            Syn.: Lepidocyrtus dahlii(sic) Schäffer, C, 1898:419 IOS
             Syn.: Lepidocyrtus dahlii(sic) Carpenter, GH, 1916:40 ISS
             Syn.: Lepidocyrtus (truncati-group) Dahlii(sic) Denis, J-R, 1924:259 ISS
             Syn.: Discocyrtus dahlii(sic) Yosii, 1960 ISS
             Syn.: Lepidocyrtus (Ascocyrtus) dahlii(sic) Yoshii, R, 1982:24 ISS
             Syn.: Discocyrtus dahlii(sic) Deharveng, L et Najt, J in Tillier, S, 1988:18 ISS
             Syn.: Lepidocyrtus (Ascocyrtus) dahlii(sic) Yayuk, RS(sic), 1989:11 ISS
             Syn.: Ascocyrtus (Dahlcyrtus) dahlii(sic) Yoshii, R et Suhardjono, YR, 1989:44 ISS
             Syn.: Ascocyrtus (Dahlcyrtus) dahlii(sic) Yoshii, R et Suhardjno, YR, 1992:31 ISS
             Syn.: Lepidocyrtus (Dahlcyrtus) dahlii(sic) Wang, F, Chen, J-X et Christiansen, K, 2003:837 ISS
             Syn.: Ascocyrtus dahlii(sic) Zhang, F, Bedos, A et Deharveng, L, 2016:19 ISS



Species decui Gruia, M, 1984:23

Lepidocyrtus (Ascocyrtus) decui from Cuba
After Gruia M, 1984 Pl.III Fig.1

Species gisini Izarra, 1972

Species kuakea Christiansen, K et Bellinger, P, 1992:287

Species magnificus Carpenter, 1924

Lepidocyrtus magnificus from India
After Mandal GP & Hazra AK, 2009 Fig.26a
             Syn.: Lepidocyrtus magnijicus(sic) Yadav, RS, Prasad, S et Kumar, D, 2018:17 ISS
             Syn.: Lepidocyrtus magnijicus(sic) Yadav, RS, Prasad, S et Kumar, D, 2018:18 ISS
             Syn.: Lepidocyrtus magnijicus(sic) Yadav, RS, Prasad, S et Kumar, D, 2018:19 ISS


Species scaber Ritter, 1911

Species suborientalis (Yosii, R, 1959:35) Ellis, WN & Bellinger, PF, 1973:18, t.t.

Species tongensis Yosii, R, 1964:12


Subgenus Fractocyrtus Cipola NG et Bellini BC in Cipola, NG, de Morais, JW et Bellini, BC, 2018:22
Species americanus Cipola NG et Bellini BC in Cipola, NG, de Morais, JW et Bellini, BC, 2018:23, t.t.

Lepidocyrtus (Fractocyrtus) americanus from Brazil
After Cipola NG, de Morais JW & Bellini BC, 2018 Fig.14A

Species mateosi Cipola NG et Bellini BC in Cipola, NG, de Morais, JW et Bellini, BC, 2018:34

Lepidocyrtus (Fractocyrtus) mateosi from Brazil
After Cipola NG, de Morais JW & Bellini BC, 2018 Fig.14B


Genus Acrocyrtus Yosii, R, 1959:28

  Syn.: Acrocyrtus Carocyrtus Yoshii, R et Suhardjono, YR, 1989:48
  Syn.: Acrocyrtus Onerocyrtus Yoshii, R et Suhardjono, YR, 1989:44


Species aequidentatus (Denis, 1948) Baquero, E, Mandal, G & Jordana, R, 2015:21

Species baii (Nguyen, TT, 2005:8) Xu, G-L, Pan, Z-X & Zhang, F, 2013:3

Species carosus (Yosii, R, 1959:31) Yoshii, R & Suhardjono, YR, 1989:46

Lepidocyrtus (Acrocyrtus) carosus
from Singapore
After Yosii R, 1959 Fig.17A

Species cheni Pan, Z-X, Chatterjee, T et Zhang, F, 2011:231

Acrocyrtus cheni from India
After Pan ZX, Chatterjee T & Zhang F, 2011 Fig.1,2

Species cryptocephalus (Handschin, E, 1929:240) Pan, Z-X, Chatterjee, T & Zhang, F, 2011:231

Lepidocyrtus cryptocephalus from South-India
After Handschin E, 1929 Fig.14

Species ductieni (Nguyen, TT, 2005:8) Baquero, E, Mandal, G & Jordana, R, 2015:21

Species eurylabialis Yoshii, R et Suhardjono, YR, 1989:53

Species finis Xu, G-L, Pan, Z-X et Zhang, F, 2013:9

Acrocyrtus finis from China
After Xu G-L, Pan Z-X & Zhang F, 2013 Fig.28

Species gentingensis (Yoshii, R, 1982:20) Yoshii, R & Suhardjono, YR, 1989:45

Lepidocyrtus (Acrocyrtus) carosus gentingensis from Malaysia
After Yoshii R, 1982 Fig.11A

Species heterolepis (Yosii, R, 1959:33) Yoshii, R & Suhardjono, YR, 1989:45

             Syn.: Lepidocyrtus (Acrocyrtus) heterolepsis(sic) Wang, F, Chen, J-X et Christiansen, K, 2003:825 ISS
             Syn.: Lepidocyrtus heterolepsis(sic) Yadav, RS, 2017:1725 ISS
             Syn.: Lepidocyrtus heterolepsis(sic) Yadav, RS, 2017:1726 ISS

Species himachalensis (Baquero E & Jordana R in Baquero, E, Mandal, G & Jordana, R, 2015:19) Baquero, E, Mandal, G & Jordana, R, 2015:20

Acrocyrtus himachalensis from India
After Baquero E & al, 2015 Fig.10-11

Species javanicus (Oudemans, JT in Weber, M, 1890:85) Yoshii, R & Suhardjono, YR, 1989:45

Species liewthatchimi (Yoshii, R, 1982:12) Yoshii, R & Suhardjono, YR, 1989:45

Lepidocyrtus (Acrocyrtus) liewthatchimi from Malaysia
After Yoshii R, 1982 Fig.6A

Species lobatus (Denis, 1948) Baquero, E, Mandal, G & Jordana, R, 2015:21

Species malayanus (Yosii, R, 1959:28) Yoshii, R & Suhardjono, YR, 1989:51

Lepidocyrtus (Acrocyrtus) malayanus
from Singapore
After Yosii R, 1959 Fig.15A
Lepidocyrtus (Acrocyrtus) malayanus from India
After Mandal GP & Hazra AK, 2009 Fig.28a

  Subspecies sabahnus Yoshii, R, 1982:17

Lepidocyrtus (Acrocyrtus) malayanus sabahnus from Malaysia
After Yoshii R, 1982 Fig.9A
Lepidocyrtus (Acrocyrtus) malayanus sabahnus from Malaysia
After Yoshii R, 1982 Fig.9B
Lepidocyrtus (Acrocyrtus) malayanus sabahnus from Malaysia
After Yoshii R, 1982 Fig.9C

Species mastani (Yoshii, R, 1982:14) Yoshii, R & Suhardjono, YR, 1989:45

Lepidocyrtus (Acrocyrtus) mastani from Malaysia
After Yoshii R, 1982 Fig.7A

Species moluccensis Yoshii, R et Suhardjno, YR, 1992:27

Species onerosus (Yoshii, R, 1982:15) Gruia, M, 1984:25, t.t.

Lepidocyrtus (Acrocyrtus) onerosus from Malaysia
After Yoshii R, 1982 Fig.8A

Species parvidentatus (Schäffer, C, 1898:417) Baquero, E, Mandal, G & Jordana, R, 2015:19

Species ralumensis (Schäffer, C, 1898:418) Yosii, 1960

            Syn.: Acrocyrtus (Carocyrtus) ochraceus Yoshii, R et Suhardjono, YR, 1989:50
             Syn.: Lepidocyrtus (Carocyrtus) ocharceus(sic) Wang, F, Chen, J-X et Christiansen, K, 2003:837 ISS


Acrocyrtus (Carocyrtus) ralumensis from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.17A

Species segamanus (Yoshii, R, 1982:8) Yoshii, R & Suhardjono, YR, 1989:45

Lepidocyrtus (Acrocyrtus) segamanus from Malaysia
After Yoshii R, 1982 Fig.3A,E
Remark by Jordana, R. 2008.12.10: in litt.: Species combination to be revised. This species clearly has 6 + 6 single eyes as in Pseudosinella.


