http://www.collembola.org/taxa/smi1idae.htm - Last updated on 2023.05.26 by Frans Janssens
Checklist of the Collembola: Sminthurididae

Familia Sminthurididae Börner, 1906:163 sensu Medeiros G & al, 2022:2 key

Genus Number
of species
Boernerides1
Debouttevillea1
Denisiella15
Parasminthurides2
Pedonides1
Pseudosminthurides1
Pygicornides2
Sinnamarides1
Sminthurides66
Sminthuridia1
Sphaeridia70
Stenacidia3
Yosiides2
Sminthurididae166
 Genera with number of species
  Syn.: ... Smynthuridae(sic) Lubbock, 1862, partim Lubbock, 1870
  Syn.: Collembola Sminthurides(sic) Willem, V, 1900:55 ISS
  Syn.: Symphypleona Smynthuridae(sic) Kloet, GS et Hincks, WD, 1945:8 ISS
  Syn.: Collembola Sminthuridae(sic) Sun, X, Bedos, A et Deharveng, L, 2018:4 lapsus ISS
  Syn.: Symphypleona Sminthuridae(sic) Mateos, E et Lukic, M, 2019:222 lapsus ISS
  Syn.: Eusymphypleona Sminthuridida(sic) Bretfeld, 1986
  Syn.: Symphypleona Sminthuridida(sic) Bretfeld, G, 1994:13
  Syn.: Eusymphypleona Sminthuridida(sic) Bretfeld, G, 1999:29


  Syn.: Symphypleona sminthurididae(sic) Massoud, Z et Betsch, J-M, 1972:55 ISS
  Syn.: Symphypleona Sminthiridididae(sic) Susanti, WI, Widyastuti,R, Scheu, S et Potapov, A, 2021:11 ISS


Sminthurides aquaticus from Belgium
2000 © Hopkin, S.P.
Sphaeridia serrata from the USA
2006 © Maddison, D.R.


Genus Pseudosminthurides Sánchez-García, A et Engel, MS, 2016:4, -

Species stoechus Sánchez-García, A et Engel, MS, 2016:4, -

Pseudosminthurides stoechus in Cretaceous amber from Spain
After Sánchez-García A & Engel MS, 2016 Fig.2


Tribus Sphaeridiini Richards, 1968:18 sensu Janssens F, 2022

Remark by Janssens, F. 2022.11.13: Based on Betsch (1980:125-127) and Medeiros & al (2022:7,19), tentatively we define Sphaeridiini as genera with the distal tibiotarsus III organ absent.
Remark by Pentinsaari, M. 2023.03.07: The tribe Sphaeridiini Richards, 1968 (Sminthurididae) is a junior homonym of Sphaeridiini Latreille, 1802, used at both subfamily and tribe ranks in Coleoptera: Hydrophilidae, type genus Sphaeridium Fabricius, 1775.

            Syn.: ... Sminthuridini Salmon, 1956
            Syn.: ... Sminthuridini Massoud, Z et Delamare Deboutteville, C, 1964:106




Genus Sphaeridia Linnaniemi, WM, 1912:248

  Syn.: ... Sminthurus Krausbauer, 1898 partim
  Syn.: ... Sminthurides Börner, 1901 partim
  Syn.: Sminthuridae Sminthurides Börner, C, 1901:6
  Syn.: ... Sphyrotheca Salmon, 1951
  Syn.: ... Sphyrotheca Massoud, Z et Delamare Deboutteville, C, 1964:101
  Syn.: Sminthurides Stenacidia Salmon, 1956
  Syn.: Sminthurides Stenacidia Massoud, Z et Delamare Deboutteville, C, 1964:106
  Syn.: ... Asphyrotheca Stach, 1956
  Syn.: ... Asphyrotheca Massoud, Z et Delamare Deboutteville, C, 1964:106
  Syn.: ... Indotheca Stach, 1956
  Syn.: ... Indotheca Massoud, Z et Delamare Deboutteville, C, 1964:101


  Syn.: Symphypleona Sphaerida(sic) Russell, DJ, 2008:126 ISS
Sphaeridia sp. ♀ from the USA
2019.09.06 © Lime, A.
Sphaeridia sp. ♀ from the USA
Mucro narrow elongate
Inner edge serrated, outer edge smooth
2019.09.06 © Lime, A.

  Syn.: Sminthurididae Sphaerida(sic) Palacios-Vargas, JG, Castaño-Meneses, G et Valladares, L, 2021:155 sp ISS
Sphaeridia sp. ♀ from Taiwan
Preparing to moult
2021.11.09 © Cheng, H.-J.
Sphaeridia sp. ♀ from Taiwan
2021.11.10 © Cheng, H.-J.
Sphaeridia sp. ♀ from Taiwan
2021.11.10 © Cheng, H.-J.
Sphaeridia sp. ♂ from Taiwan
Grasping antennae
2021.11.11 © Cheng, H.-J.
Sphaeridia pumilis ♂
Clasping mechanism
After da Silva Medeiros G & al, 2022 Fig.1A
Modified after Betsch J-M, 1980 Fig.46A
Sphaeridia sp. ♀ from Taiwan
2022.01.15 © Cheng, H.-J.
Sphaeridia sp. juv. from Taiwan
Tripod grooming
2022.01.18 © Cheng, H.-J.
Sphaeridia sp. ♂ & ♀ from Taiwan
♀ rejects ♂
2022.01.19 © Cheng, H.-J.
Sphaeridia sp. ♂ from Taiwan
2022.02.24 © Cheng, H.-J.
Sphaeridia sp. ♀ from Taiwan
Freshly deposited egg covered in excrements
2022.04.25 © Cheng, H.-J.
Sphaeridia sp. ♀ & ♂ from Taiwan
Mating
♂ deposits spermdrop on gonopore of ♀
2022.04.27 © Cheng, H.-J.



Species sp.1 Janssens F sensu Ospina-Sanchez, Soto-Adames & Gonzalez, 2020

            Syn.: Sphaeridia sp.nov.1 Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:24


Species aserrata Mari Mutt, JA, 1987:374

Sphaeridia aserrata ♂ from Colombia
After Mari Mutt JA, 1987 Fig.42
Sphaeridia aserrata ♀ from Colombia
After Mari Mutt JA, 1987 Fig.45

Species asiatica Rusek, 1971

Sphaeridia asiatica ♀ from Taiwan
2022.01.05 © Cheng, H.-J.
Sphaeridia asiatica juv. from Taiwan
2022.01.06 © Cheng, H.-J.
Sphaeridia asiatica ♂ & ♀ from Taiwan
Courtship
2022.01.06 © Cheng, H.-J.
Sphaeridia asiatica ♂ & ♀ from Taiwan
Courtship
♀ streaks ant.4 of ♂
2021.11.09 © Cheng, H.-J.
Sphaeridia asiatica ♀ from Taiwan
2022.01.13 © Cheng, H.-J.

Species aspinosa Bretfeld & Trinklein, 2000

Species betschi Arlé, 1984

Species biclava Bretfeld & Trinklein, 2000

Species biniserrata (Salmon, 1951) Massoud, Z & Delamare Deboutteville, C, 1964:101

Sphaeridia biniserrata juv. from India
After Bano R, 2016 p.64

Species bivirgata Bretfeld, 2002

Species boettgeri Bretfeld & Gauer, 1994

Species brevipila (Murphy, 1960) Murphy, 1966

Species cardosi Arlé, 1984

Species carioca Arlé, 1984

Species catapulta Bretfeld & Gauer, 1994

Species cerastes Bretfeld & Gauer, 1994

Species chisacae Bretfeld & Gauer, 1994

Species clara Bretfeld & Gauer, 1994

Species cornuta Murphy, 1966

Species coronata Bretfeld & Gauer, 1994

Species decemdigitata Bretfeld & Schulz, 2012

Species delamarei Bretfeld, 1997

Species denisi Massoud, Z et Delamare Deboutteville, C, 1964:95

Sphaeridia denisi from Peru
After Massoud Z & Delamare Deboutteville C, 1964 Fig.10B
             Syn.: Sphaeridia Denisi(sic) Massoud, Z et Delamare Deboutteville, C, 1964:113 ISS


Species duckei Bretfeld, 2002

Species fernandoi Murphy, 1966

Species fibulifera Bretfeld & Gauer, 1994

Species fluminensis Arlé, 1984

Species foliata Bretfeld, 1997

Species franklinae Bretfeld & Gauer, 1994

Species furcata Dunger, W et Bretfeld, G, 1989:7

Species gladiolifer Delamare Deboutteville, C et Massoud, Z, 1964:58

Species heloisae Arlé, 1984

Sphaeridia heloisea from Peru
2009.01-03.dd © Contreras-Valdez, A.

             Syn.: Sphaeridia heloisea(sic) Ernesto, MV, Liberal, CN, Ferreira, AS, Alves, ACF, Zeppelini, D, Martins, CF, Pereira-Colavite, A, Creao-Duarte, AJ, Vasconcellos, A, 2018:5 ISS


Species indica Prabhoo, 1971

Species inflata Bretfeld, 1997

Species irmleri Bretfeld & Gauer, 1994

Species jaliensis Bretfeld, 1997

Species leutrensis Dunger, W et Bretfeld, G, 1989:3

Species lobata Bretfeld & Gauer, 1994

Species mandibulata Bretfeld & Gauer, 1994

Species martii Bretfeld & Gauer, 1994

Species massoudi Murphy, 1966

Species minima (Schött, H, 1893:7) Stach, J, 1956:48

             Syn.: Sphaeridia minimus(sic) Massoud, Z et Delamare Deboutteville, C, 1964:112 ISS
             Syn.: Sphaeridia minimus(sic) Massoud, Z et Delamare Deboutteville, C, 1964:113 ISS
             Syn.: Sphaeridia minimus(sic) Massoud, Z et Delamare Deboutteville, C, 1964:114 ISS

Species murphyi Yosii, 1966

Species neopumilis Bretfeld & Gauer, 1994

Species obtusa Murphy, 1966

Species panguanae Bretfeld & Schulz, 2012

Species paroara Arlé, 1984

Species peruensis Bretfeld & Schulz, 2012

Species pilleata Bretfeld & Gauer, 1994

Species pippetti Murphy, 1966

Species posterospina Murphy, 1966

Species proxima Murphy, 1960

Species pumilis (Krausbauer, 1898 tt) Agrell, 1934, B

            Syn.: Sminthurus minimus Schtscherbakow, 1898 =?
            Syn.: Sminthurides (Sphaeridia) minimus Paclt, J, 1961:222 =?
            Syn.: Sminthurides globosus Axelson, 1902
            Syn.: Sphaeridia serrata Denis, 1948 =?
            Syn.: Sphaeridia serrata Stach, J, 1956:53 =?
            Syn.: Sphaeridia serrata Jeannenot, 1959 =?
            Syn.: Sphaeridia serrata Massoud, Z et Delamare Deboutteville, C, 1964:109 =?
            Syn.: Sphaeridia tunicata Stach, J, 1956:48
            Syn.: Sphaeridia tunicata Jeannenot, 1959 =?
            Syn.: Sphaeridia tunicata Massoud, Z et Delamare Deboutteville, C, 1964:109
            Syn.: Sminthurides (Stenacidia) violacea Salmon, 1956
            Syn.: Sphaeridia pumilis f. spinosa Bretfeld, G, 1995:18


             Syn.: Sminthurides pumilio(sic) Börner, 1901 ISS
             Syn.: Sminthurides pumilio(sic) Axelson, 1905 ISS
             Syn.: Sminthurides (Sphaeridia) pumilio(sic) Linnaniemi, 1912 ISS
             Syn.: Sminthurides (Sphaeridia) pumilio(sic) Handschin, E in Dahl, F, 1929:107 ISS
             Syn.: Sminthurides (Sphaeridia) pumilio(sic) Denis, 1931 ISS
             Syn.: Sminthurides (Sphaeridia) pumilio(sic) Kseneman, M, 1932:22 ISS
             Syn.: Sminthurides (Sphaeridia) pumilio(sic) Denis, J-R, 1933:272 ISS
             Syn.: Sminthurides pumilio(sic) Denis, J-R, 1933:320 ISS
             Syn.: Sphaeridia pumilio(sic) Agrell, 1934 ISS
Sminthurides (Sphaeridia) pumilis ♀ from the USA
After Maynard EA, 1951 Plate.XXII Fig.372

             Syn.: Sphaeridia pumilio(sic) Travé, J, Gadea, E et Delamare Deboutteville, C, 1954:211 ISS
             Syn.: Sminthurides pumilio(sic) Obenberger, 1955 ISS
             Syn.: Sphaeridia pumilio(sic) Delamare Deboutteville, C et Bassot, J-M, 1957:77 ISS
             Syn.: Sphaeridia pumilio(sic) Delamare Deboutteville, C et Bassot, J-M, 1957:84 ISS
             Syn.: Sphaeridia pumilio(sic) Delamare Deboutteville & Massoud, 1963 ISS
Sphaeridia pumilis ♀ from Belgium
Premating grooming
After Blancquart J-P & Mertens J, 1977 Fig.2
Sphaeridia pumilis ♂ & ♀ from Belgium
Premating contact
After Blancquart J-P & Mertens J, 1977 Fig.3
Sphaeridia pumilis ♂ & ♀ from Belgium
Production of sperm droplet
After Blancquart J-P & Mertens J, 1977 Fig.4
Sphaeridia pumilis ♂ & ♀ from Belgium
Postmating posture
After Blancquart J-P & Mertens J, 1977 Fig.5
Sminthurides (Sphaeridia) pumilis from the USA
After Krausbauer, 1898 in Christiansen K & Bellinger P, 1998 Fig.884A
Neozygites sp. from Denmark. Bar=10µm
Conidia infecting Sphaeridia pumilis
After Dromph & al, 2001 Fig.3
Sphaeridia pumilis from the UK
Male antenna
2004 © Hopkin, S.P.
Sphaeridia pumilis from France
2007.08.13 © Lebeaux, P.
Sphaeridia pumilis from Belgium
2007.12.18
© Salehian, S., Beladjal, L. & Mertens, J.
Sphaeridia pumilis female from Belgium
2007.12.18
© Salehian, S., Beladjal, L. & Mertens, J.
Sphaeridia pumilis juv. from the UK
2012.06.04 © Murray, A.
 
Sphaeridia pumilis ♂ from the UK
2012.06.04 © Murray, A.
 
Sphaeridia pumilis ♂ from the UK
Male clasping antenna
2012.06.04 © Murray, A.
Sphaeridia pumilis ♀ from the UK
2012.06.07 © Murray, A.
 
Sphaeridia pumilis ♀ from the UK
2012.06.07 © Murray, A.
Sphaeridia pumilis 1st instar from the UK
2012.06.07 © Murray, A.
Sphaeridia pumilis juv. from the UK
2012.06.07 © Murray, A.
Sphaeridia pumilis juv. from the UK
2012.06.11 © Murray, A.
Sphaeridia pumilis ♀ from the UK
2012.06.12 © Murray, A.
Sphaeridia pumilis ♂ from the UK
With clasping antennae
2012.07.08 © Nurcombe, E.
Sphaeridia pumilis ♂ from the UK
Male grasping antenna
2012.08.19 © Murray, A.
Sphaeridia pumilis ♀ from the UK
2012.08.24 © Nurcombe, E.
Sphaeridia pumilis ♀ from Holland
2012.10.04 © van Duinen, J.
Sphaeridia pumilis ♀ from Portugal
2012.10.13 © Murray, A.
Sphaeridia pumilis ♀ from the UK
2012.10.15 © Nurcombe, E.
Sphaeridia pumilis ♀ from the UK
2012.12.08 © Murray, A.
Sphaeridia pumilis ♀ from the UK
2013.01.13 © Murray, A.
Sphaeridia pumilis ♀ from the UK
2013.09.24 © Nurcombe, E.
Sphaeridia pumilis ♀ from Holland
2013.10.23 © van Duinen, J.
Sphaeridia pumilis ♂ from Holland
With clasping antennae
2013.10.23 © van Duinen, J.
Sphaeridia pumilis ♀ from the UK
2013.11.09 © Murray, A.
Sphaeridia pumilis ♀ from Holland
2014.10.03 © van Duinen, J.
Sphaeridia pumilis ♀ from Holland
2014.11.07 © van Duinen, J.
Sphaeridia pumilis ♀ & ♂ from Holland
End of copulation
2014.11.11 © van Duinen, J.
Sphaeridia pumilis ♀ from Germany
2014.11.15 © Beck, K.
Sphaeridia pumilis ♂ from Belgium
Simple clasping mechanism with 2 straight spines (*)
2017.07.20 © Huskens, M.-L.
Sphaeridia pumilis ♂ from France
2017.11.18 © Tiky, S.
Sphaeridia pumilis ♀ from Belgium
2018.06.30 © Huskens, M.-L.
Sphaeridia pumilis ♀ from Belgium
2018.06.30 © Huskens, M.-L.
Sphaeridia pumilis ♀ from Belgium
2018.06.30 © Huskens, M.-L.
Sphaeridia pumilis ♀ & ♂ from Holland
End of mating
2018.08.15 © Wieringa, P.
Sphaeridia pumilis ♂ from France
2018.11.24 © Tiky, S.
Sphaeridia pumilis ♀ from France
2020.05.07 © Reinach, Br.
Sphaeridia pumilis ♀ from France
Antenna unmodified
2020.05.07 © Reinach, Br.
Sphaeridia pumilis ♂ from Belgium
2020.09.08 © Huskens, M.-L.
Sphaeridia pumilis ♀ from Australia
2021.03.28 © Hobern, D.
Sphaeridia pumilis ♀ from Belgium
Egg production
2021.04.19 © Huskens, M.-L.
Sphaeridia pumilis ♀ from Belgium
Egg deposition
2021.04.19 © Huskens, M.-L.
Sphaeridia pumilis ♀ from Belgium
Egg covered in faeces
2021.04.19 © Huskens, M.-L.
Sphaeridia pumilis ♂ & ♀ from Belgium
Begin of mating
2021.05.01 © Huskens, M.-L.
Sphaeridia pumilis ♂ & ♀ from Belgium
♀ walks over ♂
While ♂ produces sperm droplet
2021.05.01 © Huskens, M.-L.
Sphaeridia pumilis ♂ & ♀ from Belgium
♂ deposits sperm droplet directly on gonopore of ♀
2021.05.01 © Huskens, M.-L.
Sphaeridia pumilis ♀ & ♂ from Belgium
End of mating
2021.05.01 © Huskens, M.-L.
Sphaeridia pumilis ♀ from Belgium
♀ scares off ♂ after mating
2021.05.01 © Huskens, M.-L.
Sphaeridia pumilis ♂ from Belgium
Antenna clasping mechanism (*)
2022.02.12 © Huskens, M.-L.
Sphaeridia pumilis ♂ from Belgium
Antenna clasping mechanism
2022.11.09 © Huskens, M.-L.
Sphaeridia pumilis ♀ from the USA
2022.12.08 © G, S.



