http://www.collembola.org/taxa/smi2inae.htm - Last updated on 2022.05.30 by Frans Janssens
Checklist of the Collembola: Sminthurinae

Subfamilia Sminthurinae Lubbock, 1862, sensu Deharveng, L, 2004:427 key

Genus Number
of species
Allacma (Allacma)9
Allacma (Trochopilosa)1
Allacma10
Archeallacma1
Austrosminthurus1
Brevimucronus1
Caprainea2
Galeriella1
Grinnellia2
Janusius5
Katiannasminthurus1
Keratosminthurus2
Mucrovirga1
Novokatianna4
Pararrhopalites17
Richardsitas3
Sminthurconus2
Sminthuricinus4
Sminthurus82
Spatulosminthurus6
Temeritas43
Sminthurinae188
 Genera with number of species
  Syn.: Symphypléones(sic) Arrhopalitini Delamare Deboutteville, C et Bassot, J-M, 1957:82


  Syn.: Sminthurides pseudoviolaceus Scott, 1964


Sminthurinae from Russia
2006 © Macroclub.ru.
Sminthurus sp. nov. from the USA
2007.04.30 © Cowen, R.


Genus Grinnellia Christiansen, KA et Nascimbene, P, 2006:319, -

Species sp. Christiansen, KA et Nascimbene, P, 2006:321, -

Species ventis Christiansen, KA et Nascimbene, P, 2006:320, -

Grinnellia ventis in Cretaceous amber
from the Dominican Republic
2006 © Christiansen, K. & Nascimbene, P.
Grinnellia ventis reconstruction
from the Dominican Republic
2006 © Christiansen, K. & Nascimbene, P.


Genus Mucrovirga Christiansen, KA et Nascimbene, P, 2006:325, -

Species incompleta Christiansen, KA et Nascimbene, P, 2006:326, -


Genus Sminthurconus Christiansen, KA et Nascimbene, P, 2006:328, -

Species grimaldi Christiansen, KA et Nascimbene, P, 2006:323, -

Species sp. Christiansen, KA et Nascimbene, P, 2006:329, -


Genus Sminthuricinus Christiansen, KA et Nascimbene, P, 2006:321, -

Species deceptus Christiansen, KA et Nascimbene, P, 2006:323, -

            Syn.: Sminthuricinus decepta(sic) Christiansen, KA et Nascimbene, P, 2006:323,- IOS

Species sp. Christiansen, KA et Nascimbene, P, 2006:325, -

Species sp. Christiansen, KA et Nascimbene, P, 2006:325, -

Species sp. Christiansen, KA et Nascimbene, P, 2006:325, -


Genus Brevimucronus Christiansen, KA et Pike, E, 2002:179, -

  Syn.: Dicyrtomidae? Brevimucronus Medeiros, GDS, Greenslade, P et Bellini, BC, 2020:13,-

Species anomalus Christiansen, KA et Pike, E, 2002:179, -


Genus Archeallacma Sánchez-García, A et Engel, MS, 2016:12, -

Species dolichopoda Sánchez-García, A et Engel, MS, 2016:12, -

Archeallacma dolichopoda in Cretaceous amber from Spain
After Sánchez-García A & Engel MS, 2016 Fig.8AB


Genus Katiannasminthurus Sánchez-García, A et Engel, MS, 2016:16, -

Species xenopygus Sánchez-García, A et Engel, MS, 2016:16, -

Katiannasminthurus xenopygus in Cretaceous amber from Spain
After Sánchez-García A & Engel MS, 2016 Fig.12

            Syn.: Sminthurus? sp. Simón-Benito, Ortuño & Espantaleón, 2002 -



Genus Pararrhopalites Bonet, F et Tellez, C, 1947:193 key  key

  Syn.: Sminthurini Neosphyrotheca Salmon, JT, 1964:142
  Syn.: Sminthurini Neosphyrotheca Salmon, JT, 1964:609
  Syn.: Sminthuridae Neosphyrotheca Betsch, J-M, 1980:149


  Syn.: Sminthuridae Paparrhopalites(sic) Mari Mutt, JA et Bellinger, PF, 1990:148 ISS
  Syn.: Sminthuridae Pararrhpalites(sic) Greenslade, P et Kitching, RL, 2011:339 ISS
  Syn.: Sminthuridae Parropalites(sic) Greenslade, P, 2013:46 ISS
  Syn.: Sminthuridae Pararrhopalites.(sic) Zeppelini, D, Lima, ECA, Brito, RA et Soares, GA, 2018:1 ISS
Pararrhopalites sp. from Brazil
After Cipola NG & al, 2019 Fig.17

  Syn.: Sminthurinae Parrarrhopalites(sic) Medeiros, GDS, Greenslade, P et Bellini, BC, 2020:12 ISS


Species anops Bonet, F et Tellez, C, 1947:194, t.t.