Species solomonensis (Yosii, 1960) Yoshii, R & Suhardjno, YR, 1992:29

Species spinodensus (Nguyen, TT, 2005:8) Baquero, E, Mandal, G & Jordana, R, 2015:21

Species timoricus (Yoshii, R & Suhardjono, YR, 1992:84) Baquero, E, Mandal, G & Jordana, R, 2015:24

Species transiens (Yoshii, R, 1982:11) Yoshii, R & Suhardjono, YR, 1989:45

Species vietnamensis Nguyen, TT, 2005:8

Species yosiii (Salmon, JT, 1964:529) Baquero, E, Mandal, G & Jordana, R, 2015:21

            Syn.: Lepidocyrtus yosii(sic) Salmon, JT, 1964:529 IOS
            Syn.: Acrocyrtus bipunctatus Yosii, 1961, nec Packard, 1873


             Syn.: Lepidocyrtus (Acrocyrtus) yosii(sic) Wang, F, Chen, J-X et Christiansen, K, 2003:837 ISS



Species zhujiensis Xu, G-L, Pan, Z-X et Zhang, F, 2013:3

Acrocyrtus zhujiensis from China
After Xu G-L, Pan Z-X & Zhang F, 2013 Fig.1


Genus Acanthurella Börner, C, 1906:174

Species amethystina Handschin, 1925

Species betlaensis Mandal, GP, Suman, KK et Bhattacharya, KK, 2016:41

Acanthurella betlaensis from India
After Mandal GP, Suman KK & Bhattacharya KK, 2016 p.43 Img.1

Species braueri (Börner, C, 1906:176) Carpenter, GH, 1916:42, t.t.

             Syn.: Acanthurella Braueri(sic) Carpenter, GH, 1916:42 ISS

Species brunnea Handschin, 1925

Species glauca Schött, 1925

Species halei Womersley, H, 1937:156

Species hamata (Salmon, 1951) Zhang, F, Deharveng, L, Greenslade, P & Chen, J-X, 2009:496

Species javana (Börner, C, 1906:176) Handschin, 1925

             Syn.: Acanthurella javanus(sic) Zhang, F, Bedos, A et Deharveng, L, 2016:19 ISS

Species zonata Schött, 1925


Genus Rhynchocyrtus de Mendonça, MC et Fernandes, LH, 2007:46

Species klausi de Mendonça, MC et Fernandes, LH, 2007:46, t.t.

Rhynchocyrtus klausi from Brazil
After de Mendonça M.C. & Fernandes L.H., 2007 Fig.22.


Genus Rambutsinella Deharveng, L et Bedos, A, 1996:280

Rambutsinella sp.
2006 © Boldsystems.org

Species honchongensis Deharveng, L et Bedos, A, 1996:281, t.t.

Species grinnellia (Wang, F, Chen, J-X et Christiansen, K, 2004:373)

             Syn.: Pseudosinella grinnelli(sic) Wang, F, Chen, J-X et Christiansen, K, 2004:370 ISS

Species scopae Deharveng, L et Bedos, A, 1996:283


Genus Pseudosinella Schäffer, 1897 key

  Syn.: ... Tullbergia Lie-Pettersen, 1897, nec Lubbock, 1876
  Syn.: ... Troglosinella Delamare Deboutteville, 1949


  Syn.: Collembola Pseudosineila(sic) da Gama, MM, 1988:80 ISS
  Syn.: Entomobryidae Psudosinella(sic) Palacios-Vargas, JG et Castaño-Meneses, G in Basset, Y, Novotny, V, Miller, SE et Kitching, RL, 2003:164 ISS
Pseudosinella sp. from Portugal
2006.03.07 © Rasteiro, F.
Pseudosinella sp. from the USA
2007 © Montgomery, G.
Pseudosinella sp. from the UK
2008.04.03 © Spears, G.
Pseudosinella sp. from the UK
2009.03.29 © Barton, T.
Pseudosinella sp. from Croatia
2010.11.19 © Lukic, M.
Pseudosinella sp. from Holland
2011.07.27 © van Duinen, J.
Pseudosinella sp. from the USA
2011.10.02 © Thomas, D.J.
Pseudosinella sp. from the USA
Typical antennal segmentation
2011.12.11 © Kouri, J.
Pseudosinella sp. from Portugal
2012.10.11 © Murray, A.

  Syn.: Entomobryidae Peudosinella(sic) Schulz, H-J, 2015:17 ISS


Species sp.1 Paclt, J, 1961:234

Species sp.1a da Gama, MM, 1973:48

Species sp.2 Bellinger, PF et Christiansen, KA, 1974:39

Species sp.3 Palacios-Vargas, JG, 1981:90

Species sp.4 Palacios-Vargas, JG, 1981:90

Species sp.5 Palacios-Vargas, JG, 1981:90

Species sp.6 Christiansen, K, 1982:4

Species sp.7 Christiansen, 1982

Species sp.8 Maes, J-M et Palacios-Vargas, JG, 1988:5

Species sp.9 Mateos, E, 1988:101

Species sp.10 Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:14

Species sp.11 Zeppelini Filho, D et Castaño Meneses, G, 1995:10

Species sp.12 Christiansen, K et Bellinger, P, 1996:38

Species sp.13 Christiansen, K et Bellinger, P, 1996:38

Species sp.14 Christiansen, K et Bellinger, P, 1996:38

Species sp.15 Christiansen, K et Bellinger, P, 1996:38

Species sp.16 Christiansen, K et Bellinger, P, 1996:38

Species sp.17 Christiansen, K et Bellinger, P, 1996:38

Species sp.18 Palacios-Vargas, JG et Gamboa-Vargas, JA, 1997:90

Species sp.19 Palacios-Vargas, JG et Gamboa-Vargas, JA, 1997:90

Species sp.20 Loranger, G, Ponge, J-F, Blanchart, E et Lavelle, P, 1998:24

Species sp.21 Arbea, JI, 2005:43

Species sp.22 Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:895

Species sp.23 Elliott, WR, 2007:143

Species sp.24 Zhang, F et Deharveng, L, 2015:490

Species sp.25 Lukic, M, 2019:309

Pseudosinella sp. from Croatia
2010.11.19 © Lukic, M.
Pseudosinella sp. undescr. from Croatia
After Lukic M, 2019 Fig.2B

Species absoloni Rusek, 1967

Species aelleni da Gama, MM, 1973:52

             Syn.: Pseudosinella ælleni(sic) da Gama, MM, 1973:52 ISS
             Syn.: Pseudosinella ælleni(sic) da Gama, MM, 1973:53 ISS

Species aeolica Dallai, R, 1973:530

Species aera Christiansen & Bellinger, 1980

Species aggtelekiensis (Stach, J, 1930:296)

             Syn.: Pseudosinella aggtelekensis(sic) Gisin, H, 1960:251,sd ISS
             Syn.: Pseudosinella aggtelekensis(sic) Dethier, M, 1998:56 ISS
Pseudosinella aggtelekiensis from Slovakia
2012 © Kovác, L.



Species aidamar Luciáñez & Simón, 1994

Species alba (Packard, AS, 1873:37) Absolon, K, 1901:86, B

            Syn.: Tullbergia ocellata Lie-Pettersen, 1896


Pseudosinella alba from the USA
After Christiansen K, 1960 Pl.1 Fig.17

             Syn.: Pseudosinella albus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:45 ISS
Pseudosinella alba from the UK
2012.12.23 © Murray, A.
Pseudosinella alba from the UK
2012.12.23 © Murray, A.
Pseudosinella alba from France
2013.03.03 © Sandra.
Pseudosinella alba from France
Eyepatch with 2 distinct ocelli: a dorsal and ventral ocellus
2013.03.16 © Sandra.
Pseudosinella alba from Canada
2014.01.22 © Hochhalter, R.
Pseudosinella alba from Canada
2014.01.22 © Hochhalter, R.
Pseudosinella alba from Canada
2014.01.22 © Hochhalter, R.