Species robusta Bretfeld & Gauer, 1994

Species salmoni Massoud, Z et Delamare Deboutteville, C, 1964:106

            Syn.: Sminthurides (Stenacidia) violacea Salmon, 1956
            Syn.: Sminthurides (Stenacidia) violacea Massoud, Z et Delamare Deboutteville, C, 1964:106


             Syn.: Sphaeridia Salmoni(sic) Massoud, Z et Delamare Deboutteville, C, 1964:113 ISS

Species schalleri Massoud, Z et Delamare Deboutteville, C, 1964:85

             Syn.: Sphaeridia Schalleri(sic) Massoud, Z et Delamare Deboutteville, C, 1964:113 ISS

Species serrata (Folsom & Mills, 1938) Stach, J, 1956:48

             Syn.: Sphaeridia serratus(sic) Massoud, Z et Delamare Deboutteville, C, 1964:113 ISS
             Syn.: Sminthurus (Sphaeridia) serrata(sic) Massoud, Z et Delamare Deboutteville, C, 1964:115 ISS
             Syn.: Sphaeridia serratus(sic) Miranda Rangel & Palacios-Vargas, 1992 ISS
Sphaeridia serrata from the USA
2006 © Maddison, D.R.
Sphaeridia serrata ♀ & ♂ from the USA
2009 © Haney, J.
Sphaeridia serrata ♀ from the USA
2012 © Steffen, J. & al.



Species sphaera (Salmon, JT, 1946:58) Massoud, Z & Delamare Deboutteville, C, 1964:106

Species spinifer (da Gama, 1964) Yosii, 1966

Species spinifrons Murphy, 1966

Species spira Bretfeld & Gauer, 1994

Species squamifera Bretfeld & Gauer, 1994

Species sturmi Bretfeld & Gauer, 1994

Species superstes Bretfeld in Bretfeld, Poliakov & Broza, 2000

Species torifera Bretfeld & Schulz, 2012

Species tropica Bretfeld & Schulz, 2012

Species tschirnhausi Bretfeld & Schulz, 2012

Species tunicata Yosii, 1954

Species vampyra Bretfeld & Schulz, 2012

Species vanharteni Bretfeld, 2000

Species winteri Massoud, Z et Delamare Deboutteville, C, 1964:92

             Syn.: Sphaeridia Winteri(sic) Massoud, Z et Delamare Deboutteville, C, 1964:113 ISS

Species zaheri Yosii, 1966


Genus Debouttevillea Murphy, DH, 1965:31

  Syn.: Sminthuridinae Deboutevillea(sic) Murphy, DH, 1965:31 IOS
Debouttevillea marina ♂ from Singapore
Clasping mechanism
After da Silva Medeiros G & al, 2022 Fig.1D
Modified after Betsch J-M, 1980 Fig.46J



Species marina Murphy, DH, 1965:31, t.t.


Genus Denisiella Folsom & Mills, 1938 key  key

  Syn.: Sminthurididae Spinosellina Betsch, J-M, 1980:124 partim = 1st instar


Denisiella sp. ♂ from Madagascar
Clasping mechanism
After da Silva Medeiros G & al, 2022 Fig.1C
Modified after Betsch J-M, 1980 Fig.46G

Species sp.1 Christiansen & Bellinger, 1981

Species sp.2 Bellini, BC et Zeppelini, D, 2004:588

Species betschi Palacios-Vargas, JG, Soares Ferreira, A et Zeppelini, D, 2018:118

Denisiella betschi from Brazil
Frontal cephalic chaetotaxy
After Palacios-Vargas JG & al, 2018 Fig.20

Species bretfeldi Schulz, H-J et van Harten, A, 2013:15

Denisiella bretfeldi ♀ from the UAE
After Schulz H-J & van Harten A, 2013 Pl.5
Denisiella bretfeldi ♂ from the UAE
After Schulz H-J & van Harten A, 2013 Pl.6

Species caatingae Palacios-Vargas, JG, Soares Ferreira, A et Zeppelini, D, 2018:123

Denisiella caatingae from Brazil
Frontal cephalic chaetotaxy
With nasal organ
After Palacios-Vargas JG & al, 2018 Fig.33
             Syn.: Denisiella caatinagae(sic) Palacios-Vargas, JG, Soares Ferreira, A et Zeppelini, D, 2018:127 ISS


Species colombiana Ospina, M et Palacios-Vargas, JG, 2009:63

Species diomedesi Palacios-Vargas, JG, 2007:64

Species lithophila Snider, RJ, 1988:124

             Syn.: Sminthurides lithophila(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS
Sminthurides (Denisiella) lithophila ♀ from the USA
After Snider, 1988 in Christiansen K & Bellinger P, 1998 Fig.886.1A
Sminthurides (Denisiella) lithophila ♀ from the USA
After Snider, 1988 in Christiansen K & Bellinger P, 1998 Fig.886.1B
Sminthurides (Denisiella) lithophila ♂ from the USA
After Snider, 1988 in Christiansen K & Bellinger P, 1998 Fig.886.1C
Sminthurides (Denisiella) lithophila ♂ from the USA
After Snider, 1988 in Christiansen K & Bellinger P, 1998 Fig.886.1D



Species maesorum Palacios-Vargas, JG, 1995:28

Species nayarita Palacios-Vargas, JG et Bernava, V, 1999:447

Species ramosa (Folsom, JW, 1932:72) Stach, J, 1956:21

Species rhizophorae Palacios-Vargas, JG, Soares Ferreira, A et Zeppelini, D, 2018:113

Denisiella rhizophorae from Brazil
Frontal cephalic chaetotaxy
After Palacios-Vargas JG & al, 2018 Fig.8

Species seurati (Denis, 1925) Stach, J, 1956:11, t.t.

Species serroseta (Börner, C in Schultze, 1908:58) Stach, J, 1956:56

            Syn.: Sminthurides (Stenacidia) serroseta(sic) Börner, C in Schultze, 1908:58 IOS

            Syn.: Sminthurides distilacinia(sic) Crampton, 1922 IOS


             Syn.: Sminthurides (Stenacidia) serroseta(sic) Börner in Voeltzkowi, 1907 ISS
             Syn.: Sminthurides (Denisiella) serroseta(sic) Folsom & Mills, 1938 ISS
             Syn.: Sminthurides (Stenacidia) serrosetosa(sic) Stach, J, 1956:56 ISS
             Syn.: Sminthurides serrosetosa(sic) Stach, J, 1956:56 ISS
             Syn.: Sminthurides (Denisiella) serrosetosa(sic) Stach, J, 1956:56 ISS
             Syn.: Denisiella serrosetosa(sic) Stach, J, 1956:56 ISS
             Syn.: Sminthurides (Denisiella) serroseta(sic) Paclt, 1959 ISS
             Syn.: Sminthurides (Denisiella) serroseta(sic) Greenslade, P, 1997-:www0312 ISS
Denisiella serroseta ♀ from the UAE
After Schulz H-J & van Harten A, 2013 Pl.7

             Syn.: Sminthurides (Stenacidia) serroseta(sic) Thibaud, J-M, 2013:240 ISS




Species sexpinnata (Denis, 1931) Stach, J, 1956:21

             Syn.: Denisiella sexpinnatus(sic) Snider, RJ, 1988:128 ISS
             Syn.: Denisiella sexpinnatus(sic) Palacios-Vargas, JG, 1995:33 ISS
             Syn.: Sminthurides (Denisiella) sexspinnatus(sic) Skidmore, RE, 1995:60 ISS
             Syn.: Denisiella sexpinnatus(sic) Palacios-Vargas, JG et Bernava, V, 1999:447 ISS
             Syn.: Denisiella sexpinnatus(sic) Ospina, M et Palacios-Vargas, JG, 2009:67 ISS

Tribus Sminthuridini Börner, 1913 sensu Janssens F, 2022

Remark by Janssens, F. 2022.11.13: Based on Betsch (1980:125) and Medeiros & al (2022:7,19), tentatively we define Sminthuridini as genera with the distal tibiotarsus III organ present.



Genus Sminthurides Börner, 1900 key  key

  Syn.: Podurides(sic) Sminthurus Reuter, OM, 1883:3
  Syn.: Sminthurus Pilosi Schött, H, 1891:11 partim
  Syn.: Sminthurides(sic) Prosminthurus Willem, V, 1900:55
  Syn.: Sminthurides Spinosella Palissa, A, 1961:356
  Syn.: Sminthuridae Spinosella Greenslade, P in Houston, WWK, 1994:130
  Syn.: Sminthurides Spinosellina Palissa, A in Brohmer, P, Ehrmann, P et Ulmer, G, 1964:240
  Syn.: Sminthurididae Spinosellina Betsch, J-M, 1980:124 partim = 1st instar
Remark by Betsch, J.-M. 1980:124: [Spinosellina] s'agit exclusivement du premier stade des genres Sminthurides, Jeannenotia, Denisiella.

  Syn.: Sminthurididae Spinosellina Betsch, J-M, 1980:125
  Syn.: Sminthurididae Spinosellina Pomorski, RJ et Skarzynski, D, 1992:71
  Syn.: Sminthuridae Spinosellina Greenslade, P in Houston, WWK, 1994:130
  Syn.: Sminthuridida(sic) Spinosellina Bretfeld, G, 1999:34 partim
  Syn.: Sminthurididae Spinosellina Palissa, A in Schwoerbel, J et Zwick, P, 2000:120 = juv
  Syn.: Sminthurides (Spinosella) dentospinalis Palissa, A, 1961:354
Sminthurides (Spinosella) dentospinalis from Germany
Dentes with 2 basal spines; ventral aspect
Modified after Palissa A, 1961 Fig.5
Remark by Palissa, A. 1964:240: Lawrence, 1963 halt dentospinalis für ein Jugendform einer anderen Sminthurides-Art.

  Syn.: Spinosellina dentospinalis Massoud, Z et Betsch, J-M, 1972:73 = 1st instar Sminthurididae
  Syn.: Spinosellina dentospinalis Fjellberg, 1974 = juv
  Syn.: Sminthurides (Spinosella) dentospinalis Potapov, 1995
  Syn.: Sminthurides (Spinosellina) dentospinalis Palissa, A in Schwoerbel, J et Zwick, P, 2000:120 = juv
Remark by Palissa, A. 2000:120: S. dentospinalis ist ein Jugendstadium einer anderen Sminthurides-Art (Fjellberg, 1974).




Remark by ICZN. 2007:759: [Sminthurides] Placed on the Official List of Names in Zoology. Opinion 291.

Sminthurides penicillifer var. incomptus ♀ from Germany
Bothriotricha ABC in obtuse angle that opens anteriorly
2014.09.17 © Beck, K.

  Syn.: Sminthuridae Sminthurus(sic) Glime, JM, 2015:5 lapsus ISS
Sminthurides sp. from India
After Bano R, 2016 p.63
Det.: Janssens F
Sminthurides sp. ♂ & ♀ from Japan
Courtship
While female is moulting
2020.03.23 © Hara, Y.

  Syn.: Sminthurididae Siminthurides(sic) Ferreira, AS, Arango, A et Palacios-Vargas, JG, 2021:16 ISS
Sminthurides sp. from Taiwan
Grooming
2021.07.30 © Cheng, H.-J.
Sminthurides aquaticus ♂
Clasping mechanism
After da Silva Medeiros G & al, 2022 Fig.1F
Modified after Betsch J-M, 1980 Fig.46D

  Syn.: Symphypleona Sminturides(sic) Sys, S, Weißbach, S, Jakob, L, Gerber, S, et Schneider, C, 2022:2736 ISS


Species sp.1 Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:895

Species sp.2 Janssens F & Dell T, 2008

Sminthurides nr pseudassimilis from Australia
2002.11.05 © Dell T.
Sminthurides nr pseudassimilis from Australia
Furca with lamellate mucrones
2002.11.05 © Dell T.
Sminthurides nr pseudassimilis from Australia
Female specimen
Antenna with annulated 4th antennomere
2002.11.05 © Dell T.

Species sp.3 Soto-Adames, FN et Taylor, SJ, 2013:163

Species sp.5 Janssens F & Tiky S, 2018

Sminthurides sp.5 ♂ from France
2018.02.25 © Tiky, S.
Sminthurides sp.5 ♂ from France
2018.02.25 © Tiky, S.
Sminthurides sp.5 ♀ from France
2018.02.25 © Tiky, S.
Sminthurides ?sp.5? ♀ from the UK
2018.04.28 © Robson, S.

Species sp.6 Janssens F sensu Ferreiro AS & al, 2021

            Syn.: Sminthurides sp.undescribed Ferreira, AS, Arango, A et Palacios-Vargas, JG, 2021:1


Species sp.7 Janssens F sensu Palacios-Vargas JG, 2022





Species-group aquaticus Massoud Z & Betsch J-M, 1972:74
             Syn.: Sminthurides Aquaticus(sic) Ferreira, AS, Arango, A et Palacios-Vargas, JG, 2021:2,gr ISS

Species aquaticus (Bourlet, 1842 tt) Börner, C, 1901:96, B

            Syn.: Sminthurus apicalis Reuter, 1880
Sminthurus apicalis ♀ from Scandinavia
After Reuter OM, 1883 Fig.1
Sminthurus apicalis ♀ from Scandinavia
After Reuter OM, 1883 Fig.2
Sminthurus apicalis ♀ from Scandinavia
Grooming with drop of saliva
After Reuter OM, 1883 Fig.3
Sminthurus apicalis ♂ from Scandinavia
After Reuter OM, 1883 Fig.4
Sminthurus apicalis from Scandinavia
Male grasping antenna
After Reuter OM, 1883 Fig.9


             Syn.: Sminthurus apiculis(sic) Schött, H, 1893:36 ISS
Sminthurus apicalis from Finnland
Mucro with 3 lamellae: inner serrate, outer + median smooth
After Levander KM, 1894:10

            Syn.: Sminthurides aquaticus f. principalis Schött
            Syn.: Sminthurus aquaticus var. levanderi Reuter OM in Schött, H, 1893:37
            Syn.: Sminthurus apicalis var. levanderi Reuter, 1890 u
            Syn.: Sminthurus apicalis var. Levanderi(sic) Reuter, OM, 1892:232 nn IOS
            Syn.: Sminthurus aquaticus var. Levanderi(sic) Reuter, OM, 1895:4 IOS
            Syn.: Sminthurus aquaticus var. Levanderi(sic) Reuter, OM, 1895:6 IOS
            Syn.: Sminthurus aquaticus var. Levanderi(sic) Reuter, OM, 1895:13 IOS
Sminthurides f. levanderi ♀ from the USA
After Maynard EA PlateXXII Fig.369
Sminthurides f. levanderi ♀ from the USA
After Maynard EA PlateXXII Fig.368

             Syn.: Sminthurides aquaticus var. levanderis(sic) Isa, 1963 ISS
Sminthurides aquaticus var. levanderi from France
2008.03.17 © Lebeaux, P.
 
Sminthurides aquaticus var. levanderi from France
2008.03.17 © Lebeaux, P.
 