Species christianseni (Palacios-Vargas, JG & Zeppelini, D, 1996:20) Zeppelini, D, 2006:7

Species ecuadorensis Bretfeld & Trinklein, 2000

Species fasciatus (Salmon, 1951) Betsch, 1980

            Syn.: Sphyrotheca hispida Yosii, 1959


Species hennigi (Palacios-Vargas, JG & Zeppelini, D, 1996:16) Zeppelini, D, 2006:4

            Syn.: Arrhopalites hennigius(sic) Palacios-Vargas, JG et Zeppelini, D, 1996:16 IOS


             Syn.: Arrhopalites hennigius(sic) Mari Mutt, JA et Bellinger, PF, 1996:172 ISS
             Syn.: Arrhopalites hennigius(sic) Zeppelini Filho, D, 1996:33 ISS
             Syn.: Arrhopalites (Novarrhopalites) hennigius(sic) Zeppelini Filho, D, 1996:58 ISS
             Syn.: Arrhopalites hennigius(sic) Zeppelini Filho, D, [1997]:12 ISS
             Syn.: Arrhopalites hennigius(sic) Zeppelini Filho, D, 1997:122 ISS
             Syn.: Arrhopalites hennigius(sic) Zeppelini, DF et Palacios-Vargas JG, 1999:80 ISS
             Syn.: Arrhopalites hennigius(sic) Baquero, E, Herrando-Pérez, S et Jordana, R, 2005:86 ISS
             Syn.: Arrhopalites hennigii(sic) Zeppelini, D, 2006:2 ISS
             Syn.: Arrhopalites hennigius(sic) Palacios-Vargas, JG, 2006:9 ISS
             Syn.: Pararrhopalites hennigii(sic) Zeppelini, D, 2006:4 ISS
             Syn.: Pararrhopalites hennigii(sic) Zeppelini, D, Lima, ECA, Brito, RA et Soares, GA, 2018:9 ISS
             Syn.: Pararrhopalites hennigii(sic) Zeppelini, D, Brito, RA, Zampaulo, R et de Lima, ECA, 2020:27 ISS



Species indianus Baijal & Agarwal, 1972

Species noonae (Lawrence, 1968)

Species oculatus Bonet, F et Tellez, C, 1947:198

Species palaciosi Zeppelini, D et Brito, RA, 2014:1736

Species papaveroi (Zeppelini, DF & Palacios-Vargas JG, 1999:78) Zeppelini, D, 2006:10

            Syn.: Arrhopalites papaveroi Palacios-Vargas, JG et Zeppelini, D, 1996:20 nn
            Syn.: Arrhopalites papav(sic) Zeppelini Filho, D, 1996:39
            Syn.: Arrhopalites sp2 Aegla Zeppelini Filho, D, 1996:27
            Syn.: Arrhopalites sp. 2 (Aegla) Zeppelini Filho, D, 1996:37
            Syn.: Arrhopalites (Arrhopalites) Sp 2 Zeppelini Filho, D, 1996:58
            Syn.: Arrhopalites sp.2 Zeppelini Filho, D, [1997]:12
            Syn.: Arrhopalites sp. 2 Zeppelini Filho, D, 1997:122
            Syn.: Arrhopalites sp5 papabon Zeppelini Filho, D, 1996:27
            Syn.: Arrhopalites sp5 Zeppelini Filho, D, 1996:39


Species popei Lawrence, PN, 1968:329

Pararrhopalites popei from the South Pacific
After Lawrence PN, 1968 Fig.4

Species queirozi Brito, RA, de Lima, ECA et Zeppelini, D, 2019:410

Species sideroicus Zeppelini, D et Brito, RA, 2014:1734

Species ubiquitus Zeppelini D, Lima ECA et Brito RA in Zeppelini, D, Lima, ECA, Brito, RA et Soares, GA, 2018:2

            Syn.: Pararrhopalites ubiquum(sic) Zeppelini D, Lima ECA et Brito RA in Zeppelini, D, Lima, ECA, Brito, RA et Soares, GA, 2018:2 IOS
            Syn.: Pararrhopalites.(sic) ubiquum(sic) Zeppelini, D, Lima, ECA, Brito, RA et Soares, GA, 2018:1 IOS
Remark by Zeppelini, D., Lima, E.C.A., Brito, R.A. & Soares, G.A. 2018:5: Pararrhopalites ubiquum n.sp. comes from Latin ubique (ubiquitous), in allusion to its distribution in many caves and MSS in the iron ore mining areas in Minas Gerais State.
Remark by Janssens, F. 2018.10.27: "ubique", in Latin, is a compound adverb of "ubi"(="where")+"que"(="any,ever") and is an invariable part of speech. Therefore, the declension "ubiquum" is invalid. However, the adjective "ubiquitous", in English, = "ubiquitus" in Latin. The latter also matches the gender of "Pararrhopalites" in line with the ICZN code.