             Syn.: Pseudosinella abla(sic) Zhang, F et Deharveng, L, 2015:494 ISS
             Syn.: Pseudosinella abla(sic) Zhang, F et Deharveng, L, 2015:501 ISS
             Syn.: Pseudosinella abla(sic) Zhang, F et Deharveng, L, 2015:303 ISS
Pseudosinella alba from France
Eyepatch with 2 ocelli
2016.11.11 © Legrand, T.
Pseudosinella alba from France
Antenna about 1.25x head diagonal
2016.11.11 © Legrand, T.



Species albida (Stach, 1930) Stomp, N, 1971:181

            Syn.: Lepidocyrtus (Pseudosinella) decem-oculatus(sic) Wahlgren, E, 1906:17
            Syn.: Lepidocyrtus wahlgreni Börner, 1907 =?


Species alfonsii Dallai, R et Malatesta, E, [1983]:187

Species alpina Gisin, 1950

Species ambigua Zeppelini, D, Brito, RA et Lima, ECA, 2018:69

Species anderseni Gisin, 1967

Species angelae da Gama, 2004

Species annemariae Stomp, N, 1972:281

Species antennata Bonet, 1929

Species apuanica Dallai, R, 1970:462

Species aramendiai Beruete E et Jordana R in Beruete, E, Baquero, E et Jordana, R, 2002:385

            Syn.: Pseudosinella picta Jordana & Beruete, 1983


Species argentea Folsom, JW, 1902:366

Pseudosinella argentea from the USA
After Christiansen K, 1960 Pl.2 Fig.34

Species argentea (Loksa, 1960) Simón-Benito, JC & Moreno, S, 2006:202

Species arrasatensis Beruete E et Jordana R in Beruete, E, Baquero, E et Jordana, R, 2002:388

Species arretzi Simon, 1979

Species ashmolae Christiansen, KA, 1998:150

             Syn.: Pseudosinella ashmolei(sic) Christiansen, KA, 1998:149 ISS

Species ashmoleorum da Gama, MM, 1988:609

Species aspinata Cassagnau, P, 1955:39

Species astarini Tshelnokov & Rasulova, 1982

Species astronomica Gisin, H et da Gama, MM, 1970:180

Species aueri Gisin, H, 1964:672

Species azerbaidzhani Tshelnokov & Rasulova, 1982

Species azorica da Gama, MM, 1988:608

Species bachae Luciañez & Simon, 1994

             Syn.: Pseudosinella bachai(sic) Simón-Benito, JC et Moreno, S, 2006:200 ISS
Remark by Jordana, R. & Baquero, E. 2007:6: P. bachae and P. gajui can be differentiated only by the shape of the tenent hair of the tibiotarsus (Luciáñez & Simón, 1994). In our opinion these two species are the same because, in addition, P. gajui has been described using only one specimen from the same locality and date of P. bachae. For this reason we consider P. gajui a new synonym of P. bachae.



Species baghdadica Rusek, 1981

Species balazuci Gisin, H et da Gama, MM, 1969:153

Species barcelonensis Gisin, H et da Gama, MM, 1969:149

Species bellingeri Wang, F, Christiansen, K et Chen, J-X, 2002:63

Species beruetei Jordana, R et Baquero, E, 2005:34

            Syn.: Pseudosinella duodecimoculata Gisin, H et da Gama, MM, 1970:296


             Syn.: Pseudosinella berueti(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:67 ISS

Species bessoni Deharveng, 1988

Species bidenticulata Barra, 1967

Species biguttata Barra, J-A, 1997:476

Species binoculata Kseneman, M, 1935:6

Species biocellata Gruia, Poliakov & Broza, 1999

Species biunguiculata Ellis, WN, 1967:103

Pseudosinella biunguiculata from Guatemala
After Ellis WN, 1967 Fig.6a
             Syn.: Pseudosinella biunguicultata(sic) Wang, F, Chen, J-X et Christiansen, K, 2004:375 ISS
             Syn.: Pseudosinella biungiculata(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:186 ISS


Species bohemica Rusek, 1979

Species bona Christiansen, K et Bellinger, P, 1996:46

Species boneti Bagnall, 1941

Species boneti Christiansen, K, 1960:24, nec Bagnall, 1941

Pseudosinella boneti from the USA
After Christiansen, K., 1960 Pl.4 Fig.65

Species bonita Christiansen, K in Mitchell, RW et Reddell, JR, 1973:129

Species brevicornis Handschin, 1924

Species bulgarica da Gama, MM, 1974:554

Species burgalensis Jordana, R et Baquero, E, 2005:2

Species cabidochei Deharveng & Gouze, 1986

Species caladairensis Gisin, H et da Gama, MM, 1970:169

Species canariensis da Gama, MM, 1974:556

Species candida Folsom, JW, 1902:366

Species cansiliensis Fanciulli, PP et Dallai, R, 2008:23

Species caoi Chen, J-X, Wang, F et Christiansen, K, 2002:80

Species carthusiana Gisin, 1963

Species cassagnaui Gisin, H et da Gama, MM, 1970:182

Species cava Christiansen, K et Reddell, JR, 1986:135

Species cavernarum (Moniez, 1893) Carpenter & Evans, 1899

            Syn.: Tullbergia immaculata Lie-Pettersen, 1896
             Syn.: Cyphoderus Martelli(sic) Carpenter, 1897 ISS
             Syn.: Cyphoderus martelli(sic) Carpenter, 1897 ISS
            Syn.: Pseudosinella immaculata ssp. tridenticulata Börner, 1903
             Syn.: Cyphroderus(sic) albinos(sic) Guthrie, 1903 ISS


             Syn.: Sira(sic) Cavernarum(sic) Bertkau, P, 1894:57 ISS

Species ccapanensis Winter, 1963, n.n.

Species cementensis Simón-Benito, JC et Romera-Becerro, J, 2008:57

Species centralis Gisin, H et da Gama, MM, 1969:155

Species certa Christiansen & Bellinger, 1980

Species chapmani da Gama, 1979

Species charoae Villanueva & Jordana, 1988

Species chauvini Stomp & Gouze, 1984

Species chiangdaoensis Deharveng, L, 1990:281

Species christiani Stomp, 1986

Species christianseni Salmon, 1965

Pseudosinella christianseni from the USA
2006 © Grinnell College.

Species ciliata Ellis, WN, 1974:125

             Syn.: Pseudosinella ciliate(sic) Palacios-Vargas, JG et Simón-Benito, JC, 2009:214 ISS

Species cobosae Luciáñez & Simón, 1994

Species codri da Gama & Busmachiu, 2002

Species coerulea Jacquemart, 1975 in Peck SB & Jacquemart J, 2011:2 nn

Species collina Wray, 1952

             Syn.: Pseudosinella colina(sic) Palacios-Vargas, JG, Castaño-Meneses, G et Estrada, DA, 2011:120 ISS

Species concii Gisin, 1950

Species confusa Izarra, 1965

Species cordobensis Simón, Bach & Gajú, 1986

Species crypta Christiansen, K et Reddell, JR, 1986:132

Species csafordensis Winkler, D et Mateos, E, 2018:348

            Syn.: Pseudosinella csafordi(sic) Winkler, D et Mateos, E, 2018:348 IOS
Pseudosinella csafordi(sic) from Hungary
After Winkler D & Mateos E, 2018 Fig.1



Species curtii Gouze & Deharveng, 1987

Species dallaii Gisin, H et da Gama, MM, 1970:873

Pseudosinella dallaii from Croatia
2010.11.19 © Lukic, M.

Species decemoculata (Guthrie, JE, 1903:85) Poole in Poole & Gentili, 1997:46

Species decepta Gisin, H et da Gama, MM, 1969:173

            Syn.: Pseudosinella decipiens Bonet, 1929 nec Denis, 1924


Species decipiens Denis, JR, 1924:198, B

            Syn.: Pseudosinella immaculata auctt., nec Lie-Pettersen, 1896
            Syn.: Pseudosinella martelli(sic) Handschin, 1921, nec Carpenter, 1895
            Syn.: Pseudosinella marteli(sic) Handschin, 1926, nec Carpenter, 1897
            Syn.: Pseudosinella sp. Denis, 1924
            Syn.: Pseudosinella martelli var. religiosa Agrell, 1939
            Syn.: Pseudosinella handschini Bagnall, 1941
            Syn.: Pseudosinella propinqua Bagnall, 1941


Pseudosinella decipiens from Belgium
2011.09.09 © Henderickx, H.