Sminthurides aquaticus var. levanderi from France, courtship
2008.03.18 © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France
2008.04.02 © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France, courtship
2008.04.06 © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France
2008.04.06 © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France, courtship
2008.04.06 © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France
2009.09.dd © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France
2009.09.dd © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France
2009.09.dd © Lebeaux, P.
Sminthurides aquaticus var. levanderi ♂ from France
With highly modified grasping antennae
2009.09.dd © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France
Courtship
2009.09.dd © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France
Predation by spider
2009.09.13 © Lebeaux, P.
Sminthurides aquaticus var. levanderi ♀ from Holland
2011.04.dd © van Duinen, J.
Sminthurides aquaticus var. levanderi ♀ from the UK
2012.09.29 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
2012.10.10 © Barton, T.
Sminthurides aquaticus var. levanderi ♂ & ♀ from the UK
2012.11.19 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
2012.12.23 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the USA
Feeding instar with dark dorsum due to gut contents
2013.02.09 © Fleming, K.
Sminthurides aquaticus var. levanderi ♀ from the UK
2013.04.23 © Murray, A.
Sminthurides aquaticus var. levanderi ♀ from the UK
2013.11.04 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
2014.10.20 © Barton, T.
Sminthurides aquaticus var. levanderi ♂ & ♀ juv. from the UK
Courtship
2014.10.20 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
Pale basal antennomeres
2014.10.20 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
Lamellate mucro
2014.10.20 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
Fighting females
2014.11.21 © Murray, A.
Sminthurides aquaticus var. levanderi from the UK
2014.11.26 © Murray, A.
Sminthurides aquaticus var. levanderi from the UK
Black foot and lamellate mucro
2014.11.26 © Murray, A.
Sminthurides aquaticus var. levanderi ♂ & ♀ from the UK
Courtship ritual
2014.12.01 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
Aggressive? encounter of two females
2014.12.03 © Ardron,P.A.
Sminthurides aquaticus var. levanderi ♀ from the UK
Aggressive? encounter of two females
2014.12.03 © Ardron,P.A.
Sminthurides aquaticus var. levanderi ♀ from the UK
2014.12.08 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
Proleg: black foot; short empodium
2014.12.10 © Ardron,P.A.
Sminthurides aquaticus var. levanderi ♀ from the UK
Ready to moult
2014.12.10 © Ardron,P.A.
Sminthurides aquaticus var. levanderi ♀ from the UK
2015.01.11 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
2015.01.11 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
2015.01.11 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
2015.02.15 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
2015.04.26 © Ardron, P.A.
Sminthurides aquaticus var. levanderi ♂ from the UK
2015.04.26 © Ardron, P.A.
Sminthurides aquaticus var. levanderi ♀ from the UK
Guarding eggs on Lemna minor
2015.05.09 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from France
2016.11.27 © Garcelon, P.
Sminthurides aquaticus var. levanderi ♀ from France
2017.02.28 © Picard, J.
Sminthurides aquaticus var. levanderi ♀ from France
2017.02.28 © Picard, J.
Sminthurides aquaticus var. levanderi ♂ from France
2017.02.28 © Picard, J.
Sminthurides aquaticus var. levanderi ♀ from France
Mucro with strong anterior lamella (al), undulating posterior outer lamella (pol) and mucronal seta (ms)
2017.04.01 © Picard, J.
Sminthurides aquaticus var. levanderi ♀ juv. from Belgium
2017.06.03 © Huskens, M.-L.
Sminthurides aquaticus var. levanderi ♀ from France
2017.10.22 © Picard, J.
Sminthurides aquaticus var. levanderi ♀ from France
Black foot with long unguis
2017.10.22 © Picard, J.
Sminthurides aquaticus var. levanderi ♀ from France
Mucro with undulate outer lamella
2017.10.22 © Picard, J.
Sminthurides aquaticus var. levanderi ♂ & ♀ from France
Courtship
2018.04.02 © Tiky, S.
Sminthurides aquaticus var. levanderi ♀ from the UK
2018.04.09 © Phillips, E.
Sminthurides aquaticus var. levanderi ♀ & ♂ from the USA
Courtship
2019.02.08 © Fleming, K.
Sminthurides aquaticus var. levanderi juv. from Belgium
Grooming on 3 legs
2019.08.30 © Huskens, M.-L.
Sminthurides aquaticus var. levanderi juv. from Belgium
2019.08.30 © Huskens, M.-L.
Sminthurides aquaticus var. levanderi juv. from Belgium
Ocelli A to H
2019.08.30 © Huskens, M.-L.
Sminthurides aquaticus var. levanderi ♀ from Belgium
2019.09.08 © Huskens, M.-L.
Sminthurides aquaticus var. levanderi ♀ from Belgium
Mucro with anterior lamella (al), posterior external lamella (pel) and mucronal seta (ms)
2019.09.08 © Huskens, M.-L.
Sminthurides aquaticus var. levanderi ♀ from Belgium
Frontal chaetotaxy
2019.09.08 © Huskens, M.-L.
Sminthurides aquaticus var. levanderi ♀ from France
Mucro with posterior inner lamella (pil), posterior outer lamella (pol) and anterior lamella (al)
2020.01.21 © Dejoux, P.
Sminthurides aquaticus var. levanderi ♂ & ♀ from the UK
Courtship
2021.05.02 © Phillips, E.
Sminthurides aquaticus var. levanderi ♂ & ♀ from the UK
Mucro with undulating posterior outer lamella
2021.05.02 © Phillips, E.


            Syn.: Sminthurus aquaticus var. viridulus Reuter OM in Schött, H, 1893:37
            Syn.: Sminthurus apicalis var. viridulus Reuter, 1890 u
            Syn.: Sminthurus apicalis var. viridulus Reuter, OM, 1892:231 nn
             Syn.: Sminthurides aquaticus var. viridula(sic) Bretfeld, G, 1999:38 ISS
Sminthurides aquaticus var. viridulus ♀ & ♂ from Germany
2014.11.05 © Gutekunst, V.
Sminthurides aquaticus var. viridulus ♀ from Germany
2014.11.05 © Gutekunst, V.
Sminthurides aquaticus var. viridulus ♀ from Germany
Long narrow unguis and needle-like unguiculus of 1st leg
2014.11.05 © Gutekunst, V.
Sminthurides aquaticus var. viridulus ♀ from Germany
2014.11.05 © Gutekunst, V.
Sminthurides aquaticus var. viridulus ♀ from Germany
Composed eye with 6 large ocelli and 2 distinctly smaller ocelli
2014.11.05 © Gutekunst, V.
Sminthurides aquaticus var. viridulus ♀ from Germany
2014.11.05 © Gutekunst, V.
Sminthurides aquaticus var. viridulus ♀ & ♂ from Germany
With embryo's in eggs
2014.11.05 © Gutekunst, V.
Sminthurides aquaticus var. viridulus from Germany
Embryo's in eggs embedded in epidermis of Lemna minor
2014.11.05 © Gutekunst, V.
Sminthurides aquaticus var. viridulus ♀ from Poland
2015.05.14 © Beck, K.


            Syn.: Sminthurus apicalis f. principalis Reuter, 1890
            Syn.: Smynthurus(sic) amicus Folsom, JW, 1896:447
Smynthurus(sic) amicus from the USA
Fore foot with long unguis and shorter unguiculus
After Folsom JW, 1896 Plate10 Fig.6
Smynthurus(sic) amicus from the USA
Hind foot with short unguis and longer unguiculus
After Folsom JW, 1896 Plate10 Fig.8
Smynthurus(sic) amicus from the USA
Mucro with serrate inner and smooth outer lamella
After Folsom JW, 1896 Plate10 Fig.7




             Syn.: Sminthurus aqvaticus(sic) Reuter, OM, 1883:3 ISS
Sminthurus aquaticus from Sweden
After Schött H, 1893 Taf.II Fig.22
Sminthurides aquaticus from Germany
Foot of proleg with short unguiculus
After Börner C, 1901 Fig.43a
Sminthurides aquaticus from Germany
Foot of mesoleg with long unguiculus
After Börner C, 1901 Fig.43b
Sminthurides aquaticus from Germany
Foot of metaleg with long unguiculus
and tibiotarsal organ
After Börner C, 1901 Fig.43c
Sminthurides aquaticus from Germany
After Heymons R & Heymons H, 1909 Fig.22
Sminthurus aquaticus from Germany
After Anonymous Fig.F
Sminthurides aquaticus from the UK
Broad lamellate mucro
2004 © Hopkin, S.P.
Sminthurides aquaticus from Sweden
2006 © Hall, K.
Sminthurides aquaticus from the UK
2007.03.04 © Cornwall, N.J.
Sminthurides aquaticus from the UK
2007.03.18 © Cornwall, N.J.
Sminthurides aquaticus from the UK
2007.03.18 © Cornwall, N.J.
Sminthurides aquaticus from the UK
2007.03.26 © Cornwall, N.J.
Sminthurides aquaticus from the UK
2007.04.01 © Jones, D.
Sminthurides aquaticus from Germany
2007.08.13 © Schmitt, R.
Sminthurides aquaticus from Germany
2007.08.14 © Schmitt, R.
Sminthurides aquaticus from the UK
2007.08.31 © Oakshott, A.
Sminthurides aquaticus from Germany
2007.10.12 © Schmitt, R.
Sminthurides aquaticus from Belgium
2008.04.22 © Vancoppenolle B.
Sminthurides aquaticus from the UK
2009.03.29 © Michaelson, J.
Sminthurides aquaticus from the UK
2009.05.10 © Robertson, A.
Sminthurides aquaticus from the UK
2009.05.10 © Robertson, A.
Sminthurides aquaticus from the UK
2009.05.10 © Robertson, A.
Sminthurides aquaticus from the UK
2009.05.17 © Robertson, A.
Embryo in transparant egg ready to hatch
Sminthurides aquaticus from the UK
2009.06.17 © Robertson, A.
Male with clasping antennae
Sminthurides aquaticus from the UK
2009.06.17 © Robertson, A.
Male with clasping antennae
Sminthurides aquaticus from the UK
2009.06.17 © Robertson, A.
Male advancing larger female
Sminthurides aquaticus from the UK
2009.09.30 © Robertson, A.
Bothriotricha A-D
Sminthurides aquaticus from the UK
2010.07.03 © Robertson, A.
Male clasped to female
Sminthurides aquaticus from the UK
2010.08.07 © Robertson, A.
Unsubdivided 4th antennal segment
Sminthurides aquaticus from the UK
Furca with broad lamellate mucrones
2012.04.12 © Peacock, H.
Sminthurides aquaticus from the UK
2012.05.01 © Murray, A.
Sminthurides aquaticus ♂ from the UK
2012.05.02 © Murray, A.
Sminthurides aquaticus juv. from the UK
2012.05.14 © Murray, A.
Sminthurides aquaticus from the UK
Eggs in 'nests'
2012.05.19 © Murray, A.
Sminthurides aquaticus from the UK
Embryo in transparant egg
2012.05.19 © Murray, A.
Sminthurides aquaticus from the UK
Embryo in transparant egg
2012.05.22 © Murray, A.
Sminthurides aquaticus ♂ & juv. ♀ from the UK
2012.05.28 © Murray, A.
Sminthurides aquaticus juv. from the UK
2012.06.04 © Murray, A.
Sminthurides aquaticus juv. ♀ & ♂ from the UK
2012.08.12 © Nurcombe, E.
Sminthurides aquaticus juv. & ♂ from the UK
Male grasps juvenile at wrong end...
2012.10.03 © Nurcombe, E.
Sminthurides aquaticus juv. ♀ from the UK
2013.02.11 © Nurcombe, E.
Sminthurides aquaticus ♂ & ♀ from the UK
Mucro with broad lamellae
2013.02.17 © Nurcombe, E.
Sminthurides aquaticus ♂ from Germany
Antennal clasping organ
2014.09.19 © Beck, K.
Sminthurides aquaticus ♀ from Germany
Ant.4 not subdivided
2014.10.05 © Kluck, A.
Sminthurides aquaticus ♂'s from Germany
Figthing?
2014.11.09 © Kluck, A.
Sminthurides aquaticus from the UK
2014.11.14 © Murray, A.
Sminthurides aquaticus ♀ from Germany
2015.10.31 © Kluck, A.
Sminthurides aquaticus ♂ from Germany
2015.11.15 © Kluck, A.
Sminthurides aquaticus ♀ & ♂ from Germany
Male grasping antenna
2015.11.15 © Kluck, A.
Sminthurides aquaticus ♀ from Germany
Antenna
2015.11.15 © Kluck, A.
Sminthurides aquaticus ♀ from Belgium
2016.07.10 © Huskens, M.-L.
Sminthurides aquaticus juv. ♂ from Belgium
With swollen ant.2-3
2016.07.12 © Huskens, M.-L.
Sminthurides aquaticus subadult ♂ from Belgium
With precursive grasping antenna
2016.07.12 © Huskens, M.-L.
Sminthurides aquaticus juv. ♀ & ♂ from Belgium
Courtship
2016.07.13 © Huskens, M.-L.
Sminthurides aquaticus ♀ from Belgium
Ant.4 not subdivided
2016.07.19 © Huskens, M.-L.
Sminthurides aquaticus ♀ from Belgium
Grooming with a drop of saliva
2016.07.25 © Huskens, M.-L.
Sminthurides aquaticus ♀ from Belgium
Chewing cup in epidermis of Lemna minor to deposit egg
2016.07.27 © Huskens, M.-L.
Sminthurides aquaticus ♀ from Belgium
Depositing egg in epidermal cup of Lemna minor
2016.07.27 © Huskens, M.-L.
Sminthurides aquaticus ♀ from Belgium
Covering egg with faecal pellet
2016.07.27 © Huskens, M.-L.
Sminthurides aquaticus from Belgium
Embryo's embedded in epidermis of Lemna minor
2016.07.27 © Huskens, M.-L.
Sminthurides aquaticus ♂ from the UK
Male clasping antenna
2016.08.19 © Phillips, E.
Sminthurides aquaticus ♀ from the UK
Cleaning dorsum with drop of saliva
2016.08.23 © Phillips, E.
Sminthurides aquaticus ♀ from Belgium
Balancing on 3 right legs
2016.08.25 © Huskens, M.-L.
Sminthurides aquaticus ♀ + ♂ from Belgium
Begin of courtship ritual
2016.09.22 © Huskens, M.-L.
Sminthurides aquaticus ♂ from Belgium
Clasping antenna
2016.09.22 © Huskens, M.-L.
Sminthurides aquaticus ♂ juv. from Belgium
First instar
2016.09.30 © Huskens, M.-L.
Sminthurides aquaticus ♀ from the UK
Bothriotricha A-D
2016.10.09 © Barton, T.
Sminthurides aquaticus ♀ from Belgium
Balancing on 3 right legs
2016.10.25 © Huskens, M.-L.
Sminthurides aquaticus ♀ from Germany
2017.05.25 © Jäschke, J.
Sminthurides aquaticus ♀ from Belgium
2017.06.20 © Huskens, M.-L.
Sminthurides aquaticus ♂ from Belgium
Metathoracic bladder-like vesicles
2017.07.23 © Huskens, M.-L.
Sminthurides aquaticus ♀♀ from Belgium
Ventral aspect. Fighting?
Note split labium
2018.05.22 © Huskens, M.-L.
Sminthurides aquaticus ♂ from Belgium
2018.07.13 © Huskens, M.-L.
Sminthurides aquaticus ♀ from Belgium
Proleg with unguiculus (u) as long as unguis (U)
2018.10.05 © Huskens, M.-L.
Sminthurides aquaticus ♀ & ♂ from the UK
2018.10.16 © Phillips, E.
Sminthurides aquaticus ♀ from Belgium
Interantennal spot X-shaped
2019.01.14 © Huskens, M.-L.
Sminthurides aquaticus ♀ from Belgium
Ant.4 not subdivided
2019.01.14 © Huskens, M.-L.
Sminthurides aquaticus ♀ juv. & ♂ from the UK
Courtship
Male clasping antenna
2019.09.17 © Smart, J.
Sminthurides aquaticus ♀ from Belgium
Posterior abdominal dense cover of short setae
2020.07.04 © Huskens, M.-L.
Sminthurides aquaticus ♀ from Belgium
Mucro with distinct lateral lamellae
2020.07.04 © Huskens, M.-L.
Sminthurides aquaticus ♂ from France
Furca with mucrones having lateral lamellae
2020.11.18 © Reinach, Br.

             Syn.: Sminthurides Aquaticus(sic) Ferreira, AS, Arango, A et Palacios-Vargas, JG, 2021:1,gr ISS
Sminthurides aquaticus ♀ from Belgium
Proleg with unguiculus (u) as long as unguis (U)
2021.08.15 © Huskens, M.-L.
Sminthurides aquaticus ♂ & subadult ♀ from the UK
Courtship with clasping antennae
2022.02.07 © Phillips, E.





Species aureolus Maynard, EA, 1951:213

Sminthurides aureolus ♀ from the USA
After Maynard EA PlateXXII Fig.370
Sminthurides aureolus ♂ from the USA
After Maynard EA PlateXXII Fig.371

Species cihuatlensis Ferreira, AS, Arango, A et Palacios-Vargas, JG, 2021:4

Sminthurides cihuatlensis from Mexico
Facial chaetotaxy
After Ferreiro AS & al, 2021 Fig.5

            Syn.: Sminthurides sp. Cutz Pool & al.2010
            Syn.: Sminthurides sp.undescribed Ferreira, AS, Arango, A et Palacios-Vargas, JG, 2021:1
            Syn.: Sminthurides sp. Ferreira, AS, Arango, A et Palacios-Vargas, JG, 2021:4



Species globocerus Folsom & Mills, 1938

Species hyogramme Pedigo, 1966

            Syn.: Sminthurus signatus Christiansen, K et Bellinger, P, 1998:1198 =?
            Syn.: Sminthurides krausbaueri Folsom & Mills, 1938
            Syn.: Sminthurides krausbaueri Hammer, 1953
            Syn.: Sminthurides krausbaueri Christiansen, K et Bellinger, P, 1998:1198


             Syn.: Sminthurides hygromae(sic) Palacios-Vargas, 1990 ISS
             Syn.: Sminthurides hyogramma(sic) Miranda Rangel & Palacios-Vargas, 1992 ISS
             Syn.: Sminthurides hypogramme(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS
Sminthurides (Sminthurides) hyogramme ♀ from the USA
After Pedigo, 1966 in Christiansen K & Bellinger P, 1998 Fig.893B
Sminthurides (Sminthurides) hyogramme ♀ from the USA
After Pedigo, 1966 in Christiansen K & Bellinger P, 1998 Fig.893A

             Syn.: Sminthurides hygramme(sic) Cruz-Leal, JI, Escalante-Jiménez, AL, Salgado-Ortiz, J et Palacios-Vargas, JG, 2013:1666 ISS
Sminthurides hyogramme ♀ from the USA
2018.05.29 © Caterino, M.
Sminthurides hyogramme ♀ from the USA
2018.06.04 © Caterino, M.
Sminthurides hyogramme ♀ from the USA
2018.08.15 © Caterino, M.
Sminthurides hyogramme ♂ from the USA
2020.06.08 © Caterino, M.

             Syn.: Sminthurides hyograme(sic) Ferreira, AS, Arango, A et Palacios-Vargas, JG, 2021:1 ISS
Sminthurides hyogramme ♀ from the USA
2021.09.23 © Caterino, M.



Species napostaensis Najt & Rapoport, 1965

Species occultus Mills, HB, 1934:91

Sminthurides (Sminthurides) occultus ♀ from the USA
After Maynard EA PlateXXII Fig.357
Sminthurides (Sminthurides) occultus ♂ from the USA
After Maynard EA PlateXXII Fig.358
Sminthurides (Sminthurides) occultus from Manitoba
After Folsom & Mills, 1938 in Christiansen K & Bellinger P, 1998 Fig.897A
Sminthurides occultus ♀ from the USA
2012 © Steffen, J. & al.

Species olivieri Delamare Deboutteville & Massoud, 1963:185

            Syn.: Sminthurides Olivieri(sic) Delamare Deboutteville & Massoud, 1963 IOS

Species rapoporti Delamare Deboutteville & Massoud, 1963:187

            Syn.: Sminthurides Rapoporti(sic) Delamare Deboutteville & Massoud, 1963 IOS

Species ringueleti Delamare Deboutteville & Massoud, 1963:182

            Syn.: Sminthurides Ringueleti(sic) Delamare Deboutteville & Massoud, 1963 IOS

Species terrestris Maynard, EA, 1951:213

Sminthurides terrestris from the USA
After Maynard EA, 1951 Pl.XXIII Fig.395
Sminthurides terrestris from the USA
After Maynard EA, 1951 Pl.XXIII Fig.396

             Syn.: Sminthurides terretris(sic) Ferreira, AS, Arango, A et Palacios-Vargas, JG, 2021:1 ISS

Species-group penicillifer Massoud Z & Betsch J-M, 1972:74
             Syn.: Sminthurides Penicillifer(sic) Ferreira, AS, Arango, A et Palacios-Vargas, JG, 2021:2,gr ISS


Species bifidus Mills, HB, 1934:90

            Syn.: Sminthurus penicillifer Guthrie, JE, 1903:55


Sminthurides (Sminthurides) bifidus ♀ from the USA
After Maynard EA PlateXXII Fig.366
Sminthurides (Sminthurides) bifidus ♀ from the USA
After Maynard EA PlateXXII Fig.367
Sminthurides bifidus ♀ from the USA
After Maynard EA PlateXXIV Fig.427

             Syn.: Sminthurides (Sminthurides) bifidis(sic) Richards, 1956 ISS
             Syn.: Sminthurides (Sminthurides) bifidis(sic) Richards, WR, 1968:19 ISS
             Syn.: Sminthurides bificus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS
Sminthurides bifidus ♂ & ♀ from Russia
From aquarium
2005.mm.dd © Potapov, M.
Sminthurides bifidus from Germany
Eye cluster with 7 eyes
2006 © Schulz, H.-J. & Schubert H.D.
Sminthurides bifidus from the USA
2008.01.16 © Mcmillan, A.S.
 
Sminthurides bifidus ♀ from the USA
2009.02.15 © Fleming, K.
Sminthurides bifidus from the USA
2009.05.02 © Zungri, D.
Sminthurides bifidus ♀ from the USA
2012.04.23 © Kouri, J.
Sminthurides bifidus from the USA
Male approaching female
2012.04.23 © Kouri, J.
Sminthurides bifidus from the USA
Male grasped juvenile female
2012.04.23 © Kouri, J.
Sminthurides bifidus from Germany
Habitus dorsal
2012.05.04 © Schulz, H.-J.
Sminthurides bifidus from Germany
Habitus lateral
2012.05.04 © Schulz, H.-J.
Sminthurides bifidus from Germany
Mucro SEM 400x
2012.05.04 © Schulz, H.-J.
Sminthurides bifidus from Germany
Foot complex of leg1 or leg2
2012.05.04 © Schulz, H.-J.
Sminthurides bifidus from the USA
2012.05.07 © Kouri, J.
Sminthurides bifidus ♂ from the USA
2012.05.07 © Kouri, J.
Sminthurides bifidus ♂ & ♀ from the USA
2012.05.07 © Kouri, J.
Sminthurides bifidus from the USA
2012.05.13 © Kouri, J.
Sminthurides bifidus ♀ from the USA
2012.07.11 © Janssens, F.
Leg. Kouri, J.
Sminthurides bifidus ♀ from the UK
2013.04.19 © Murray, A.
Sminthurides bifidus ♀ from the UK
2013.04.19 © Murray, A.
Sminthurides bifidus ♀ & ♂ from the UK
2013.04.19 © Murray, A.
Sminthurides bifidus ♀ from the UK
Composed eye with 6 large ocelli and 2 distinctly smaller ocelli
2013.04.23 © Murray, A.
Sminthurides bifidus ♂ from the UK
2013.04.23 © Murray, A.
Sminthurides bifidus ♀ from Germany
2015.05.23 © Gutekunst, V.
Sminthurides penicillifer ssp. bifidus ♀ from France from aquarium
2017.02.08 © Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ from France from aquarium
Ant.4 subdivided
2017.02.08 © Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ juv. & ♂ from France from aquarium
Courtship
2017.02.08 © Picard, J.
Sminthurides penicillifer ssp. bifidus ♂ from France from aquarium
With modified grasping antennae
2017.02.08 © Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ from France from aquarium
Albino form with reddish eyes
2017.03.01 © Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ from France from aquarium
2017.03.02 © Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ from France from aquarium
8 ocelli (C+D small)
2017.03.02 © Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ juv. & albino ♂ from France from aquarium
2017.03.02 © Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ from France from aquarium
Ant.4 with intersubdivisional rings
2017.04.11 © Janssens, F. Leg.: Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ from France from aquarium
Foot3 with bifid unguiculus
2017.04.21 © Janssens, F. Leg.: Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ from France from aquarium
Mucro with shortened outer lamella
2017.04.21 © Janssens, F. Leg.: Picard, J.
Sminthurides penicillifer ssp. bifidus ♂ from France from aquarium
Ant.2+3 clasping mechanism
2017.04.21 © Janssens, F. Leg.: Picard, J.
Sminthurides penicillifer ssp. bifidus ♂ & ♀ from Germany from aquarium
Courtship
2017.11.20 © Jäschke, J.
Sminthurides penicillifer ssp. bifidus ♀ from Portugal
From aquarium
2018.11.25 © Gomes, M.
Sminthurides penicillifer ssp. bifidus ♀ from Portugal from aquarium
With reddish eyes
2018.11.25 © Gomes, M.
Sminthurides penicillifer ssp. bifidus ♀ from Portugal from aquarium
With reddish eyes
2018.11.25 © Gomes, M.
Sminthurides penicillifer ssp. bifidus ♀ from Portugal from aquarium
With reddish eyes
2018.12.02 © Gomes, M.
Sminthurides penicillifer ssp. bifidus ♀ from Portugal from aquarium
With reddish eyes
2018.12.02 © Gomes, M.
Sminthurides penicillifer ssp. bifidus ♀ from Portugal from aquarium
With reddish eyes
2018.12.02 © Gomes, M.
Sminthurides penicillifer ssp. bifidus ♂ & ♀ from Portugal from aquarium
Courtship
2019.02.03 © Gomes, M.
Sminthurides penicillifer ssp. bifidus ♂ & ♀ from Portugal from aquarium
Courtship
2019.02.03 © Gomes, M.
Sminthurides penicillifer ssp. bifidus ♀ from Portugal from aquarium
With reddish eyes
2020.11.16 © Gomes, M.
Sminthurides penicillifer ssp. bifidus ♀ from Portugal from aquarium
With reddish eyes
2020.11.16 © Gomes, M.
Sminthurides penicillifer ssp. bifidus ♀ from Portugal from aquarium
With reddish eyes
2020.11.16 © Gomes, M.
Sminthurides penicillifer ssp. bifidus ♀ from Russia from aquarium
With pale eye background
2021.02.03 © Teslenko, I.
Sminthurides bifidus ♀ & ♂ from Taiwan
Courtship
2021.11.13 © Cheng, H.-J.
Sminthurides bifidus ♀ from Taiwan
2021.11.13 © Cheng, H.-J.
Sminthurides bifidus ♂ from Taiwan
With modified clasping antennae
2021.11.13 © Cheng, H.-J.
Sminthurides bifidus ♂ from Taiwan
Clasping mechanism
b1-4-3-5; 2 not visible
2021.11.13 © Cheng, H.-J.
Sminthurides bifidus ♀ from Taiwan
2021.11.13 © Cheng, H.-J.
Sminthurides bifidus ♂ & ♀ from Taiwan
Courtship
2021.11.13 © Cheng, H.-J.
Sminthurides bifidus ♀ from Holland
Ant.4 subdivision with basal and apical ring (*)
2021.11.16 © van Veen, K.
Sminthurides bifidus ♀ from Holland
Unguiculus 3 with bifid subapical filament
2021.11.16 © van Veen, K.



Species condei (Delamare Deboutteville & Massoud, 1963:177) Delamare Deboutteville & Massoud, 1963

            Syn.: Sminthurides Condéi Delamare Deboutteville & Massoud, 1963 IOS

Species fridakahloae Ferreira, AS, Arango, A et Palacios-Vargas, JG, 2021:11

Sminthurides fridakahloae from Mexico
Facial chaetotaxy
After Ferreiro AS & al, 2021 Fig.25
Sminthurides fridakahloae from Mexico
Left: modified designation of ocelli
Right: original designation
Modified after Ferreiro AS & al, 2021 Fig.38b

            Syn.: Sminthurides sp. Palacios-Vargas, 2000
            Syn.: Sminthurides sp.undescribed Ferreira, AS, Arango, A et Palacios-Vargas, JG, 2021:1
            Syn.: Sminthurides sp. Ferreira, AS, Arango, A et Palacios-Vargas, JG, 2021:14



            Syn.: Siminthurides(sic) fridakahloae Ferreira, AS, Arango, A et Palacios-Vargas, JG, 2021:16
             Syn.: Sminthurides fridakalohae(sic) Palacios-Vargas, JG, 2022:4 ISS
             Syn.: Sminthurides fridakalohae(sic) Palacios-Vargas, JG, 2022:5 ISS

Species pejelagarto Palacios-Vargas, JG, 2022:4

Sminthurides pejelagarto ♂ from Mexico
Left: original designation of ocelli
Right: modified designation
Modified after Palacios-Vargas JG, 2022 Fig.15
Sminthurides pejelagarto ♂ from Mexico
Facial chaetotaxy
After Palacios-Vargas JG, 2022 Fig.16
Sminthurides pejelagarto ♀ from Mexico
Facial chaetotaxy
After Palacios-Vargas JG, 2022 Fig.25
Remark by Janssens, F. 2022.08.25: According to the etymology, "pejelagarto" is a vernacular Mexican name of a fish from the Tabasca state. The ICZN Rec.11A discourages the use of vernacular words as scientific names. Given appropriate latinisation is the prefered means of formation of names from vernacular words, "pejelagartus" would be an example of an appropriate name in line with Rec.11A. Now it is only a matter of time for the new 'Sminthurides fishandchips' from the UK to emerge ;-)
             Syn.: Sminthurides pejelargarto(sic) Palacios-Vargas, JG, 2022:14 ISS
             Syn.: Sminthurides pejelargarto(sic) Palacios-Vargas, JG, 2022:15 ISS


Species penicillifer (Schäffer, 1896) Börner, C, 1901:94, B

            Syn.: Sminthurides penicillifer var. incompta(sic) Börner, C, 1901:94 IOS
             Syn.: Sminthurides penicillifer var. incompta(sic) Voigts, 1903 ISS
             Syn.: Sminthurides penicillifer var. incompta(sic) Heymons, R et Heymons, H in Brauer, A, 1909:12 ISS
             Syn.: Sminthurides penicillifer var. incompta(sic) Linnaniemi, WM, 1912:252 ISS
             Syn.: Sminthurides penicillifer f. incompta(sic) Handschin, E in Dahl, F, 1929:108 ISS
             Syn.: Sminthurus penicillifer var. incompta(sic) Kel'schtein, 1929 ISS
             Syn.: Sminthurides penicillifer var. incompta(sic) Stach, J, 1956:41 ISS
             Syn.: Sminthurides penicillifer var. incompta(sic) Grinbergs, 1960 ISS
             Syn.: Sminthurides penicillifer var. incompacta(sic) Palissa, A in Brohmer, P, Ehrmann, P et Ulmer, G, 1964:241 ISS
             Syn.: Sminthurides penicillifer f. incomota(sic) Tarashchuk, 1995 ISS
Sminthurides penicillifer var. incomptus ♀ from the UK
With Podura aquatica spermatophore
2010.07.09 © Michaelson, J.
Sminthurides penicillifer var. incomptus from the UK
Footcomplex of proleg; striated mucro
2013.11.01 © Murray, A.
Sminthurides penicillifer var. incomptus ♀ from the UK
Footcomplex of mesoleg; hydrophylic outer face of unguis
2013.11.01 © Murray, A.
Sminthurides penicillifer var. incomptus ♀ from the UK
Extended mouthparts
2013.11.01 © Murray, A.
Sminthurides penicillifer var. incomptus ♀ from Germany
2014.09.15 © Beck, K.
Sminthurides penicillifer var. incomptus ♀ from Germany
Feeding instar
2014.09.17 © Beck, K.
Sminthurides penicillifer var. incomptus ♀ from Germany
Ant.4 with 7 subdivisions and rings
2014.09.17 © Beck, K.
Sminthurides penicillifer var. incomptus ♀ from Germany
Mucro
2014.09.17 © Beck, K.
Sminthurides penicillifer var. incomptus ♀ from Germany
2014.09.17 © Beck, K.
Sminthurides penicillifer var. incomptus ♂ juv. from Germany
Ant.2 with two bothriotricha
2014.09.17 © Beck, K.
Sminthurides penicillifer var. incomptus ♀ from Germany
2014.09.17 © Kluck, A.
Sminthurides penicillifer var. incomptus ♀ from Germany
2014.09.17 © Kluck, A.
Sminthurides penicillifer var. incomptus ♀ from Germany
Facial interantennal spot
2014.09.20 © Beck, K.
Sminthurides penicillifer var. incomptus ♀ from Germany
Mucro with shortened outer lamella
2014.11.09 © Kluck, A.
Sminthurides penicillifer var. incomptus ♀ from Germany
Facial interantennal spot
2014.11.09 © Kluck, A.
Sminthurides penicillifer var. incomptus ♀ from the UK
Aggressive? encounter of two females
2014.12.03 © Ardron,P.A.
Sminthurides penicillifer var. incomptus ♀ from the UK
Aggressive? encounter of two females
2014.12.03 © Ardron,P.A.
Sminthurides penicillifer var. incomptus ♀ from Germany
2015.08.30 © Kluck, A.
Sminthurides penicillifer var. incomptus ♂ from Germany
2015.08.30 © Kluck, A.
Sminthurides penicillifer var. incomptus ♀ from the UK
2016.12.11 © Phillips, E.
Sminthurides penicillifer var. incomptus ♀ from the UK
Unguiculus (ung) longer than unguis (UNG)
2016.12.15 © Phillips, E.


            Syn.: Sminthurides penicillifer f. typica(sic) Börner, C, 1901:94 IOS
            Syn.: Sminthurides Pauliani(sic) Denis, 1936
            Syn.: Sminthurides malmgreni Janssens, F, 1992:99,B



Sminthurides penicillifer ♀ & ♂ from Germany
After Börner, 1901
Sminthurides penicillifer from Germany
Foreleg with long unguiculus
After Börner, 1901 Taf.II Fig.11a
Sminthurides penicillifer from Germany
Hindleg with multifaded unguiculus
After Börner, 1901 Taf.II Fig.11b

             Syn.: Sminthurides penicilifer(sic) Kseneman, M, 1932:22 ISS



Sminthurides penicillifer ♀ from Poland
After Stach J, 1956 Pl.V Fig.1
Sminthurides penicillifer ♀ from Belgium
After Janssens F, 1995 Fig.1a
Sminthurides penicillifer ♀ from Belgium
After Janssens F, 1995 Fig.1b
Sminthurides penicillifer ♀ from Belgium
After Janssens F, 1995 Fig.2b
Sminthurides penicillifer ♂ from Belgium
After Janssens F, 1995 Fig.3a
Sminthurides penicillifer ♂ from Belgium
After Janssens F, 1995 Fig.3b
Sminthurides penicillifer ♂ juv. from Germany
2014.03.23 © Beck, K.
Sminthurides penicillifer ♂ & ♀ from Germany
Courtship
2014.09.12 © Beck, K.
Sminthurides penicillifer ♂ juv. from Germany
2014.09.17 © Beck, K.
Sminthurides penicillifer ♂ juv. from Germany
2014.09.17 © Beck, K.
Sminthurides penicillifer ♂ from Germany
Male antennal clasping organ
2014.09.17 © Beck, K.
Sminthurides penicillifer ♂ juv. from Germany
With undeveloped pearly ant.2-3
2014.09.23 © Beck, K.
Sminthurides penicillifer ♂ & ♂ from Germany
Juvenile ♂ grabbed by adult ♂
2014.11.09 © Kluck, A.
Sminthurides penicillifer ♂ & ♀ juv. from the UK
Courtship practise
2014.12.09 © Ardron,P.A.
Sminthurides penicillifer ♀ from France
Dorsal aspect
2016.11.27 © Garcelon, P.
Sminthurides penicillifer ♀ juv. & ♂ from France
Courtship
2020.09.30 © Reinach, Br.
Sminthurides penicillifer ♀ juv. & ♂ from France
Courtship
2020.10.03 © Reinach, Br.