Species decui Gruia, 1995

Species delhezi Stomp, N, 1971:469

Species denisi Gisin, 1954

Species difficilis Denis, 1927

Species dispadentata Salmon, 1948

Species dobati Gisin, H, 1965:1

            Syn.: Pseudosinella martelii Denis, 1935


Species dodecophthalma Gisin, H et da Gama, MM, 1969:170

             Syn.: Pseudosinella dodecophtalma(sic) Stomp, N, 1971:185 ISS
             Syn.: Pseudosinella dodecopthalma(sic) Wang, F, Chen, J-X et Christiansen, K, 2002:243 ISS
             Syn.: Pseudosinella dodecopthalma(sic) Wang, F, Chen, J-X et Christiansen, K, 2004:384 ISS

Species dodecopsis Gisin, H et da Gama, MM, 1969:168

  Subspecies caerulea Gisin, H et da Gama, MM, 1970:176

             Syn.: Pseudosinella dodecopsis ssp. coerulea(sic) da Gama, MM, 1984:60 ISS
Pseudosinella dubia from the USA
After Christiansen, K., 1960 Pl.1 Fig.6.


Species dubia Christiansen, K, 1960:9


Species dungeri Winkler, D et Mateos, E, 2018:357

Pseudosinella dungeri from Hungary
After Winkler D & Mateos E, 2018 Fig.21

Species duodecimocellata Handschin, 1928, B

             Syn.: Pseudosinella duodecimocullata(sic) Handschin, E in Dahl, F, 1929:89 ISS
             Syn.: Pseudosinella duodecimoculata(sic) Marlier, GJ, 1942:9,B ISS
             Syn.: Pseudosinella duodecimoculata(sic) Obenberger, 1955 ISS
             Syn.: Pseudosinella duodecimocullata(sic) da Gama, MM, 1973:56 ISS
             Syn.: Pseudosinella 12-ocellata(sic) da Gama, MM, 1984:60 ISS
Pseudosinella duodecimpunctata from the USA
After Christiansen, K., 1960 Pl.1 Fig.1.


Species duodecimoculata Bonet, F, 1931:324


             Syn.: Pseudosinella 12-oculata(sic) da Gama, MM, 1984:60 ISS

Species duodecimpunctata Denis, 1931

Species duprei Beruete E et Jordana R in Beruete, E, Baquero, E et Jordana, R, 2002:389

Species edax Gisin, 1967

Species efficiens Gisin, H et da Gama, MM, 1969:289

Species encrusae Gisin, H et da Gama, MM, 1969:164

Species enriquei Arbea, JI, 2017:402

Pseudosinella enriquei from Spain
After Arbea JI, 2017 Fig.1.1
            Syn.: Pseudosinella suboculata Arbea, JI, 2017:408 partim


Species erehwon Christiansen & Bellinger, 1996:40

Species eskualduna Beruete E et Jordana R in Beruete, E, Baquero, E et Jordana, R, 2002:390


Pseudosinella espana from the USA
After Christiansen, K., 1960 Pl.2 Fig.38.
Species espana Christiansen, K, 1960:19


Species espanita Christiansen, K et Bellinger, P, 1996:40

Species espanoli Simón & Selga, 1977

             Syn.: Pseudosinella españoli(sic) Beruete, E et Jordana, R, 2001:159 ISS
             Syn.: Pseudosinella espagnoli(sic) Arbea, JI, 2006:7 ISS
             Syn.: Pseudosinella espagnoli(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2007:13 ISS
             Syn.: Pseudosinella espagnoli(sic) Simón-Benito, JC et Romera-Becerro, J, 2008:57 ISS
             Syn.: Pseudosinella espagnoli(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:186 ISS
             Syn.: Pseudosinella espagnoli(sic) Arbea, JI, 2013:305 ISS

Species espartana Beruete & Jordana, 2001

Species extra Christiansen, K et Bellinger, P, 1996:45

Species fallax (Börner, 1903) Bagnall, 1941

            Syn.: Pseudosinella intermixta Kloet, GS et Hincks, WD, 1945:7
            Syn.: Pseudosinella soniae Dallai, R, 1968:102


Pseudosinella fallax from the UK
2012.08.02 © Murray, A.
Pseudosinella fallax from the UK
Eyepatch with 3 ocelli
2012.08.02 © Murray, A.

Species fasciata Womersley, 1934

Species federicoi Simón-Benito, JC et Palacios-Vargas, JG, 2008:182

Species feneriensis Gisin, H, 1962:21

Species finca Christiansen, K in Mitchell, RW et Reddell, JR, 1973:130

Species fjellbergi da Gama, MM, 1974:556

Species flatua Christiansen, K et Bellinger, P, 1996:41

Species folsomi (Mills, HB, 1931:10)

Species fonsa Christiansen, K et Bellinger, P, 1996:42

Species fujiokai Yosii, R, 1964:13

Species gamae Gisin, 1967

             Syn.: Pseudosinella gama(sic) Soto-Adames, FN, 2010:30 ISS

Species genghizkhani Yosii, 1966

Species georgia Christiansen, K et Bellinger, P, 1998:1097

Species germani da Gama, 1973

Species ghirkhani Tshelnokov & Rasulova, 1982

Species gineti Cassagnau, P, 1955:39

Species gisini Christiansen, K, 1960:14

Pseudosinella gisini from the USA
After Christiansen K, 1960 Pl.2 Fig.25
             Syn.: Pseudosinella gilini?(sic) Camousseight, A, 1999:6 ISS
  Subspecies carolina Christiansen, K et Bellinger, P, 1996:48

  Subspecies virginia Christiansen, K et Bellinger, P, 1996:47



Species gleycola Rapoport, 1962

Species goughi Gisin, H et da Gama, MM, 1972:276

             Syn.: Pseudosinella gougi(sic) Jordana, R et Baquero, E, 2005:28 ISS

Species granda Christiansen, K et Bellinger, P, 1996:43

Species gruiae da Gama & Busmachiu, 2002

Species guanhaensis Zeppelini, D, Brito, RA et Lima, ECA, 2018:73

Species gutierrezae Simón-Benito, JC et Palacios-Vargas, JG, 2008:63

Species hahoteana Soto-Adames FN, Bernard EC & Wynne JJ in Bernard EC, Soto-Adames FN & Wynne JJ, 2015:257

Pseudosinella hahoteana from Rapa Nui
After Bernard EC & al, 2015 Fig.7C

Species halophila Bagnall, 1939

             Syn.: Pseudosinella halofila(sic) Jordana, R et Baquero, E, 2007:2 ISS

Species hauseri da Gama, MM, 1973:45

Species helenae Arbea, JI et Jordana, R, 1990:27

Species hercynica Rusek, 1979

Species hermanni Dallai, 1976

Species heteromurina (Stach, J, 1930:295) da Gama, MM, 1974:552

            Syn.: Pseudosinella jeanneli Gisin, H et da Gama, MM, 1971:223


Pseudosinella heteromurina from Croatia
2010.11.19 © Lukic, M.


Species hirsuta (Delamare Deboutteville, 1949) Christiansen, K, 1960:22

Pseudosinella hirsuta from the USA
After Christiansen, K., 1960 Pl.3 Fig.51.
Pseudosinella hirsuta from the USA
1998 © Taylor, S.