             Syn.: Sminthurides Penicillifer(sic) Ferreira, AS, Arango, A et Palacios-Vargas, JG, 2021:1,gr ISS

  Subspecies ssp.1 Janssens F & Murray A, 2012

Remark by Janssens, F. 2017.03.03: With white circumlateral band.

            Syn.: Sminthurides penicillifer ssp.1 f. ldovi Janssens F, 2018
Sminthurides nr penicillifer ssp.1 f. ldovi ♀ & juv. from Ukraine
2015.07.06 © L'dov, S.
Sminthurides nr penicillifer ssp.1 f. ldovi ♀ from Ukraine
2015.07.06 © L'dov, S.
Sminthurides nr penicillifer ssp.1 f. ldovi juv. from Ukraine
2015.07.06 © L'dov, S.
Sminthurides nr penicillifer ssp.1 f. ldovi ♀ from Ukraine
2016.06.22 © L'dov, S.
Sminthurides nr penicillifer ssp.1 f. ldovi ♀ from Ukraine
Chewing cavity in epidermis of Lemna
2018.04.10 © L'dov, S.
Sminthurides nr penicillifer ssp.1 f. ldovi ♀ from Ukraine
Egg deposition in cavity
2018.04.10 © L'dov, S.
Sminthurides nr penicillifer ssp.1 f. ldovi ♀ from Ukraine
Verifying egg position
2018.04.10 © L'dov, S.

            Syn.: Sminthurides penicillifer ssp.1 f. principalis Janssens F, 2018
Sminthurides nr penicillifer ♀ from the UK
2012.04.21 © Murray, A.
Sminthurides nr penicillifer ♀ from the UK
2012.04.21 © Murray, A.
Sminthurides nr penicillifer ♀ from the UK
2012.04.21 © Murray, A.
Sminthurides nr penicillifer ♀ from the UK
2012.04.21 © Murray, A.
Sminthurides nr penicillifer ♀ from the UK
2012.04.26 © Murray, A.
Sminthurides nr penicillifer ♂ from the UK
2012.04.26 © Murray, A.
Sminthurides nr penicillifer from the UK
Courtship ritual
2012.04.26 © Murray, A.
Sminthurides nr penicillifer from the UK
2012.04.30 © Murray, A.
 
Sminthurides nr penicillifer from the UK
2012.04.30 © Murray, A.
 
Sminthurides nr penicillifer from the UK
Footcomplex with split empodium
2012.05.01 © Shaw, P.
Sminthurides nr penicillifer from the UK
2012.05.02 © Murray, A.
 
Sminthurides nr penicillifer from the UK
2012.05.02 © Murray, A.
 
Sminthurides nr penicillifer juv. from the UK
2012.05.24 © Murray, A.
Sminthurides nr penicillifer juv. ♂ from the UK
2012.05.28 © Murray, A.
Sminthurides nr penicillifer ♀ from the UK
2012.06.02 © Murray, A.
Sminthurides nr penicillifer ♀ & ♂ from the UK
2012.06.04 © Murray, A.
Sminthurides nr penicillifer ♀ from the UK
2012.06.04 © Murray, A.
 
Sminthurides nr penicillifer ♀ from the UK
2012.06.04 © Murray, A.
 
Sminthurides nr penicillifer ♀ from the UK
2012.06.04 © Murray, A.
Sminthurides nr penicillifer ♀ & ♂ from the UK
2012.06.04 © Murray, A.
Sminthurides nr penicillifer juv. ♂ from the UK
2012.06.06 © Murray, A.
 
Sminthurides nr penicillifer from the UK
Lamellate mucro
2012.06.dd © Shaw, P.
Sminthurides nr penicillifer ♀ from the UK
2013.09.24 © Murray, A.
Sminthurides nr penicillifer from the UK
2013.09.24 © Murray, A.
Sminthurides nr penicillifer from the UK
2013.10.05 © Nurcombe, E.
Sminthurides nr penicillifer ♀ from the UK
2013.10.08 © Nurcombe, E.
Sminthurides nr penicillifer ♀ from the UK
2013.10.08 © Nurcombe, E.
Sminthurides nr penicillifer ♂ & ♀ juv. from the UK
Courtship ritual
2013.10.08 © Nurcombe, E.
Sminthurides nr penicillifer ♀ from the UK
Eyepatch with 8 ocelli: 6 large 2 small
2014.07.07 © Nurcombe, E.
Sminthurides nr penicillifer ♂ & ♀ from the UK
Courtship ritual 1. ♂ clasps to ♀
2014.07.07 © Nurcombe, E.
Sminthurides nr penicillifer ♂ & ♀ from the UK
Courtship ritual 2. ♀ lifts up ♂
2014.07.07 © Nurcombe, E.
Sminthurides nr penicillifer ♂ & ♀ from the UK
Clasping mechanism
2014.07.07 © Nurcombe, E.
Sminthurides nr penicillifer ♂ & ♀ from Germany
Courtship
2014.09.12 © Beck, K.
Sminthurides nr penicillifer ♂ & ♀ from Germany
♂ approaches ♀ in courtship
2014.09.19 © Beck, K.
Sminthurides nr penicillifer ♂ & ♀ from Germany
♂ grasps antenna of ♀ in courtship
2014.09.19 © Beck, K.
Sminthurides nr penicillifer ♀ from Germany
Facial pigmentation; lamellate mucro
2014.09.20 © Beck, K.
Sminthurides nr penicillifer ♀ from Germany
Anal segment
2014.09.20 © Beck, K.
Sminthurides nr penicillifer ♀ & ♂ from Holland
Courtship
2015.07.02 © van Leeuwen, J.
Sminthurides nr penicillifer ♂ & ♀ from Germany
Courtship
2015.08.13 © Jäschke, J.
Sminthurides nr penicillifer ♀ from Germany
Eyepatches
2015.08.13 © Jäschke, J.
Sminthurides nr penicillifer ♂ from Holland
2018.07.18 © Kamsteeg, G.
Sminthurides nr penicillifer ♀ from Holland
2018.07.26 © Kamsteeg, G.
Sminthurides nr penicillifer ♂ from Holland
2018.07.26 © Kamsteeg, G.
Sminthurides nr penicillifer ♀ & ♂ from Holland
Courtship
2018.07.26 © Kamsteeg, G.
Sminthurides nr penicillifer ♀ & ♂ from Holland
Courtship
2018.07.26 © Kamsteeg, G.
Sminthurides nr penicillifer juv. from Holland
With egg
2018.07.26 © Kamsteeg, G.
Sminthurides nr penicillifer ♀ from France
2019.08.27 © Dejoux, P.
Sminthurides nr penicillifer ♂ from Holland
2019.09.10 © Kamsteeg, G.
Sminthurides nr penicillifer ♀ from Holland
2019.09.10 © Kamsteeg, G.
Sminthurides nr penicillifer ♀ from Holland
Caudal aspect
2019.09.10 © Kamsteeg, G.
Sminthurides nr penicillifer ♀ juv. from Holland
Pale juvenile
2019.09.10 © Kamsteeg, G.
Sminthurides nr penicillifer ♀ juv. from Holland
2019.09.12 © Kamsteeg, G.
Sminthurides nr penicillifer ♀ juv. from Holland
Cephalic pattern
2019.09.12 © Kamsteeg, G.
Sminthurides nr penicillifer ♂ from Holland
2019.09.12 © Kamsteeg, G.
Sminthurides nr penicillifer ♀ juv. & ♂ from Holland
Courtship
2019.09.23 © Kamsteeg, G.
Sminthurides nr penicillifer ♀ & ♂ from Holland
Courtship
2020.07.31 © Kamsteeg, G.
Sminthurides nr penicillifer ♂ from Holland
Dorsal pattern
2020.07.31 © Kamsteeg, G.
Sminthurides nr penicillifer ♂ from Holland
Ant.2 with 2 bothriotricha (*)
2020.07.31 © Kamsteeg, G.
Sminthurides nr penicillifer ♂ from Holland
Clasping antenna
2020.07.31 © Kamsteeg, G.
Sminthurides nr penicillifer ♀ from Holland
Dorsal pattern
2020.07.31 © Kamsteeg, G.
Sminthurides nr penicillifer ♀ from Holland
Facial pattern
2020.07.31 © Kamsteeg, G.
Sminthurides nr penicillifer ♀ & ♂ from Holland
Begin of courtship
2021.05.17 © Kamsteeg, G.
Sminthurides nr penicillifer ♀ & ♂ from Holland
End of courtship
Male clasped to female antennae
2021.05.17 © Kamsteeg, G.

            Syn.: Sminthurides penicillifer ssp.1 f. kamsteegi Janssens F, 2019
Sminthurides nr penicillifer ♀ from Holland
Bluish colour form
2019.09.23 © Kamsteeg, G.
Sminthurides nr penicillifer ♀ from Holland
Bluish colour form
2019.09.23 © Kamsteeg, G.





Species spegazzinii Börner, 1907

            Syn.: Sminthurus fuscus Parona, 1895
            Syn.: Sminthurus fuscus Börner, 1907
            Syn.: Sminthurus viridis Parona, 1895 partim
            Syn.: Sminthurus viridis Börner, 1907 partim


             Syn.: Sminthurides spegazzini(sic) Bonet, F, 1933:124 ISS
             Syn.: Sminthurides spegazzini(sic) Bonet, F, 1933:126 ISS
             Syn.: Sminthurides Spegazzinii(sic) Murphy, 1960 ISS
             Syn.: Sminthurides Spegazzinii(sic) Delamare Deboutteville, C et Massoud, Z, 1963:175 ISS
             Syn.: Sminthurides spegazzinni(sic) Ferreira, AS, Arango, A et Palacios-Vargas, JG, 2021:20 ISS

Species-group undefined

Species acanthus Hüther, 1967

Species afer Marlier, 1944:5

Species annulicornis Axelson, WM, 1905:793

             Syn.: Sminthurides ann.(sic) Bretfeld, G et Griegel, A, 1999:223 ISS

Species antennatus Baijal & Verma, 1986

Species appendiculatus Imms, 1912

Species armatus Bretfeld, 2000

Species avicenniae Murphy, 1960

Species bicuspidatus Bretfeld, G, 2001:121

Species biwaensis Yosii, R, 1970:12

            Syn.: Sminthurides biwae(sic) Yosii, R, 1970:12 IOS
             Syn.: Sminthurides biwae(sic) Gough, HJ, 1975:653 ISS
             Syn.: Sminthurides biwae(sic) Gough, HJ, 1975:654 ISS
             Syn.: Sminthurides biwae(sic) Yosii, 1977 ISS
             Syn.: Sminthurides biwae(sic) Bretfeld, G, 1999:35 ISS
             Syn.: Sminthurides biwae(sic) Bretfeld, G, 1999:36 ISS
             Syn.: Sminthurides biwae(sic) Bretfeld, G, 1999:39 ISS



Species bothrium Nguyen, 2001

Species cruciatus Axelson, WM, 1905:792

Sminthurides cruciatus from Finland
Pale form
After Linnaniemi, 1912 Plate XIV Fig.24
Sminthurides cruciatus from Finland
Dark form
After Linnaniemi, 1912 Plate XIV Fig.25
Sminthurides cruciatus ♀ from Finland
Head
After Linnaniemi, 1912 Plate XIV Fig.27
Sminthurides cruciatus from Poland
After Stach J, 1956 Plate III Fig.1
Sminthurides cruciatus from Russia
2003.mm.dd © Potapov, M.
Sminthurides cruciatus subadult ♀ & ♂ from Holland
Courtship
2017.09.28 © van Duinen, J.
Sminthurides cruciatus ♀ from Holland
2017.09.28 © van Duinen, J.
Sminthurides cruciatus ♀ from Holland
2017.09.28 © van Duinen, J.
Sminthurides cruciatus ♀ from Holland
Caudal aspect
2017.09.28 © van Duinen, J.
Sminthurides cruciatus ♂ from Holland
Caudal aspect
2017.09.28 © van Duinen, J.
Sminthurides cruciatus ♀ from Holland
Dark form
2017.09.28 © van Duinen, J.
Sminthurides cruciatus ♀ from Holland
Dark form
2017.09.28 © van Duinen, J.
Sminthurides cruciatus ♀ from Holland
Dark form
2017.09.28 © van Duinen, J.

Species gambiae Murphy, 1960

Species hessei (Poppe, 1886)

Species hospes Börner, 1907

Species inaequalis Börner, 1903

            Syn.: Sminthurides sexoculatus Unknown YYYY =?


             Syn.: Sminthurides inequalis(sic) Stach, J, 1956:19 ISS
Sminthurides inaequalis ♀ from Italy
After Dallai R, 1973 Fig.14a
Sminthurides inaequalis ♂ from Italy
After Dallai R, 1973 Fig.14a



Species lepus Mills, HB, 1934:91

Sminthurides lepus from the USA
After Mills HB, 1934 Plate 2 Fig.13
Sminthurides (Sminthurides) lepus ♂ from the USA
After Maynard EA, 1951 Pl.XXII Fig.359
Sminthurides (Sminthurides) lepus ♀ from the USA
After Maynard EA, 1951 Pl.XXII Fig.360
Sminthurides (Sminthurides) lepus ♀ from the USA
After Maynard EA, 1951 Pl.XXII Fig.361
Sminthurides (Sminthurides) lepus ♀ from the USA
After Maynard EA, 1951 Pl.XXII Fig.362
Sminthurides (Sminthurides) lepus ♀ from the USA
After Maynard EA, 1951 Pl.XXII Fig.363
Sminthurides lepus ♂ from the USA
After Maynard EA, 1951 Pl.XXIV Fig.422
Sminthurides lepus ♂ from the USA
After Maynard EA, 1951 Pl.XXIV Fig.423
Sminthurides lepus from the USA
2008.06.04 © Schoenherr, J.
Sminthurides lepus from the USA
2008.06.04 © Schoenherr, J.
Sminthurides lepus from the USA, female & male
2008.06.04 © Schoenherr, J.
Sminthurides lepus from the USA
2008.06.04 © Schoenherr, J.

Species lolelua Christiansen, K et Bellinger, P, 1992:340

Species longicornis (Joseph G, 1882) Börner, C, 1901:137

            Syn.: Arrhopalites? sp. Stach, J, 1956:239


Species macgregori Murphy, 1960

Species macnamarai Folsom & Mills, 1938

            Syn.: Sminthurides assimilis Folsom & Mills, 1938 =?
Sminthurides (Sminthurides) assimilis ♀ from the USA
After Maynard EA, 1951 Pl.XXII Fig.364
Sminthurides (Sminthurides) assimilis ♀ from the USA
After Maynard EA, 1951 Pl.XXII Fig.365
Sminthurides assimilis ♀ from the USA
After Maynard EA, 1951 Pl.XXV Fig.464
Sminthurides assimilis ♀ from the USA
After Maynard EA, 1951 Pl.XXV Fig.465
Sminthurides assimilis ♂ from the USA
After Maynard EA, 1951 Pl.XXV Fig.466




Sminthurides (Sminthurides) macnamarai from Canada
After Christiansen K & Bellinger P, 1998 Fig.895A
Sminthurides macnamarai? from the USA
After Dukes CD & al, 2022 Fig.5B

Species macroceros Arlé, R, 1961:36

Species malmgreni (Tullberg, T, 1877:30) Axelson, 1905, B

            Syn.: Sminthurus Malmgrenii(sic) Tullberg, T, 1877:30 IOS
            Syn.: Sminthurus elegantulus Reuter, OM in Lindberg, SO, 1881:204
Sminthurus Malmgrenii(sic) var. elegantulus from Sweden
After Schött H, 1893 Taf.II Fig.20

             Syn.: Sminthurus Malmgreni(sic) var. elegantula(sic) Carl, J, 1899:353 ISS
             Syn.: Sminthurides malmgreni var. elegantula(sic) Börner, C, 1901:94 ISS
             Syn.: Sminthurides malmgreni var. elegantula(sic) Axelson, WM, 1905:792 ISS
             Syn.: Sminthurides Malmgreni(sic) var. elegantula(sic) Heymons, R et Heymons, H in Brauer, A, 1909:16 ISS
             Syn.: Sminthurides malmgreni var. elegantula(sic) Linnaniemi, WM, 1912:255 ISS
             Syn.: Sminthurides malmgreni var. elegantula(sic) Linnaniemi, WM, 1912:257 ISS
             Syn.: Sminthurides malmgreni var. elegantula(sic) Linnaniemi, WM, 1912:258 ISS
             Syn.: Sminthurides malmgreni var. elegantula(sic) Davies, WM, 1927:27 ISS
             Syn.: Sminthurides (Sminthurides) malmgreni f. elegantula(sic) Handschin, E in Dahl, F, 1929:109 ISS
Sminthurides (Sminthurides) malmgreni f. elegantulus ♀ from the USA
After Maynard EA PlateXXV Fig.454
Sminthurides (Sminthurides) malmgreni f. elegantulus ♂ from the USA
After Maynard EA PlateXXV Fig.455

             Syn.: Sminthurides malmgreni var. elegantula(sic) Stach, J, 1956:238 ISS
            Syn.: Sminthurides elegantulus ssp. malmgrenii(sic) Salmon, JT, 1964:563 lapsus
Sminthurides malmgreni f. elegantulus ♀ from Sweden
2006 © Hall, K.
Sminthurides malmgreni f. elegantulus ♀ from Holland
2012.01.25 © van Duinen, J.
Sminthurides malmgreni f. elegantulus ♂ from the UK
2012.08.02 © Murray, A.
Sminthurides malmgreni f. elegantulus ♀ from Holland
2014.11.07 © van Duinen, J.
Sminthurides malmgreni f. elegantulus ♂ from Holland
2014.11.07 © van Duinen, J.
Sminthurides malmgreni f. elegantulus ♀ from Belgium
2016.08.11 © Huskens, M.-L.
Sminthurides malmgreni f. elegantulus ♀ from Belgium
2016.08.12 © Huskens, M.-L.
Sminthurides malmgreni f. elegantulus ♀ from Belgium
Caudal aspect
2016.09.23 © Huskens, M.-L.
Sminthurides malmgreni f. elegantulus ♀ from Belgium
Cleaning antenna with drop of saliva
2016.09.23 © Huskens, M.-L.
Sminthurides malmgreni f. elegantulus ♀ from Belgium
Eyepatch with 6 large + 2 small ocelli
2016.09.23 © Huskens, M.-L.
Sminthurides malmgreni f. elegantulus ♀ from Germany
2017.08.22 © Jäschke, J.
Sminthurides malmgreni f. elegantulus ♀ from the UK
2019.02.27 © Smart, J.
Sminthurides malmgreni f. elegantulus ♀ from the UK
2019.02.27 © Smart, J.
Sminthurides malmgreni f. elegantulus ♀ from the UK
2019.02.27 © Smart, J.


            Syn.: Smynthurus(sic) socialis Folsom, JW, 1896:446
Smynthurus(sic) socialis from the USA
Fore foot with long unguis
After Folsom JW, 1896 Plate10 Fig.1
Smynthurus(sic) socialis from the USA
Hind foot with short unguis
After Folsom JW, 1896 Plate10 Fig.3
Smynthurus(sic) socialis from the USA
Mucro with serrate inner and smooth outer lamella
After Folsom JW, 1896 Plate10 Fig.2
Smynthurus(sic) ♂ socialis from the USA
Modified clasping antenna
After Folsom JW, 1896 Plate10 Fig.4

            Syn.: Sminthurides malmgreni var. nigrescens Börner, C, 1901:96
Sminthurides malmgreni f. nigrescens ♀ & ♂ from the UK
2008.02.dd © Ardron, P.A.
Sminthurides malmgreni f. nigrescens ♂ & ♀ from the UK
2008.02.dd © Ardron, P.A.
Sminthurides malmgreni f. nigrescens ♀ from Germany
2009.12.12 © Marquard, F.
Sminthurides malmgreni f. nigrescens ♀ from Norway
2009.04.16 © Fjellberg, A.
Sminthurides malmgreni f. nigrescens ♀ from Germany
2010.02.09 © Marquard, F.
Sminthurides malmgreni f. nigrescens ♀ from Poland
2015.05.14 © Beck, K.
Sminthurides malmgreni f. nigrescens ♀ from Poland
2015.05.14 © Beck, K.
Sminthurides malmgreni f. nigrescens ♀ from Holland
2016.02.13 © van Duinen, J.
Sminthurides malmgreni f. nr nigrescens ♀ from the USA
2016.12.23 © Dobbs, Br.
Sminthurides malmgreni f. nigrescens ♀ from France
2017.10.31 © Tiky, S.
Sminthurides malmgreni ♀ f. nigrescens from Belgium
2023.02.15 © Huskens, M.-L.
Sminthurides malmgreni ♀ f. nigrescens from Belgium
2023.02.15 © Huskens, M.-L.
Sminthurides malmgreni ♀ f. nigrescens from Belgium
2023.02.15 © Huskens, M.-L.


            Syn.: ... abrupta Ågren, 1903
            Syn.: Sminthurides maculatus Ågren, 1903
             Syn.: Sminthurides (Sminthurides) malmgreni f. maculata(sic) Handschin, E in Dahl, F, 1929:109 ISS
             Syn.: Sminthurides malmgreni f. maculata(sic) Stach, J, 1956:31 ISS
Sminthurides malmgreni ♀
Medium form
After Stach J, 1956 Pl.II Fig.5
Sminthurides malmgreni ♀ light form
After Janssens F, 1995 Fig.2c
Sminthurides malmgreni ♂ f. maculatus from the UK
2012.04.26 © Murray, A.
Sminthurides malmgreni ♂ & ♀ f. maculatus from the UK
Courtship
2012.05.11 © Murray, A.
Sminthurides malmgreni ♂ & ♀ juv. f. maculatus from Germany
Courtship ritual
2015.03.27 © Landgraf, A.
Sminthurides malmgreni ♀ f. maculatus from Belgium
Grooming balancing on 3 right legs
2016.11.04 © Huskens, M.-L.
Sminthurides malmgreni ♀ f. maculatus from Belgium
2017.02.19 © Huskens, M.-L.
Sminthurides malmgreni ♀ f. maculatus from Belgium
Mucro with lamellae
2017.02.19 © Huskens, M.-L.
Sminthurides malmgreni f. maculatus ♀ from France
With dark thoracic midstripe
2018.05.13 © Picard, J.
Sminthurides malmgreni f. maculatus ♀ juv. & ♂ from France
Courtship
♂ with distinctly dark thorax
2018.05.13 © Picard, J.

            Syn.: Sminthurides quadrilineatus Ågren, 1903
             Syn.: Sminthurides (Sminthurides) malmgreni f. quadrilineata(sic) Handschin, E in Dahl, F, 1929:109 ISS
             Syn.: Sminthurides malmgreni f. quadrilineata(sic) Stach, J, 1956:32 ISS
Sminthurides malmgreni ♀ light form
After Stach J, 1956 Pl.II Fig.4
Sminthurides malmgreni f. quadrilineatus from France
2008.03.02 © Lebeaux, P.
Sminthurides malmgreni f. quadrilineatus from France
Male with clasping antennae
2008.03.02 © Lebeaux, P.
Sminthurides malmgreni f. quadrilineatus from France
2008.04.13 © Lebeaux, P.
Sminthurides malmgreni f. nr quadrilineatus ♀ from Canada
2010 © Biodiv. Inst. Ontario.
Sminthurides malmgreni f. quadrilineatus ♂ & ♀ from France
Courtship
2016.12.08 © Garcelon, P.
Sminthurides malmgreni f. quadrilineatus ♀ from France
2017.10.22 © Picard, J.
Sminthurides malmgreni f. quadrilineatus ♀ from France
Caudal aspect
2017.10.22 © Picard, J.
Sminthurides malmgreni f. quadrilineatus ♂ from France
Ant.2-3 clasping mechanism
2017.10.22 © Picard, J.
Sminthurides malmgreni f. quadrilineatus ♂ from France
2017.11.12 © Tiky, S.
Sminthurides malmgreni f. quadrilineatus ♀ from France
Pale form
2017.11.25 © Picard, J.
Sminthurides malmgreni f. quadrilineatus ♂ & ♀ from France
Courtship ritual
2018.03.11 © Garcelon, P.
Sminthurides malmgreni f. quadrilineatus ♀ from France
2018.03.11 © Garcelon, P.
Sminthurides malmgreni f. quadrilineatus ♀ from France
2018.12.15 © Picard, J.
Sminthurides malmgreni f. quadrilineatus ♀ from France
2019.12.12 © Tiky, S.
Sminthurides malmgreni f. quadrilineatus ♀ from France
2019.12.12 © Tiky, S.

            Syn.: Sminthurides malmgreni var. immaculata(sic) Axelson, WM, 1905:792 IOS
            Syn.: Sminthurides malmgreni var. immaculata(sic) Linnaniemi, WM, 1912:256 IOS
            Syn.: Sminthurides (Sminthurides) malmgreni f. immaculata(sic) Handschin, E in Dahl, F, 1929:109 IOS
             Syn.: Sminthurides malmgreni f. unimaculata(sic) Agrell, 1941 lapsus ISS
            Syn.: Sminthurides malmgreni f. immaculata(sic) Stach, J, 1956:31 IOS
            Syn.: Sminthurides malmgreni f. immaculata(sic) Palissa, A in Brohmer, P, Ehrmann, P et Ulmer, G, 1964:243 IOS
            Syn.: Sminthurides malmgreni var. immaculatus Christiansen, K et Bellinger, P, 1981:1085
Sminthurides malmgreni f. immaculatus ♀ from France
2008.04.13 © Lebeaux, P.
Sminthurides malmgreni f. immaculatus ♀ from the UK
2011 © Shaw, P.
Sminthurides malmgreni f. immaculatus ♀ from Germany
2014.10.23 © Horak, T.
Sminthurides malmgreni f. immaculatus ♀ from France
2018.01.06 © Picard, J.
Sminthurides malmgreni f. immaculatus ♂ & f. principalis ♀ from France
Courtship
2018.04.01 © Picard, J.


            Syn.: Sminthurides cruciatus Ionescu, 1915
            Syn.: Sminthurides ludovicianus Folsom & Mills, 1938 partim
            Syn.: Sminthurides malmgreni var. palustris Folsom & Mills, 1938
Sminthurides (Sminthurides) malmgreni f. palustris ♂ from the USA
After Maynard EA PlateXXV Fig.452
Sminthurides (Sminthurides) malmgreni f. palustris ♀ from the USA
After Maynard EA PlateXXV Fig.453
Sminthurides (Sminthurides) malmgreni from the USA
After Folsom & Mills, 1938 in Christiansen K & Bellinger P, 1998 Fig.896A

            Syn.: Sminthurides malmgreni f. principalis Stach, J, 1956:32
Sminthurides malmgreni f. principalis ♀ from the UK
2008.02.dd © Ardron, P.A.
Sminthurides malmgreni f. principalis ♂ & ♀ from the UK
2008.02.dd © Ardron, P.A.
Sminthurides malmgreni f. principalis ♀ from Norway
2009.12.18 © Fjellberg, A.
Sminthurides malmgreni f. principalis ♀ & ♂ from France
Courtship
2018.04.01 © Picard, J.
Sminthurides malmgreni f. principalis ♀ & f. immaculata ♂ from France
Courtship
2018.04.01 © Picard, J.
Sminthurides malmgreni f. principalis ♀ from France
2018.04.01 © Picard, J.

            Syn.: Sminthurides (Spinosella) dentospinalis Lawrence, 1963 = juv
            Syn.: Sminthurides sp. Greenslade, 1987
            Syn.: Sminthurides sp. Greenslade, 1990
            Syn.: Sminthurides sp. Greenslade, 2002
            Syn.: Sminthurus pilleatus Yao AL & Kluge G, 1995
Sminthurus pilleatus
1995 © Yao, A.-L. & Kluge, G.
Modified after Palissa, 1964 Fig.2
Sminthurus pilleatus
2000 © Chen, S.

            Syn.: Sminthurides malmgreni f. dorsomaculatus Janssens F & Picard J., 2018
Sminthurides malmgreni ♀ from the USA
2016.12.23 © Dobbs, Br.
Sminthurides malmgreni f. dorsomaculatus ♀ from France
Dorsal spot only; ventrolateral band reduced
2018.03.04 © Picard, J.





            Syn.: Sminthurus Malmgrenii(sic) Schött, H, 1893:34 IOS
Sminthurus Malmgrenii(sic) from Sweden
After Schött H, 1893 Taf.II Fig.20

             Syn.: Sminthurus Malmgreni(sic) Reuter, OM, 1895:13 ISS
            Syn.: Smynthurus(sic) malmgrenii(sic) Folsom, JW, 1896:447 IOS
Sminthurides malmgreni ♀ dark form
After Stach J, 1956 Pl.II Fig.6

             Syn.: Sminthurides Malmgreni(sic) Delamare Deboutteville & Massoud, 1963 ISS
Sminthurides malmgreni from Germany
After Palissa, A., 1964 Fig.2

             Syn.: Sminthurides malmgremi(sic) Richards, WR, 1968:16 ISS
Sminthurides (Sminthurides) malmgreni from the USA
After Folsom & Mills, 1938 in Christiansen K & Bellinger P, 1998 Fig.896B

             Syn.: Sminthurides malgreni(sic) Shaw, P, 2003:5 ISS
             Syn.: Sminthurides malmgeni(sic) Hogg, D et Hebert, PDN, 2004:752 ISS
Sminthurides malmgreni from the UK
Mucrones
2004 © Hopkin, S.P.
Sminthurides malmgreni from the UK
2007.03.04 © Cornwall, N.J.
 
Sminthurides malmgreni ♂ juv. from France
2009.09.dd © Lebeaux, P.
Sminthurides malmgreni from Norway
2009.11.24 © Fjellberg, A.
Sminthurides malmgreni ♀ from Holland
2011.02.16 © van Duinen, J.
Sminthurides malmgreni ♂ from Holland
2011.02.16 © van Duinen, J.
Sminthurides malmgreni juv. from France
2012.01.15 © Lebeaux, P.

             Syn.: Sminthurides malmegreni(sic) Porco, D, Skarzynski, D, Decaëns, T, Hebert, PDN et Deharveng, L, 2013:8 ISS
Sminthurides malmgreni from the UK
2013.02.15 © Nurcombe, E.
Sminthurides malmgreni juv. from the UK
2013.02.17 © Nurcombe, E.
Sminthurides malmgreni ♀ & Dicyrtomina saundersi ♂ from Holland
Love at first sight ;-)
2014.11.11 © van Duinen, J.
Sminthurides malmgreni ♀ from the UK
2015.01.19 © Murray, A.
Sminthurides malmgreni ♀ from Belgium
2016.04.17 © Huskens, M.-L.
Sminthurides malmgreni ♀ from the USA
2017.10.27 © Dobbs, Br.
Sminthurides malmgreni ♂ subadult from France
Ant.2 > ant.2
2017.10.29 © Picard, J.
Sminthurides malmgreni ♂ & ♀ from the UK
Courtship ritual: male clasped to female
2017.11.09 © Horton, M.
Sminthurides malmgreni ♂ from the UK
Clasping mechanism
2017.11.09 © Horton, M.
Sminthurides malmgreni ♂ from the UK
Evaluating a juvenile
2017.12.07 © Horton, M.
Sminthurides malmgreni ♂ from the UK
Clasping mechanism
2017.12.07 © Horton, M.
Sminthurides malmgreni ♂ from the UK
Fight between males?
2017.12.07 © Horton, M.
Sminthurides malmgreni ♀ & ♂ from France
Courtship
2019.02.20 © Tiky, S.
Sminthurides malmgreni ♀ from Belgium
2019.07.07 © Huskens, M.-L.
Sminthurides malmgreni ♀ from Belgium
2020.11.18 © Huskens, M.-L.
Sminthurides malmgreni ♀ from Belgium
Unguiculus of 3rd foot with long filament (*)
2020.11.18 © Huskens, M.-L.





Species melanotus Börner, 1908

Species millsi (Marlier, 1944:6) Stach, 1956, s.i.