             Syn.: Pseudosinella hisurta(sic) Palacios-Vargas, JG in Sánchez-Ramos, G, Reyes-Castillo, P et Dirzo, R, 2005:360 ISS

Species horaki Rusek, 1985

Species hrabei Rusek, 1979

             Syn.: Pseudosinella hrabrei(sic) Simón-Benito, JC et Moreno, S, 2006:202 ISS
             Syn.: Pseudosinella hrabrei(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:186 ISS

Species huautla Christiansen, K, 1982:11

             Syn.: Pseudosinella hautla(sic) Palacios-Vargas, JG et Mejía Recamier, BE, 2011:52 ISS

Species huescensis Gisin, H et da Gama, MM, 1969:165

Species huetheri Stomp, N, 1971:176

            Syn.: Pseudosinella hütheri(sic) Stomp, N, 1971:176 IOS
            Syn.: Pseudosinella sp. Stomp, N, 1971:175
            Syn.: Pseudosinella wahlgreni Hüther, 1969


             Syn.: Pseudosinella hutheri(sic) da Gama, MM, 1984:60 ISS
             Syn.: Pseudosinella hütheri(sic) Stomp, N et Weiner, WM, 2005:230 ISS
             Syn.: Pseudosinella hutheri(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:61 ISS



Species hui Wang, F, Chen, J-X et Christiansen, K, 2003:240

Species illiciens Gisin, 1967

Species immaculata (Lie-Pettersen, 1896) Folsom, 1902, t.t.

            Syn.: Pseudosinella argentata(sic) Folsom, 1902
            Syn.: Lepidocyrtus cavernarum Stach, 1922
            Syn.: Cyphoderus martelii Carpenter, 1895
             Syn.: Pseudosinella marteli(sic) Carpenter, 1897 ISS
             Syn.: Pseudosinella martelli(sic) Handschin, E in Dahl, F, 1929:91 ISS
             Syn.: Pseudosinella martelli(sic) Obenberger, J, 1955:99 ISS


             Syn.: Pseudosinella inmaculata(sic) Beruete, E, Baquero, E et Jordana, R, 2002:392 ISS
             Syn.: Pseudosinella immaculate(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2007:18 ISS


Pseudosinella immaculata from the UK
2011.10.22 © Jones, A.
Pseudosinella immaculata from the UK
2011.10.22 © Jones, A.
Pseudosinella immaculata from the UK
2013.04.11 © Murray, A.
Pseudosinella immaculata from the UK
2013.12.12 © Murray, A.

Species imparipunctata Gisin, 1953

Species impediens Gisin, H et da Gama, MM, 1969:147

             Syn.: Pseudosinella impidens(sic) Soto-Adames, FN, 2015:3 ISS
             Syn.: Pseudosinella impidens(sic) Soto-Adames, FN, 2016:3 ISS


Species inaequalis Bagnall, 1941

Species inaequalis Stach, 1960, nec Bagnall, 1941

Species infernalis Gisin, H, 1964:670

Species inflata Bonet, F, 1931:334

Species infrequens Gisin, H et da Gama, MM, 1969:145

Species insoloculata Salmon, JT, 1941:396

Species insubrica Gisin, H et da Gama, MM, 1969:292

            Syn.: Pseudosinella cavernarum Carl, 1906
            Syn.: Sirodes lamperti Handschin, 1926
            Syn.: Pseudosinella vandeli f. longicornis Denis, 1937 =?
            Syn.: Pseudosinella vandeli ssp. alpina Gisin, 1960 partim


             Syn.: Pseudosinella insubricia(sic) Christiansen, K, 1982:22 ISS

Species insularum Dallai, 1969

Species intemerata Gisin, H et da Gama, MM, 1969:175

Species intermixta (Folsom, 1924) Winter, 1963

             Syn.: Pseudosinella intermixtus(sic) Winter, 1963 ISS
             Syn.: Pseudosinella inermixta(sic) Katz, AD, Taylor, SJ, Soto-Adames, FN, Addison, A, Hoese, GB, Sutton, MR et Toulkeridis, T, 2016:94 ISS

Species ioni da Gama & Busmachiu, 2002

Species jacetanica Jordana, R et Baquero, E, 2007:9

Species jeanpierrei Beruete E et Jordana R in Beruete, E, Baquero, E et Jordana, R, 2002:390

Species jesusi Beruete E et Jordana R in Beruete, E, Baquero, E et Jordana, R, 2002:392

             Syn.: Pseudosinella jesus(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2007:18 ISS

Species jordanai Simón-Benito, JC et Palacios-Vargas, JG, 2007:12

Species josemarii Soto-Adames, FN, 2010:6

Species joupani Denis, 1933

             Syn.: Pseudosinella joupai(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:62 ISS

Species kalalauensis Christiansen & Luther, 1986:49

            Syn.: Pseudosinella kalaluensis(sic) Christiansen & Luther, 1986 IOS

Species kenchristianseni Gruia, M, 1984:21

             Syn.: Pseudosinella kenchrystianseni(sic) Velázquez, DR et Novoa, NM, 2002:7 ISS

Species kersmaekersi Stomp, 1974

Species ksenemani Gisin, 1944, B

Species kwartirnikovi da Gama, MM, 1973:57

Species lafargensis Simón-Benito, JC et Romera-Becerro, J, 2008:57

Species lahainaensis Christiansen & Luther, 1986:46

             Syn.: Pseudosinella lahainensis(sic) Christiansen & Luther, 1986 ISS
             Syn.: Pseudosinella lahianensis(sic) Wang, F, Chen, J-X et Christiansen, K, 2004:375 ISS
             Syn.: Pseudosinella lahainensis(sic) Wang, F, Chen, J-X et Christiansen, K, 2004:375 ISS

Species lamperti (Schäffer, 1900) Gisin, 1963

             Syn.: Sirodes Lamperti(sic) Absolon, K, 1901:379 ISS
  Subspecies germanica da Gama, MM, 1973:54



Species larisae da Gama & Busmachiu, 2002

Species lava Christiansen, KA, 1998:152

Species lechuzasensis Winter, 1963, n.n.

Species leclerci Stomp, Thibaud & Massoud, 1982

             Syn.: Pseudosinella lecrerci(sic) Beruete, E et Jordana, R, 2001:159 ISS

Species leivaensis Simón-Benito, JC et Palacios-Vargas, JG, 2008:62

Species lenkorani Tshelnokov & Rasulova, 1982

Species leoni Christiansen, K, 1982:19

Species lesi Jordana R, Baquero E et Janssens F in Baquero, E, Janssens, F, Les, J et Jordana, R, 2006:72

Pseudosinella sp. nov. from Spain
2005 © Les, J.

Species lleidensis da Gama, MM, 1984:86

Species logrognensis Simón-Benito, JC et Palacios-Vargas, JG, 2008:59

             Syn.: Pseudosinella logrongnesis(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:61 ISS

Species luenganensis Simón-Benito, JC et Palacios-Vargas, JG, 2007:11

             Syn.: Pseudosinella luengasensis(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2007:16 ISS

Species lunaris Gisin, H et da Gama, MM, 1970:173

Species luquei Beruete E et Jordana R in Beruete, E, Baquero, E et Jordana, R, 2002:393

            Syn.: Pseudosinella duodecimoculata Bonet, F, 1931:324 partim
            Syn.: Pseudosinella luquei Authorunknown, 1999 nn


Species mallorquinensis Palacios-Vargas, JG et Simón-Benito, JC, 2009:214

Species manuelae Gruia, M, 1973:55

Species marcuzzii Rusek, 1985

             Syn.: Pseudosinella marcuzzi(sic) Wang, F, Christiansen, K et Chen, J-X, 2002:53 ISS
             Syn.: Pseudosinella marcuzzi(sic) Wang, F, Chen, J-X et Christiansen, K, 2004:382 ISS

Species maritima Bagnall, 1941

Species maros Deharveng & Suhardjono, 2004

Species mauli Stomp, N, 1972:279, B

Species meganporteri Soto-Adames, FN, 2010:10

             Syn.: Pseudosinella megaporteri(sic) Zeppelini, D, Brito, RA et Lima, ECA, 2018:73 ISS

Species melatensis Gisin, H et da Gama, MM, 1969:163

Species metallica Jacquemart, 1980:2

Species miratio Christiansen, KA, 1998:154

Species moldavica da Gama & Busmachiu, 2002

Species montis Yosii, 1971

Species montiscarmeli Gruia, Poliakov & Broza, 1999

Species monzoni Winter, 1963, n.n.

Species mucronata Gouze & Deharveng, 1987

Species mutabilis Wang, F, Christiansen, K et Chen, J-X, 2003:603

Species najtae Jordana R et Baquero E in Jordana, R, Barranco, P, Amezcua, A et Baquero, E, 2017:110

Pseudosinella najtae from Spain
After Jordana R & Baquero E in Jordana R & al, 2017 Fig.2D

Species nata Christiansen & Bellinger, 1980

Species navarrensis Ardanaz & Jordana, 1985

Species neosensillata Winter, 1963, n.n.

Species nonoculata Salmon, JT, 1941:397

Species noseki Rusek, 1985

Species obanae Gruia, 1998

Species occidentalis Dallai, 1976

             Syn.: Pseudosinella occidentals(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2007:17 ISS

Species ocellata (Absolon K, 1901:378) Thibaud, J-M & Deharveng, L in Jubertie, C & Decu, V, 1994:268

Species octoculata (Caroli, 1914) Womersley, 1934

             Syn.: Pseudosinella octooculata(sic) Womersley, 1934 ISS
Lepidocyrtus octoculatus from Palestine
After Handschin E, 1942 Fig.5a



Species octophthalma Gisin, H et da Gama, MM, 1970:178

             Syn.: Pseudosinella octophtalma(sic) Stomp, N, 1971:184 ISS

Species octopunctata Börner, 1901, B

            Syn.: Lepidocyrtus octoculatus Handschin, E in Dahl, F, 1929:89
            Syn.: Lepidocyrtus octoculatus Gisin, H, 1960:243 =?


Pseudosinella octopunctata from Japan
After Yosii R, 1942 Abb.17.1

             Syn.: Pseudosinella octopunctala(sic) Stomp, N, 1971:185 ISS
Pseudosinella nr octopunctata from the UK
2012.12.29 © Murray, A.
Pseudosinella nr octopunctata from the UK
2012.12.29 © Murray, A.
Pseudosinella nr octopunctata from the UK
2012.12.29 © Murray, A.



Species ops Christiansen, K et Bellinger, P, 1998:1107


Pseudosinella orba from the USA
After Christiansen, K., 1960 Pl.3 Fig.43.
Species orba Christiansen, K, 1960:20

Species oromii da Gama, 1996

Species oxapampensis Winter, 1963, n.n.

Species oxybarensis Gisin, H et da Gama, MM, 1969:160

Species paclti Rusek, 1961

            Syn.: Pseudosinella cavernarum Paclt, 1957


Pseudosinella paclti from Slovakia
2012 © Kovác, L.

Species palaciosi Christiansen, K et Reddell, JR, 1986:141

Species pallida Gruia, M, 1977:78

Species paprivata Ellis, WN, 1976:312

Species parattenuata Simón-Benito, JC et Palacios-Vargas, JG, 2008:184

            Syn.: Pseudosinella petterseni var. attenuata Bonet, 1934


Species passaueri Stomp & al., 1991

Species paxi Nosek, 1964

Species pecki Christiansen & Bellinger, 1980

Species perezi Arbea, JI, 2013:304

Pseudosinella perezi from Spain
After Arbea JI, 2013 Fig.13

Species peruensis Jacquemart, 1980:2

Species petrustrinatii Reddell & Elliott, 1973

             Syn.: Pseudosinella petrustinatii(sic) Cutz-Pool, LQ, García-Gómez, A et Palacios-Vargas, JG, 2007:50 ISS

Species petterseni Börner, 1901, B

            Syn.: Pseudosinella martelli var. talpae Agrell, 1939


Species picta (Börner, 1903) Handschin, E, 1942:413

Species pieltaini Bonet, 1929

Species pseudofolsomi Winter C, 1963:503 n.n.

Remark by Winter, C. 1963:395: 37 von mir neu aufgestellte Arten bzw. Formen sind in der vorliegende Arbeit mit n.sp. bzw. n.f. (ohne Autorennamen) bezeichnet und müssen bis zu ihrer Beschreibung (voraussichtlich im kommenden Jahr) als Nomina nuda betrachtet werden.
Species plachteri Stomp, 1986

Species pongei da Gama, 1979

Species pori Gruia, Poliakov & Broza, 1999

Species praecipiens Gisin, 1967

Species problematica Gisin, H et da Gama, MM, 1971:219

Species pseudodecepta Arbea, JI et Jordana, R, 1990:25

Species pseudolanuginosa Yosii, R, 1942:493

Lepidocyrtus pseudolanuginosus from Japan
After Yosii R, 1942 Abb.16.1
            Syn.: Pseudosinella duodecimocellata Handschin, 1928
            Syn.: Pseudosinella duodecimocellata Yosii, 1954
            Syn.: Pseudosinella duodecimocellata Yosii, R, 1977:161


Species pseudopetterseni Jordana, R et Baquero, E, 2007:2

Species pygmaea da Gama & Busmachiu, 2004

Species pyrenaea Bonet, F, 1931:345

             Syn.: Pseudosinella pyrenaica(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2007:18 ISS

Species rabonica Gruia, M, 1984:19

Species racovitzai Gisin, H et da Gama, MM, 1971:221

             Syn.: Pseudosinella racoviitzai(sic) Wang, F, Chen, J-X et Christiansen, K, 2004:377 ISS

Species rapoporti Izarra, 1965

Species recipiens Gisin, 1967

Species reddelli Christiansen in Mitchell & Reddell, 1973:130

Species riojana Beruete, E et Jordana, R, 2001:156

Species rochezi Palacios-Vargas, JG et Mejía Recamier, BE, 2011:49

Pseudosinella rochezi from Mexico
After Palacios-Vargas JG & Mejía Recamier BE, 2011 Fig.1

Species rolfsi Mills, 1932

             Syn.: Pseduosinella(sic) rolfsia(sic) Brand, RH et Dunn, CP, 1998:239 ISS
             Syn.: Pseudosinella rolfi(sic) Simón-Benito, JC et Moreno, S, 2006:202 ISS
Pseudosinella rolfsi from the USA
2006.11.28 © McClarin, J.
Det. Christiansen, K.
Pseudosinella rolfsi from the USA
2006.11.28 © McClarin, J.
Det. Christiansen, K.
Pseudosinella rolfsi from the USA
2006.11.28 © McClarin, J.
Det. Christiansen, K.
Pseudosinella rolfsi from the USA
2006.11.28 © McClarin, J.
Det. Christiansen, K.
Pseudosinella rolfsi from the USA
2006.11.28 © McClarin, J.
Det. Christiansen, K.
Pseudosinella rolfsi from the USA
2006.11.28 © McClarin, J.
Det. Christiansen, K.
Pseudosinella rolfsi from the USA
2007.11.14 © Murray, T.
Pseudosinella rolfsi from the USA
2007.11.14 © Murray, T.
Pseudosinella rolfsi from the USA
2008.03.14 © Murray, T.
Pseudosinella rolfsi from the USA
2009.03.10 © Cowen, R.
Pseudosinella rolfsi from the USA
2011.03.05 © Nanz, S.
Pseudosinella rolfsi from the USA
2011.11.06 © Murray, T.
Pseudosinella rolfsi from the USA
2012.03.24 © Kouri, J.
Pseudosinella rolfsi from the USA
2014.11.07 © Silsbee, D.
Pseudosinella rolfsi from the USA
In nest of Formica
2017.08.20 © Bailey, J.
Pseudosinella rolfsi from the USA
2019.01.15 © Bairstow, A.



Species salisburgiana Gisin, H et da Gama, MM, 1969:157

Species sandelsorum Gruia, M, 1977:80

Species sanguinea (Jackson, AD, 1909:531)

Lepidocyrtus sanguineus from the USA
Adult and two juvenile instar forms
After Jackson, A.D., 1909 Pl.30 fig.1, 5 & 6.

Species selgae Simón, 1978

Species sensillifera Yoshii, R et Suhardjono, YR, 1989:60

Species sera Christiansen & Bellinger, 1980

Species serrana Luciañez & Simón, 1994

Species sexoculata Schött, 1902

            Syn.: Entomobrya (Sinella) aglis Harvey, 1900
             Syn.: Sinella agilis(sic) Gisin ISS


Pseudosinella sexoculata from the USA
After Christiansen, K., 1960 Pl.1 Fig.12
Pseudosinella sexoculata from the USA
2008.11.22 © Justis, S.
Pseudosinella sexoculata from the USA
2008.11.22 © Justis, S.
Pseudosinella sexoculata from the UK
2012.12.29 © Murray, A.
Pseudosinella sexoculata from the UK
2012.12.29 © Murray, A.