            Syn.: Sminthurides (Sminthurides) Millsi Marlier, 1944 IOS

Species monnioti Massoud & Betsch, 1966

Species parvulus (Krausbauer, 1898) Heymons, R & Heymons, H in Brauer, A, 1909:13, B

            Syn.: Sminthurides quinquelineatus Gisin, 1943
            Syn.: Sminthurides quinquearticulatus Gisin, 1944
             Syn.: Sminthurides 5-articulatus(sic) Cassagnau, P, 1953:227 ISS
            Syn.: Sminthurides alpinus Cassagnau, P, 1960:407


Sminthurides parvulus from Germany
After Heymons R & Heymons H, 1909 Fig.18
Sminthurides parvulus from the UK
Mucrones gutter-like
2004 © Hopkin, S.P.
Sminthurides parvulus ♀ from Luxembourg
Ant.4 with 5 subdivisions
After Marx MT & Weber D, 2013 Fig.4
Sminthurides parvulus ♀ from the UK
Fourth antennal segment with 5 subdivisions
2013.01.05 © Murray, A.
Sminthurides parvulus ♂ from the UK
2013.01.06 © Murray, A.
Sminthurides parvulus ♂ from the UK
Male grasping antenna
2013.01.06 © Murray, A.
Sminthurides parvulus ♀ juv. from the UK
Mucro gutter-like (*)
2015.02.13 © Murray, A.
Sminthurides parvulus ♀ juv. from the UK
Ant.4 with 5 subdivisions
2015.02.13 © Murray, A.
Sminthurides parvulus ♀ juv. from Holland
2016.08.25 © van Duinen, J.
Sminthurides parvulus ♂ & ♀ from France
Courtship
2018.01.06 © Picard, J.
Sminthurides parvulus ♂ from France
Mucro gutter-like (*)
2018.01.18 © Picard, J.
Sminthurides parvulus ♂ from France
Metathoracic bladder-like vesicles of male
2018.01.18 © Picard, J.
Sminthurides parvulus ♂ from France
Metathoracic bladder-like vesicles of male
2018.01.20 © Picard, J.
Sminthurides parvulus ♂ from France
Male clasping mechanism
b3-5 on common papilla
2018.02.10 © Picard, J.
Sminthurides parvulus ♂ from France
Male clasping mechanism
2018.02.10 © Picard, J.
Sminthurides parvulus ♀ from France
Ant.4 with 5 subdivisions
2018.02.12 © Picard, J.
Sminthurides parvulus ♀ from France
2018.02.17 © Picard, J.
Sminthurides parvulus ♀ juv. from Belgium
2021.03.04 © Huskens, M.-L.
Sminthurides parvulus ♀ juv. from Belgium
2021.03.04 © Huskens, M.-L.
Sminthurides parvulus ♀ juv. from Belgium
Ant.4 with 5 subdivisions
2021.03.04 © Huskens, M.-L.
Sminthurides parvulus ♀ from Belgium
Ant.4 with 5 subdivisions
2021.04.28 © Huskens, M.-L.
Sminthurides parvulus ♀ from Belgium
Ant.4 with 5 subdivisions
2021.11.16 © Huskens, M.-L.
Sminthurides parvulus ♀ from Belgium
2021.11.16 © Huskens, M.-L.
Sminthurides parvulus ♀ from Belgium
Dorso-caudal aspect
2021.11.16 © Huskens, M.-L.

Species plicatus (Schött, H, 1891:13) Folsom & Mills, 1938

Sminthurus plicatus from the USA
After Schött H, 1891 Taf.II Fig.2
Sminthurides (Sminthurides) plicatus from the USA
After Scott & Yosii in Christiansen K & Bellinger P, 1981 Fig.898A
Sminthurides plicatus ♀ from Canada
2009 © Porco, D.
Sminthurides plicatus ♂? juv. from the USA
2017.10.25 © Dobbs, Br.
Sminthurides plicatus ♂? juv. from the USA
2017.10.25 © Dobbs, Br.
Sminthurides plicatus ♀ from the USA
Dark form
2017.11.30 © Dobbs, Br.
Sminthurides plicatus ♀ from the USA
Dark form
2017.11.30 © Dobbs, Br.
Sminthurides plicatus ♀ from the USA
2017.12.08 © Dobbs, Br.
Sminthurides plicatus ♀ from the USA
Antennae with 2 white rings
2017.12.08 © Dobbs, Br.
Sminthurides plicatus ♀ from the USA
2017.12.08 © Dobbs, Br.
Sminthurides plicatus ♀ from the USA
2017.12.08 © Dobbs, Br.
Sminthurides plicatus ♀ from the USA
2017.12.09 © Dobbs, Br.
Sminthurides plicatus juv. from the USA
2017.12.09 © Dobbs, Br.
Sminthurides plicatus ♂? juv. from the USA
2017.12.09 © Dobbs, Br.
Sminthurides plicatus ♀ from the USA
2017.12.10 © Dobbs, Br.
Sminthurides plicatus ♀ from the USA
2017.12.11 © Dobbs, Br.
Sminthurides plicatus ♂ from the USA
2017.12.11 © Dobbs, Br.
Sminthurides plicatus ♂ from the USA
2017.12.11 © Dobbs, Br.
Sminthurides plicatus ♀ from the USA
2017.12.13 © Dobbs, Br.
Sminthurides plicatus ♀ from the USA
2017.12.14 © Dobbs, Br.
Sminthurides plicatus ♀ juv. from the USA
2017.12.15 © Dobbs, Br.
Sminthurides plicatus ♀ from the USA
2017.12.23 © Dobbs, Br.
Sminthurides plicatus ♀ from the USA
2021.05.07 © Pavy, R.

Species potamobius Yosii, R, 1970:10

Sminthurides potamobius ♀ juv. from Japan
After Yosii R, 1970 Fig.7A
Sminthurides potamobius ♀ from Taiwan
2022.01.19 © Cheng, H.-J.

Species pseudassimilis Stach, J, 1956:43

            Syn.: Sminthurides sp. Axelson, 1905
            Syn.: Sminthurides signatus var. Linnaniemi, 1907
            Syn.: Sminthurides assimilis Linnaniemi, 1912, nec Krausbauer, 1898
            Syn.: Sminthurides assimilis Folsom & Mills, 1938, nec Krausbauer, 1898
            Syn.: Sminthurides signatus var. distincta Linnaniemi, WM, 1912:262
            Syn.: Sminthurides signatus Womersley, H, 1939:222
            Syn.: Sminthurides pseudassimilis var. potamobius Gough, HJ, 1975:653 =?
             Syn.: Sminthurides pseudassimilis var. biwae(sic) Gough, HJ, 1975:653 =? ISS
             Syn.: Sminthurides pseudassimilis var. biwae(sic) Gough, HJ, 1975:654 =? ISS


             Syn.: Sminthurides pseudassimillis(sic) Zhang, S, Xie, Z, Dou, Y, Sun, X, Chang, L et Wu, D, 2022:9 ISS

Species quinquesensillatus Hüther, 1967

Species rupium Najt, 1967

Species schoetti Axelson, WM, 1903:12, B

            Syn.: Sminthurides Schötti(sic) Axelson, WM, 1903:12 IOS
            Syn.: Sminthurides sp. Schött, 1902
             Syn.: Sminthurides Schötti(sic) var. bilineata(sic) Axelson, WM, 1903:13 ISS
             Syn.: Sminthurides schötti(sic) var. bilineata(sic) Linnaniemi, WM, 1912:269 ISS
             Syn.: Sminthurides (Sminthurides) schötti(sic) f. bilineata(sic) Handschin, E in Dahl, F, 1929:110 ISS
             Syn.: Sminthurides schoetti var. bilineata(sic) Stach, J, 1956:37 ISS
Sminthurides schoetti var. bilineatus ♀ from Belgium
2017.03.08 © Huskens, M.-L.
Sminthurides schoetti var. bilineatus ♀ from Holland
With 2 parallel dark longitudinal lines middorsally on abdomen
2017.11.02 © van Duinen, J.
Sminthurides schoetti var. bilineatus ♀ from Holland
2017.11.02 © van Duinen, J.
Sminthurides schoetti var. bilineatus ♀ from Holland
Ant.1:2:3:4 = 1:1.7:2.7:5.7
2017.11.02 © van Duinen, J.
Sminthurides schoetti var. bilineatus ♂ from France
2018.01.28 © Picard, J.
Sminthurides schoetti var. bilineatus ♀ from France
2018.01.28 © Picard, J.
Sminthurides schoetti var. bilineatus ♀ from France
2018.01.28 © Picard, J.
Sminthurides schoetti var. bilineatus ♀ from France
Ant.4 with 4 subdivisions
2018.01.28 © Picard, J.
Sminthurides schoetti var. bilineatus ♀ from France
Mucro without lamellae
2018.01.28 © Picard, J.
Sminthurides schoetti var. bilineatus ♂ & ♀ from France
Courtship
2018.02.03 © Picard, J.
Sminthurides schoetti var. bilineatus ♂ from France
Mucro with apical constriction (*)
2018.02.03 © Picard, J.
Sminthurides schoetti var. bilineatus ♂ & ♀ from France
Courtship
2018.02.03 © Picard, J.
Sminthurides schoetti var. bilineatus ♂ from France
2018.10.27 © Picard, J.
Sminthurides schoetti var. bilineatus ♀ from France
2018.10.27 © Picard, J.
Sminthurides schoetti var. bilineatus ♀ from France
2018.10.27 © Picard, J.
Sminthurides schoetti var. bilineatus ♀ from Belgium
2019.06.10 © Huskens, M.-L.
Sminthurides schoetti var. bilineatus ♀ from Belgium
2020.02.18 © Huskens, M.-L.
Sminthurides schoetti var. bilineatus ♀ from Belgium
2020.02.18 © Huskens, M.-L.
Sminthurides schoetti var. bilineatus ♀ from Belgium
Caudal aspect
2020.02.18 © Huskens, M.-L.

            Syn.: Sminthurides Schötti(sic) var. ornata(sic) Axelson, WM, 1903:13 IOS
             Syn.: Sminthurides schötti(sic) var. ornata(sic) Linnaniemi, WM, 1912:269 ISS
             Syn.: Sminthurides (Sminthurides) schötti(sic) f. ornata(sic) Handschin, E in Dahl, F, 1929:110 ISS
             Syn.: Sminthurides schoetti var. ornata(sic) Stach, J, 1956:38 ISS
Sminthurides schoetti var. ornatus ♂ subadult from the UK
With distinct lateral white spot
2012.07.08 © Nurcombe, E.
Sminthurides schoetti var. ornatus ♂ subadult from the UK
Swollen ant.1+2
2012.07.08 © Nurcombe, E.
Sminthurides schoetti var. ornatus ♂ subadult from the UK
2012.07.08 © Nurcombe, E.
Sminthurides schoetti var. ornatus ♂ subadult from the UK
2012.07.08 © Nurcombe, E.
Sminthurides schoetti var. ornatus ♂ subadult from the UK
Ant.2 > ant.3
2012.07.08 © Nurcombe, E.


            Syn.: Sminthurus signatus Paclt, J, 1961:235



             Syn.: Sminthurides schötti(sic) Axelson, 1904 ISS
             Syn.: Sminthurides schötti(sic) Axelson, 1905 ISS
             Syn.: Sminthurides schötti(sic) Linnaniemi, WM, 1912:267 ISS
             Syn.: Sminthurides (Sminthurides) schötti(sic) Handschin, E in Dahl, F, 1929:110 ISS
             Syn.: Sminthurides (Sminthurides) schötti(sic) Kloet, GS et Hincks, WD, 1945:9 ISS
             Syn.: Sminthurides schötti(sic) Joosse-Van Damme, ENG et Naaktgeboren, C, 1961:34 ISS
             Syn.: Sminthurides schötti(sic) Paclt, J, 1961:235 ISS
Sminthurides schoetti from the UK
Mucrones
2004 © Hopkin, S.P.
Sminthurides schoetti
2006 © Boldsystems.org
 
Sminthurides schoetti juv. from the UK
2012.06.21 © Murray, A.
 
Sminthurides schoetti juv. from the UK
2012.06.27 © Nurcombe, E.
Sminthurides schoetti juv. from the UK
2012.06.27 © Nurcombe, E.
Sminthurides schoetti ♂ subadult from the UK
Distinctly pale basal antennal segment
2012.07.08 © Nurcombe, E.
Sminthurides schoetti from the UK
Pale colour form
2012.07.08 © Nurcombe, E.
Sminthurides schoetti ♀ from the UK
Subdivided 4th antennal segment
2012.07.08 © Nurcombe, E.
Sminthurides schoetti ♀ from the UK
2012.09.07 © Nurcombe, E.
 
Sminthurides schoetti ♂ from the UK
2012.10.09 © Nurcombe, E.
Sminthurides schoetti ♂ from the UK
2012.10.09 © Nurcombe, E.
Sminthurides schoetti ♀ from the UK
2012.10.09 © Nurcombe, E.
Sminthurides schoetti ♀ & ♂ from the UK
2012.10.09 © Nurcombe, E.
Sminthurides nr schoetti ♀ juv. from Portugal
2012.10.16 © Murray, A.
Sminthurides schoetti juv. from the UK
2012.12.21 © Barton, T.
Sminthurides schoetti ♂ subadult from the UK
Swollen ant.1+2; ant.2 > ant.3
2013.09.18 © Nurcombe, E.
Sminthurides schoetti juv. from the UK
2013.09.24 © Nurcombe, E.
Sminthurides schoetti ♀ from the UK
2013.09.24 © Nurcombe, E.
Sminthurides schoetti ♂ subadult; from Holland
Ant.2 > ant.3
2014.10.05 © van Duinen, J.
Sminthurides schoetti ♀ from Holland
2014.10.09 © van Duinen, J.
Sminthurides schoetti ♂ & ♀ from Holland
Courtship
2014.11.17 © van Duinen, J.
Sminthurides schoetti ♂ & ♀ from Holland
Courtship
2014.11.17 © van Duinen, J.
Sminthurides schoetti ♂ subadult from France
Ant.2 swollen; ant.2 > ant.3
2017.09.07 © Picard, J.
Sminthurides schoetti ♀ juv. from France
2017.12.02 © Picard, J.
Sminthurides schoetti ♀ juv. from France
2017.12.02 © Picard, J.
Sminthurides schoetti ♀ from France
Mucro with apical constriction (*)
2018.02.10 © Picard, J.
Sminthurides schoetti ♂ from France
Male antennal clasping mechanism
2018.02.10 © Picard, J.
Sminthurides schoetti ♂ from France
Male antennal clasping mechanism
2018.02.10 © Picard, J.
Sminthurides schoetti ♀ & ♂ from France
Courtship ritual
2018.03.26 © Picard, J.
Sminthurides schoetti ♀ from France
2019.02.05 © Tiky, S.
Sminthurides schoetti ♀ from France
2019.02.05 © Tiky, S.
Sminthurides schoetti ♂ & ♀ from France
Courtship
2019.02.12 © Tiky, S.
Sminthurides schoetti ♂ & ♀ from France
Courtship
2019.02.12 © Tiky, S.
Sminthurides schoetti ♀ from France
Mucro with constriction (*)
2019.02.12 © Tiky, S.
Sminthurides schoetti ♀ & ♂ from Belgium
Courtship
2020.03.04 © Huskens, M.-L.
Sminthurides schoetti ♀ from Belgium
2020.03.04 © Huskens, M.-L.
Sminthurides schoetti ♀ from Belgium
2020.03.04 © Huskens, M.-L.
Sminthurides schoetti ♀ from Belgium
2020.03.04 © Huskens, M.-L.
Sminthurides schoetti ♀ from Belgium
2020.03.04 © Huskens, M.-L.
Sminthurides schoetti ♀ juv. from Belgium
2020.03.04 © Huskens, M.-L.
Sminthurides schoetti ♂ from Belgium
2020.03.05 © Huskens, M.-L.
Sminthurides schoetti ♂ from Italy
2020.06.25 © Colombo, M.
Sminthurides schoetti ♂ from Italy
2020.06.25 © Colombo, M.
Sminthurides schoetti ♂ from Italy
Male antennal clasping mechanism
2020.06.25 © Colombo, M.
Sminthurides schoetti ♀ from Belgium
2020.11.27 © Huskens, M.-L.
Sminthurides schoetti juv. ♂ from Belgium
With thick ant.2+3
With unsubdivided ant.4
2021.02.03 © Huskens, M.-L.
Sminthurides schoetti ♀ from Belgium
With thin ant.2+3
With subdivided ant.4
2021.02.03 © Huskens, M.-L.
Sminthurides schoetti ♀ from Belgium
2021.02.25 © Huskens, M.-L.
Sminthurides schoetti ♀ from Belgium
Feeding instar
2021.02.25 © Huskens, M.-L.
Sminthurides schoetti ♀ from Belgium
2021.08.05 © Huskens, M.-L.
Sminthurides schoetti ♂ from Belgium
Antenna clasping mechanism
b6 absent (*)
2022.02.12 © Huskens, M.-L.





Species sensillatus Massoud & Betsch, 1966

Species sexoculatus Betsch, J-M et Massoud, Z, 1970:213

Sminthurides sexoculatus ♀ from France
Lateral cephalic chaetotaxy
After Betsch J-M & Massoud Z, 1970 Fig.13C
Sminthurides sexoculatus ♂ from France
Facial chaetotaxy
After Betsch J-M & Massoud Z, 1970 Fig.14B
Sminthurides sexoculatus ♂ from France
Male clasping antenna
After Betsch J-M & Massoud Z, 1970 Fig.14D
Sminthurides sexoculatus ♀ from France
2017.03.20 © Picard, J.
Sminthurides sexoculatus ♀ from France
2017.03.20 © Picard, J.
Sminthurides sexoculatus subadult ♀ & ♂ from France
2017.03.20 © Picard, J.
Sminthurides sexoculatus ♀ from France
2017.03.22 © Picard, J.
Sminthurides sexoculatus ♀ from France
Pointed mucro
2017.03.22 © Picard, J.
Sminthurides sexoculatus ♂ from France
Male clasping mechanism
2018.02.17 © Picard, J.
Sminthurides sexoculatus ♀ from France
2019.02.05 © Tiky, S.
Sminthurides sexoculatus ♂ from France
2019.02.05 © Tiky, S.
Sminthurides sexoculatus ♂ from France
2019.02.05 © Tiky, S.
Sminthurides sexoculatus ♀ from France
Sharply pointed mucro
2019.02.05 © Tiky, S.
Sminthurides sexoculatus ♀ from France
2019.02.06 © Tiky, S.
Sminthurides sexoculatus ♀ from France
Eye with 6 ocelli
2019.02.06 © Tiky, S.

Species sexsensillatus Hüther, 1967

Species signatus (Krausbauer, 1898 partim) Börner, 1900

            Syn.: Sminthurus assimilis Krausbauer, 1898


Sminthurides signatus from Germany
After Heymons R & Heymons H, 1909 Fig.17
Sminthurides signatus from the UK
Mucrones
2004 © Hopkin, S.P.
Sminthurides signatus from the UK
2012.07.22 © Murray, A.
 
Sminthurides signatus from the UK
2012.07.22 © Murray, A.
Sminthurides signatus from the UK
2012.08.14 © Murray, A.
Sminthurides signatus ♂ from the UK
2012.08.14 © Murray, A.
Sminthurides signatus juv. from the UK
2012.09.09 © Nurcombe, E.
Sminthurides signatus juv. from the UK
2012.09.09 © Nurcombe, E.
Sminthurides signatus from the UK
2012.10.09 © Nurcombe, E.
Sminthurides signatus from the UK
2012.10.09 © Nurcombe, E.
Sminthurides signatus from the UK
2012.10.09 © Nurcombe, E.
Sminthurides signatus from the UK
2012.10.21 © Murray, A.
Sminthurides signatus ♀ from the UK
2012.10.25 © Nurcombe, E.
Sminthurides signatus ♀ from the UK
2012.10.25 © Nurcombe, E.
Sminthurides signatus ♀ from the UK
2013.01.04 © Nurcombe, E.
Sminthurides signatus ♀ from the UK
2013.01.04 © Nurcombe, E.
Sminthurides signatus ♂ from the UK
2013.01.04 © Nurcombe, E.
Sminthurides signatus ♀ from the UK
2013.01.04 © Nurcombe, E.
Sminthurides signatus ♀ juv. & ♂ from France
Courtship
2017.10.15 © Picard, J.
Sminthurides signatus ♂ from France
Courtship
2017.10.15 © Picard, J.
Sminthurides signatus ♂ from France
Mucro with narrow lamellae
2017.10.15 © Picard, J.
Sminthurides signatus ♂ from France
2017.09.16 © Picard, J.
Sminthurides signatus ♂ from France
2017.09.16 © Picard, J.
Sminthurides signatus ♂ from France
2017.10.22 © Picard, J.
Sminthurides signatus ♀ & juv. from France
2017.10.22 © Picard, J.
Sminthurides signatus ♀ from France
Ant.4 with 4 subdivisions
Ant.4.1=ant.4.4
2017.10.22 © Picard, J.
Sminthurides signatus ♀ from France
2017.10.22 © Picard, J.
Sminthurides signatus ♀ from France
2017.10.25 © Picard, J.
Sminthurides signatus ♀ from Denmark
2019.01.06 © Schlütter, P.

Species signatus2 James, 1933

Species signatus3 Murphy, 1960

Species sundanus Yoshii, R et Suhardjono, YR, 1992:89

Sminthurides sundanus from Indonesia
After Yoshii R & Suhardjono YR, 1992 Fig.8A

Species tropicus Delamare Deboutteville, 1952

Species velii Prabhoo, 1971

             Syn.: Sminthurides velli(sic) Bano, R, 2016:56 ISS
             Syn.: Sminthurides velli(sic) Bano, R, 2016:64 ISS

Species ventanae Najt, 1969

            Syn.: Sminthurides ventanae Massoud, Z et Betsch, J-M, 1972:73 = 1st instar Sminthurididae

Species weichseli Christiansen & Bellinger, 1981

Sminthurides (Sminthurides) weichseli from the USA
After Christiansen K & Bellinger P, 1998 Fig.900A
Sminthurides weichseli from the USA
Eggs deposited in epidermal cavity
2013.09.07 © Krebs, C.
Sminthurides weichseli from the USA
Pale embryo and well pigmented embryo
2013.09.07 © Krebs, C.
Sminthurides weichseli juv. from the USA
2013.09.07 © Krebs, C.
Sminthurides weichseli ♂ & ♀ from the USA
2013.09.10 © Krebs, C.
Sminthurides weichseli ♂ & ♀ from the USA
Ant.4 with weakly developed, spherical subapical knob (*)
2013.09.10 © Krebs, C.


Genus Parasminthurides Medeiros G et Bellini BC in da Silva Medeiros, G, Nunes, RC, Zhang, F, Godeiro, NN et Bellini, BC, 2022:3

Species sp.1 Janssens F sensu da Silva Medeiros G & al, 2022

            Syn.: Parasminthurides sp. new da Silva Medeiros, G, Nunes, RC, Zhang, F, Godeiro, NN et Bellini, BC, 2022:19


Species spinosus Medeiros G et Bellini BC in da Silva Medeiros, G, Nunes, RC, Zhang, F, Godeiro, NN et Bellini, BC, 2022:8, t.t.

Parasminthurides spinosus ♂ from Brazil
Ant.2(dorsal) + ant.3(ventral) clasping mechanism
After da Silva Medeiros G & al, 2022 Fig.3B+C
Parasminthurides spinosus ♀ from Brazil
Ant.2(dorsal) + ant.3(ventral) with spine-like setae
After da Silva Medeiros G & al, 2022 Fig.3E+F
Parasminthurides spinosus ♀ from Brazil
Facial chaetotaxy
After da Silva Medeiros G & al, 2022 Fig.4A
Parasminthurides spinosus ♂ from Brazil
Trunk chaetotaxy
After da Silva Medeiros G & al, 2022 Fig.5A
Parasminthurides spinosus ♀ from Brazil
Trunk chaetotaxy
After da Silva Medeiros G & al, 2022 Fig.5B


Genus Pygicornides Betsch, J-M, 1969:355

Pygicornides torridus ♂ from Australia
Clasping mechanism
After da Silva Medeiros G & al, 2022 Fig.1G
Modified after Betsch J-M, 1980 Fig.46I

Species stagnalis (Womersley, H, 1932:16) Greenslade, P in Houston, WWK, 1994:129

Species torridus Betsch, J-M, 1969:349, t.t.


Genus Sinnamarides Betsch, J-M et Waller, A, 1991:234

  Syn.: Sminthurididae Sinnnamarides(sic) Ramos-Díaz, A, Palacios-Vargas, JG et Pinzón-Florián, OP, 2020:11 ISS
  Syn.: Sminthurididae Sinnamerides(sic) Ramos-Díaz, A, Palacios-Vargas, JG et Pinzón-Florián, OP, 2020:14 ISS
Sinnemarides sp. ♂ from French Guiana
Clasping mechanism
After da Silva Medeiros G & al, 2022 Fig.1K



Species delamarei Betsch, J-M et Waller, A, 1991:230, t.t.


Genus Sminthuridia Massoud, Z et Betsch, J-M, 1972:70

Sminthuridia sphaeridioides ♂ from Gambia
Clasping mechanism
After da Silva Medeiros G & al, 2022 Fig.1B
Modified after Betsch J-M, 1980 Fig.46B

Species sphaeridioides (Murphy, 1960) Massoud, Z & Betsch, J-M, 1972:70, t.t.


Genus Yosiides Massoud, Z et Betsch, J-M, 1972:72

Yossides himachal ♂ from Nepal
Clasping mechanism
After da Silva Medeiros G & al, 2022 Fig.1E
Modified after Betsch J-M, 1980 Fig.46C

Species chinensis Zhao L-J in Itoh, R et Zhao, L-J, 1993:31

Yosiides chinensis ♂ from China
After Itoh R & Zhao J-L, 1993 Fig.1
Yosiides chinensis ♀ from China
After Itoh R & Zhao J-L, 1993 Fig.8

Species himachal (Yosii, 1966) Massoud, Z & Betsch, J-M, 1972:72, t.t.


Genus Stenacidia Börner, C, 1906:182

  Syn.: Sminthuridae Jeannenotia Stach, J, 1956:10
  Syn.: ... Sminthurides Jeannenot, 1955
  Syn.: Sminthurididae Spinosellina Betsch, J-M, 1980:124 partim = 1st instar
  Syn.: Sminthuridida(sic) Spinosellina Bretfeld, G, 1999:59 partim


Stenacidia violacea ♀ from France
Ant.4 with long blunt setae
2017.09.20 © Picard, J.
Stenacidia stachi ♂ from the Holarctic
Clasping mechanism
After da Silva Medeiros G & al, 2022 Fig.1H
Modified after Betsch J-M, 1980 Fig.46F

Species picta Yosii, 1966

Species stachi (Jeannenot, 1955) Dallai, R, 1973:547

Species violacea (Reuter, OM in Lindberg, SO, 1881:203) auctt., t.t.

            Syn.: Stenacidia violacea var. variabilis Krausbauer, 1898
Sminthurides (Stenacidia) violaceus var. variabilis from Germany
After Heymons R & Heymons H, 1909 Fig.13

            Syn.: Arrhopalites adelaidica Womersley, 1933 partim
            Syn.: Arrhopalites coccineus Salmon, JT, 1941:408
Arrhopalites coccineus from New Zealand
After Salmon JT, 1941 Pl.70 Fig.484

            Syn.: Jeannenotia stachi ssp. australiensis Betsch, J-M et Massoud, Z, 1970:173



Sminthurus violaceus from Finland
After Schött H, 1893 Taf.II Fig.8

             Syn.: Sminthurides (Stenacidia) violacea(sic) Womersley, H, 1932:14 ISS
Sminthurides (Stenacidia) violaceus ♂ from Australia
After Womersley H, 1932 Fig.2b
Sminthurides violaceus ♀ juv. from the USA
After Maynard EA, 1951 Pl.XXIII Fig.411
Sminthurides violaceus ♂ from the USA
After Maynard EA, 1951 Pl.XXIII Fig.412
Stenacidia violacea from the UK
Male antenna
2004 © Hopkin, S.P.
Stenacidia violacea from the UK
2011.08.17 © Robertson, A.
Stenacidia violacea from the UK
2011.08.17 © Robertson, A.
Stenacidia violacea ♂ juv. from the UK
With partly developed grasping antennae
2011.08.17 © Robertson, A.
Stenacidia violacea ♀ & ♂ from the UK
Courtship
2012.06.21 © Murray, A.
Stenacidia violacea ♀ & ♂ from the UK
Courtship
2012.06.21 © Murray, A.
Stenacidia violacea ♂ from the UK
Abdominal dorsal rows of setae
2012.06.21 © Murray, A.
Stenacidia violacea ♂ from the UK
Grasping antenna
2012.06.22 © Murray, A.
Stenacidia violacea ♀ from the UK
2012.06.22 © Murray, A.
Stenacidia violacea juv. from the UK
2012.07.20 © Murray, A.
Stenacidia violacea ♂ from the UK
2012.08.08 © Nurcombe, E.
Stenacidia violacea ♂ from the UK
2012.08.08 © Nurcombe, E.
Stenacidia violacea ♀ from the UK
2012.08.19 © Nurcombe, E.
Stenacidia violacea ♀ from the UK
2012.09.07 © Nurcombe, E.
Stenacidia violacea ♂ & ♀ from the UK
♂ grasps ♀ with modified antennae
2012.09.07 © Nurcombe, E.
Stenacidia violacea ♀ & ♂ from the UK
♀ lifts up ♂ in courtship ritual
2012.09.07 © Nurcombe, E.
Stenacidia violacea ♀ juv. from the UK
2012.10.07 © Murray, A.
Stenacidia violacea ♀ juv. from the UK
2012.10.07 © Murray, A.
Stenacidia violacea ♂ from the UK
2012.10.10 © Nurcombe, E.

             Syn.: Sminthurides (Stenacidia) violacea(sic) Thibaud, J-M, 2013:239 ISS
Stenacidia violacea ♀ from France
2013.03.dd © Laprevote, S.
Stenacidia violacea ♀ from the UK
2013.04.01 © Nurcombe, E.
Stenacidia violacea ♀ from the UK
2013.09.24 © Nurcombe, E.
Stenacidia violacea ♂ from the UK
Male clasping antenna
2013.10.24 © Nurcombe, E.
Stenacidia violacea ♂ & ♀ from the UK
Male clasped to antennae of female
2013.10.24 © Nurcombe, E.
Stenacidia violacea ♂ juv. from Holland
2014.10.03 © van Duinen, J.
Stenacidia violacea ♂ from Holland
Antennal clasping organ
2014.10.04 © van Duinen, J.
Stenacidia violacea ♀ & ♀ from Holland
Courtship
2014.10.04 © van Duinen, J.
Stenacidia violacea ♂ from Holland
Antennal clasping organ
2014.10.05 © van Duinen, J.
Stenacidia violacea ♂ from Holland
2014.10.09 © van Duinen, J.
Stenacidia violacea ♀ juv. from Belgium
Facial chaetotaxy
2016.01.02 © Huskens, M.-L.
Stenacidia violacea ♀ from France
2017.09.14 © Picard, J.
Stenacidia violacea ♀ from France
Cleaning
Retractable anal sac?
2017.09.14 © Picard, J.
Stenacidia violacea ♀ from France
Unguis with tunica
2017.09.14 © Picard, J.
Stenacidia violacea ♂ from France
2017.09.20 © Picard, J.
Stenacidia violacea ♂ from France
Ant.2+3 grasping mechanism
2017.09.20 © Picard, J.
Stenacidia violacea ♀ from France
Grooming on 3 right legs
2017.09.20 © Picard, J.
Stenacidia violacea ♀ from France
With bilateral dorsal rows of setae (*)
2017.09.20 © Picard, J.
Stenacidia violacea ♂ from France
2017.11.12 © Picard, J.
Stenacidia violacea ♀ from France
2017.11.12 © Picard, J.
Stenacidia violacea ♀ from France
Unguis with hyaline membraneous tunica (*)
2017.11.12 © Picard, J.
Stenacidia violacea ♀ juv. from Belgium
2017.12.09 © Huskens, M.-L.
Stenacidia violacea ♀ juv. from France
Narrow slender mucro (*)
2018.02.10 © Picard, J.
Stenacidia violacea ♀ juv. from France
2018.02.10 © Picard, J.
Stenacidia violacea ♀ from France
Mucro narrow slender; anterior lamella missing
2018.02.24 © Picard, J.
Stenacidia violacea ♀ from France
Bothriotricha ABC in a large obtuse angle
2018.02.24 © Picard, J.
Stenacidia violacea ♂ from France
Male clasping antenna
C1 short and bulby
2018.03.03 © Picard, J.
Stenacidia violacea ♂ juv. from France
2018.04.30 © Picard, J.
Stenacidia violacea ♂ from France
Male clasping antenna
2018.04.30 © Picard, J.
Stenacidia violacea ♂ from Denmark
With short tubular collophore vesicles (*)
2018.09.17 © Schlütter, P.
Stenacidia violacea ♂ from France
2020.02.15 © Joachim, J.
Stenacidia violacea ♂ subadult from France
2020.02.15 © Joachim, J.
Stenacidia violacea ♀ juv. from France
Feeding instar
Bothriotricha A-B-C in very obtuse angle
2020.02.15 © Joachim, J.
Stenacidia violacea ♀ from France
2020.02.15 © Joachim, J.
Stenacidia violacea ♀ juv. & ♂ from France
Courtship ritual
2020.02.15 © Joachim, J.
Stenacidia violacea ♀ juv. & ♂ from France
Courtship ritual
2020.03.22 © Reinach, Br.
Stenacidia violacea ♀ from France
2020.03.22 © Reinach, Br.
Stenacidia violacea ♀ subadult from France
2020.03.25 © Reinach, Br.
Stenacidia violacea ♂ from France
2020.03.25 © Reinach, Br.
Stenacidia violacea ♂ & ♀ from France
Courtship ritual
2020.10.10 © Joachim, J.
Stenacidia violacea ♂ & ♀ from France
Courtship ritual
2020.10.10 © Joachim, J.
Stenacidia violacea ♂ from the UK
2020.10.14 © Uncle, F.
Stenacidia violacea ♂ from the UK
Antennal clasping mechanism
2020.10.14 © Uncle, F.
Stenacidia violacea ♂ from the UK
Mucro long slender pointed
Ventral aspect
2020.10.14 © Uncle, F.
Stenacidia violacea ♀ & ♂ from Spain
Deposition of spermatophore
2020.12.23 © Zeeders, M.
Stenacidia violacea ♂ & ♀ from Spain
Courtship begin
2022.02.03 © Zeeders, M.
Stenacidia violacea ♀ & ♂ from Spain
Courtship continuation part 1
♂ trembles with front & middle legs
2022.02.03 © Zeeders, M.
Stenacidia violacea ♀ & ♂ from Spain
Courtship continuation part 2
♂ trembles ant.4 while depositing spermatophore
2022.02.03 © Zeeders, M.
Stenacidia violacea ♀ & ♂ from Spain
Courtship continuation part 3
♀ climbs over ♂ to take-up spermatophore
2022.02.03 © Zeeders, M.
Stenacidia violacea juv. ♂ from Spain
2022.02.12 © Zeeders, M.




Genus Pedonides Bretfeld, G, 2010:317

Pedonides sp. ♂ from Portugal
Clasping mechanism
After da Silva Medeiros G & al, 2022 Fig.1J

Species alcochetensis Bretfeld, G, 2010:318, t.t.

Pedonides alcochetensis from Portugal
with clasping second legs
After Bretfeld G, 2010 Fig.1,13


Genus Boernerides Bretfeld, G, 1999:32

Boernerides sp. ♂ from the Mediterranean
Clasping mechanism
After da Silva Medeiros G & al, 2022 Fig.1I

Species hystrix (Börner, 1903) Bretfeld, G, 1999:34, t.t.


Stenacidia hystrix ♀ from Italy
After Dallai R, 1973 Fig.17a
Stenacidia hystrix ♂ from Italy
After Dallai R, 1973 Fig.17b
Stenacidia hystrix ♂ from Italy
After Dallai R, 1973 Fig.18a

References