Species seyleri Stomp & Tommasi-Ursone, 1989

Species simoni Jordana, R et Baquero, E, 2007:13

Species simpatica da Gama & Busmachiu, 2002

Species sollaudi Denis, JR, 1924:198

            Syn.: Pseudosinella Sollaudi(sic) Denis, JR, 1924:198 IOS


             Syn.: Pseudosinella Sollaudi(sic) Delamare Deboutteville, C, 1945:47 ISS



Species soriensis Simón-Benito, JC et Moreno, S, 2006:200

Species spelunca Salmon, JT, 1958:710

Species spinosa (Delamare Deboutteville, 1949) Christiansen, K, 1960:23

Pseudosinella spinosa from the USA
After Christiansen, K., 1960 Pl.4 Fig.62.
Pseudosinella spinosa from the USA
2007.04.27 © Wolf, G.
Pseudosinella spinosa from the USA
2007.04.27 © Wolf, G.

Species stachi Christiansen & al., 1983

Species staryi Rusek, 1981

Species stewartpecki Katz AD, Soto-Adames FN et Taylor SJ in Katz, AD, Taylor, SJ, Soto-Adames, FN, Addison, A, Hoese, GB, Sutton, MR et Toulkeridis, T, 2016:95

Species stompi Gisin, H et da Gama, MM, 1970:177

Species strinatii Christiansen, K in Mitchell, RW et Reddell, JR, 1973:130

             Syn.: Pseudosinella strinati(sic) Gruia, M, 1973:57 ISS

Species stygia Bonet, F, 1931:339

Species styriaca Neuherz, H et Nosek, J, 1975:399

Species subcentralis da Gama, 1985

Species subdobati Gisin, H et da Gama, MM, 1970:167

Species subduodecima Gisin, H et da Gama, MM, 1970:301

Species subefficiens Gisin, H et da Gama, MM, 1970:163

Species subilliciens Mateos, 1993

Species subinflata Gisin, H et da Gama, MM, 1969:151

Species suboculata Bonet, F, 1931:326

Species substygia Gisin, H et da Gama, MM, 1969:171

Species subterranea Bonet, 1929

             Syn.: Pseudosinella subterranean(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2007:18 ISS
  Subspecies baztanensis Beruete E et Jordana R in Beruete, E, Baquero, E et Jordana, R, 2002:395



Species subvirei Bonet, F, 1931:335

Remark by Jordana, R. 2008.11.27:in litt.: Probably subvirei is a synonym of virei; the only difference is the relative distance from the base of the distal impaired tooth on the claw.

Species superduodecima Gisin, H et da Gama, MM, 1970:298

Species superoculata Gisin, H et da Gama, MM, 1969:167

Species tarraconensis Bonet, 1929

  Subspecies longicornis Bonet, 1929

Species templadoi Simón & Selga, 1977

Species terricola Gisin, 1967

Species testa Christiansen & Bellinger, 1980

Species theodoridesi Gisin, H et da Gama, MM, 1969:176

            Syn.: Pseudosinella theodoridesi Cassagnau in Cauchois & Théodoridès, 1954 nn


Species thibaudi Stomp, 1977

Species tietarensis Jordana, R et Baquero, E, 2007:7

Species torcuatoensis Simón-Benito, JC et Palacios-Vargas, JG, 2008:66

Species tridenticulata Börner, 1903

Species tridentifera Rusek, 1971

Species trioculata da Gama, MM, 1988:81

Species truncata Cassagnau, P, 1955:36

Species tumula Wang, F, Chen, J-X et Christiansen, K, 2002:243

Species turiasonensis Arbea, JI, 2006:1

            Syn.: Pseudosinella sp1 Arbea, JI, 2005:43


Species turolensis Arbea, JI, 2006:5

Species tyrrhena da Gama & Dallai, 1978

Species unguiculata Bonet, 1929

Species unguilonginea Jordana & Beruete, 1983

Species unioculata Womersley, 1934

Species vallis Yosii, 1971

Species vandeli Denis, 1923, B

  Subspecies meridionalis Gisin, H, 1964:668

  Subspecies praealpina Gisin, H, 1964:669

  Subspecies relicta Gisin, H, 1964:667

Species variabilis da Gama & Busmachiu, 2004

Species vera Christiansen, K, 1982:14

Species vertamicoriensis Gisin, 1963

Species vespera Christiansen, K et Bellinger, P, 1996:44

Species violenta (Folsom, JW, 1924:10) Spencer & Stracener, 1929

Lepidocyrtus violentus from the USA
After Folsom, J.W., 1924 Fig.28.
            Syn.: Cyphodeirus albinus Guthrie, 1903
            Syn.: Pseudosinella petterseni Denis, 1931
            Syn.: Pseudosinella folsomi Denis, 1931



             Syn.: Pseudosinella violentus(sic) Poole in Poole & Gentili, 1997 ISS
Pseudosinella violenta from the USA
2006 © Murray, T.
Det. Christiansen, K.
Pseudosinella violenta from the USA
2009.01.31 © Houston, S.



Species violeta Mari Mutt, JA, 1986:17

            Syn.: Pseudosinella violets(sic) Mari Mutt, JA, 1986:17 IOS

Species virei Absolon, K, 1901:87

Remark by Gisin, H. 1960:251: Species dubium.

             Syn.: Pseudosinella viréi(sic) Obenberger, J, 1955:99 ISS

Species vita Christiansen & Bellinger, 1980

Species volca Christiansen, K, 1982:7

Species vornatscheri Gisin, H, 1964:673

Species voylesi Christiansen, K, 1982:16

             Syn.: Pseudosinella voylesii(sic) Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:28,ca ISS

Species vulcana Katz AD, Soto-Adames FN et Taylor SJ in Katz, AD, Taylor, SJ, Soto-Adames, FN, Addison, A, Hoese, GB, Sutton, MR et Toulkeridis, T, 2016:101

Species wahlgreni (Börner, 1901) Gisin, H, 1960:242

             Syn.: Pseudosinella whalgreni(sic) Beruete, E, Baquero, E et Jordana, R, 2002:388 ISS

Species xabieri Ardanaz & Jordana, 1985

Species yosiiana Salmon, 1964

            Syn.: Pseudosinella truncata Yosii, R, 1959:37


Species yuca Christiansen, K, 1982:22

Species zaragozana Arbea, JI, 2006:2

Species zygophora Schille, 1908

            Syn.: Pseudosinella wahlgreni? Skarzynski, D, Pomorski, RJ, Smolis, A, Weiner, WM, Szeptycki, A, Slawska, M et Sterzynska, M, 2002:38


             Syn.: Pseudosinella zygophorus(sic) Vysotskaja, SO et Nosek, J, 1974:253 ISS



Genus Lepidiaphanus Salmon, JT, 1949:41

Species eudyptidus Salmon, JT, 1949:42

Species kashmirensis (Arora & Singh, 1962, i.s.) Arora & Singh, 1962


Genus Pseudocyrtus Salmon, 1956

Species dentatus Prabhoo, 1967

Species projectus Salmon, 1956

Species salmoni Prabhoo, 1967


Genus Sinelloides Bonet, F, 1942:58

Species adamsoni (Carpenter, 1935) Bonet, F, 1942:58, t.t.

Species dubius Bonet, F, 1942:58


Genus Metasinella Denis, 1929

  Syn.: Entomobryinae Metacinella(sic) González, V, Díaz, M et Palacios-Vargas, JG, 1997:47 ISS

Subgenus Metasinella Mari Mutt, JA et Gruia, MM, 1983:134

Species acrobates Denis, 1929, t.t.

Metasinella acrobata(sic) from Cuba
After Massoud Z & Gruia M, 1973 Fig.7A
             Syn.: Metasinella acrobata(sic) Massoud, Z et Gruia, M, 1973:340 ISS
Subgenus Sulcuncus Mills, HB in Pearse, AS, 1938:188 key


Species borincana Mari Mutt, JA et Gruia, MM, 1983:139

Species coralia Mari Mutt, JA et Gruia, MM, 1983:141

Species falcifera (Mills, HB in Pearse, AS, 1938:190) Bonet, 1944, t.t.

             Syn.: Metasinella falciferus(sic) Zeppelini Filho, D et Castaño Meneses, G, 1995:10 ISS

Species millsi Mari Mutt, JA et Gruia, MM, 1983:145

Species nunezi Massoud, Z et Gruia, M, 1973:335

Species radkei Soto-Adames, FN et Anderson, EW, 2017:35

Species rapoporti Massoud, Z et Gruia, M, 1973:337

Metasinella rapoporti from Cuba
After Massoud Z & Gruia M, 1973 Fig.6F

Species subfusa (Wray, DL, 1953:149) Thibaud & Massoud, 1980

Species topotypica Bonet, 1944

Metasinella topotypica from Cuba
After Massoud Z & Gruia M, 1973 Fig.7B

Species wrayi Mari Mutt, JA et Gruia, MM, 1983:142


Tribus Bromacanthini Mitra, 1993 sensu Janssens, 2018 key

Remark by Mitra, S.K. 1993:62: General chaetal pattern of Lepidonella and Bromacanthus comes close to some genera under Lepidocyrtini.
             Syn.: Paronellinae Bromocanthini(sic) Soto-Adames, FN, Jordana, R et Baquero, E, 2014:15 ISS
             Syn.: Paronellidae Bromocanthini(sic) Soto-Adames, FN et Bellini, BC, 2015:331 ISS
Remark by Zeppelini D & Oliveira J.L.V.C. 2016:569: The tribe Bromacanthini is more similar to some Entomobryidae than to Paronellini+Cyphoderini.
Remark by Janssens, F. 2018.11.04: We propose to place Bromacanthini in Lepidocyrtinae, in line with the the morphological study of Zeppelini & Oliveira, 2016.




Genus Lepidonella Yosii, 1960

  Syn.: Paronellini Microparonella Carpenter, GH, 1916:43


Lepidonella
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.37
Lepidonella sp. from Vietnam
2009 © Boldsystems.org
Lepidonella sp. from Vanuatu
2010 © Deharveng, L. & Bedos, A.
Lepidonella? sp. from New Zealand
2017.12.08 © Navest, M.P.
Lepidonella? sp. from New Zealand
2018.10.22 © Navest, M.P.

Species sp.1 Bellini, BC et Zeppelini, D, 2004:588

Species annulicornis (Oudemans, JT in Weber, M, 1890:87) Mitra, SK, 1992:212

Lepidonella annulicornis from Indonesia
After Yoshii in Soto-Adames FN & Bellini BC, 2015 Fig.36
Lepidonella annulicornis from Indonesia
After Yoshii in Soto-Adames FN & Bellini BC, 2015 Fig.37

Species ceylonica (Yosii, 1966) Deharveng, L & Bedos, A, 1995:22

Cyphoderopsis ceylonica from India
After Mandal GP & Hazra AK, 2009 Fig.43a
Lepidonella ceylonica from Ceylon
After Yosii in Soto-Adames FN & Bellini BC, 2015 Fig.35

Species doveri (Carpenter, 1932) Deharveng & al, 2018

Species duodecimoculata (Prabhoo, 1971) Deharveng, L & Bedos, A, 1995:22

Lepidonella duodecimoculata from India
After Prabhoo in Soto-Adames FN & Bellini BC, 2015 Fig.31

Species flava (Carpenter, GH, 1916:44) Deharveng, L & Bedos, A, 1995:22

            Syn.: Paronella seychellesiensis Salmon, JT, 1964:545
             Syn.: Microparonella seychellesensis(sic) Yoshii, 1981 ISS
             Syn.: Microparonella seychellensis(sic) Yoshii, R et Suhardjno, YR, 1992:48 ISS
             Syn.: Paronella seychelliensis(sic) Nguyen, M et Soto-Adames, FN, 2018:193 ISS


Species incerta (Handschin, 1925) Mitra, SK, 1992:265

            Syn.: Lepidonella oudemansi Soto-Adames, FN et Bellini, BC, 2015:337 =?


Species kei (Yoshii, R & Suhardjno, YR, 1992:47) Deharveng, L & Bedos, A, 1995:22

Microparonella kei from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.27A right
Microparonella kei from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.27A left

             Syn.: Lepidonella key(sic) Soto-Adames, FN et Bellini, BC, 2015:339 ISS


Species lecongkieti Deharveng, L et Bedos, A, 1995:23

Species marimutti Soto-Adames, FN et Bellini, BC, 2015:334

            Syn.: Lepidonella marimuti(sic) Soto-Adames, FN et Bellini, BC, 2015:334 IOS
            Syn.: Microparonella incerta Mari Mutt, JA, 1987:404


             Syn.: Lepidonella marimuti(sic) Bellini, BC et Cipola, NG, 2017:176 ISS
             Syn.: Lepidonella marimuti(sic) Nguyen, M et Soto-Adames, FN, 2018:194 ISS



Species nigrofasciata (Handschin, 1928) Deharveng, L & Bedos, A, 1995:22

Lepidonella nigrofasciata from Indonesia
After Handschin in Soto-Adames FN & Bellini BC, 2015 Fig.33

Species oudemansi (Yoshii, 1983) Deharveng, L & Bedos, A, 1995:22

Lepidonella oudemansi from Malaysia
After Yoshii in Soto-Adames FN & Bellini BC, 2015 Fig.32

Species picta (Schäffer, C, 1898:410) Mitra, SK, 1992:212

Species subcarpenteri (Denis, 1948) Deharveng, L & Bedos, A, 1995:22

Lepidonella subcarpenteri from Indonesia
After Yoshii in Soto-Adames FN & Bellini BC, 2015 Fig.38

Species tokiokai Yosii, 1960 tt

Lepidonella tokiokai from the Solomon Islands
After Yosii in Soto-Adames FN & Bellini BC, 2015 Fig.34

Species zeppelinii Soto-Adames, FN et Bellini, BC, 2015:332

Lepidonella zeppelinii from Brazil
After Soto-Adames FN & Bellini BC, 2015 Fig.1
             Syn.: Lepidonella zeppelini(sic) Zeppelini, D et Oliveira, JVLC, 2016:569 ISS
             Syn.: Lepidonella zepellinii(sic) Nguyen, M et Soto-Adames, FN, 2018:192 ISS
             Syn.: Lepidonella zeppelini(sic) Nguyen, M et Soto-Adames, FN, 2018:194 ISS



Genus Bromacanthus Schött, 1925

  Syn.: Paronellinae Glacialoca Salmon, JT, 1941:405
  Syn.: ... Handschinella Yosii, R, 1959:53


Bromacanthus
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.35

Species sp.1 Yoshii, 1983

Species sp.2 Yoshii, 1983

Species elongatus Yoshii, 1981

Species flavidulus Yoshii, 1981

Species halmaherae Yoshii, 1983

Pseudoparonella (Irianella) halmaherae from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.24A

Species handschini Schött, 1925, t.t.

             Syn.: Pseudoparonella (Bromacanthus) Handschini(sic) Absolon, K et Ksenemann, M, 1942:14 ISS

Species orientalis (Handschin, 1930)

Species palawanicus Yoshii, 1983

Species sago Yoshii, R et Suhardjno, YR, 1992:49

Bromacanthus sago from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.28A

Species seramensis Yoshii, 1989

Species setigerus (Börner, C, 1906:178) Yoshii, 1981

             Syn.: Bromacanthus setigera(sic) Mitra, SK, 2003:84 ISS

Species wallacei Yoshii, R et Greenslade, P, 1993:82


Genus Trichorypha Schött, H, 1893:16

Species atrofasciata Schött, H, 1893:18, t.t.

            Syn.: Trichorypha atro-fasciata(sic) Schött, H, 1893:18 IOS
            Syn.: Trogolaphysa lamottei Nguyen, M et Soto-Adames, FN, 2018:184 =?



References