             Syn.: Pararrhopalites ubiquum(sic) Brito, RA, de Lima, ECA et Zeppelini, D, 2019:401 ISS
             Syn.: Pararrhopalites ubiquum(sic) Brito, RA, de Lima, ECA et Zeppelini, D, 2019:417 ISS



Species villiersi (Delamare Deboutteville, C et Massoud, Z, 1964:83)

Sminthurotheca villiersi from Angola
After Delamare Deboutteville C & Massoud Z, 1964 Planche I Fig.B

Species wallacei (Palacios-Vargas, JG & Zeppelini, D, 1996:18) Zeppelini, D, 2006:6

             Syn.: Arrhopalites walacei(sic) Mari Mutt, JA et Bellinger, PF, 1996:172 ISS

Species yinae Itoh & Zhao, 2000


Genus Temeritas Richards WR in Delamare Deboutteville, C et Massoud, Z, 1963:276 key

Temeritas sp. from Taiwan
Ant.4 with 40 subdivisions
2022.05.29 © Cheng, H.-J.
Temeritas sp. from Taiwan
2022.05.29 © Cheng, H.-J.

Species sp.1 Janssens F & Murray A, 2015

Temeritas sp. from Tasmania
2015.12.15 © Murray, A.

Species sp.2 Janssens F & Steve, 2017

Temeritas sp. from Australia
Mimicing spider posture
2017.03.24 © Steve & Alison.
Temeritas sp. from Australia
2021.01.07 © Steve & Alison.
Temeritas sp. juv. from Australia
2021.01.08 © Steve & Alison.
Temeritas sp. juv. from Australia
2021.01.08 © Steve & Alison.
Temeritas sp. juv. from Australia
2021.01.08 © Steve & Alison.

Species amazonensis Arlé & Oliveira, 1977

Species ambohitantelensis Betsch, 2000

Species ambrensis Betsch, 2000

Species ampasimbensis Betsch, J-M, 1980:162

Temeritas ampasimbensis from Madagascar
After Betsch J-M, 1980 Fig.52D.

Species andranobensis Betsch, 2000

Species andreazzei da Silva Medeiros, G et Bellini, BC, 2019:538

Temeritas andreazzei from Brazil
After da Silva Medeiros G & Bellini BC, 2019 Fig.1
?Temeritas andreazzei? from Brazil
2018.08.10 © Idlegrraphics.

Species angolensis Delamare Deboutteville, C et Massoud, Z, 1964:76

Temeritas angolensis from Angola
After Delamare Deboutteville C & Massoud Z, 1964 Planche I Fig.C

Species anjozorobensis Betsch, 2000

Species ankarafantsikensis Betsch, J-M, 1980:159

Temeritas ankarafantsikensis from Madagascar
After Betsch J-M, 1980 Fig.52B.

Species anosyennensis Betsch, J-M, 1980:164

Temeritas anosyennensis from Madagascar
After Betsch J-M, 1980 Fig.52G.

Species antongilensis Betsch, J-M, 1980:159

Temeritas antongilensis from Madagascar
After Betsch J-M, 1980 Fig.52C.

Species bergensis Betsch, J-M, 1980:156

Temeritas bergensis from Madagascar
After Betsch J-M, 1980 Fig.52A.

Species bharatensis Baijal & Kohli, 1972

Species caatingae Arlé & Oliveira, 1977

Species daneghanensis Bretfeld, G, 2005:46

Temeritas-daneghanebsis from Yemen
After Bretfeld G, 2005 Fig.87

Species denisi (Womersley, 1934) Wallace, 1974

            Syn.: Sminthurus denisii(sic) Womersley, 1934 IOS
            Syn.: Sminthurus longicornis Womersley H, 1932:38, nec Joseph, 1882
            Syn.: Sminthurus longicornis Womersley H, 1932:38, nec Schött, 1896


             Syn.: Temeritas denisii(sic) Greenslade, P in Houston, WWK, 1994:135 ISS
Temeritas denisi from Australia
After Womersley in Greenslade P, 2013 Fig.2

             Syn.: Sminthurus denisii(sic) Greenslade, P, 2013:45 ISS
             Syn.: Temeritas denisii(sic) Medeiros, GDS, Greenslade, P et Bellini, BC, 2020:1 ISS




Species dimna Mandal, GP, Suman, KK et Bhattacharya, KK, 2016:49

Temeritas dimna from India
After Mandal GP, Suman KK & Bhattacharya KK, 2016 p.49 Img.1

Species iarakensis Betsch, 2000

Species isabellae Greenslade, P, 2013:46

Temeritas isabellae from Australia
After Greenslade P, 2013 Fig.4

            Syn.: Sminthurus elegans Womersley, 1933, nec Fitch, 1863
Sminthurus elegans from Australia
After Womersley H, 1939 Fig.74P




Temeritas isabellae from Australia
2017 © Murray, A.

Species kilimandjarica (Wahlgren, 1908)

             Syn.: Arrhopalites kilimandjarensis(sic) Handschin, 1929 ISS

Species longipes (Börner, 1906)

Species macrocera (Denis, J-R, 1933:302) Delamare Deboutteville & Massoud, 1963, t.t.

            Syn.: Sminthurus macroceros(sic) Denis, J-R, 1933:302,tt IOS
Sminthurus macroceros(sic) from Costa Rica
After Denis J-R, 1933 Fig.137


            Syn.: Sminthurus pullus Denis, J-R, 1933:300
            Syn.: Sminthurus pullus Betsch, 1980 =? 1st instar
            Syn.: Sminthurus pullus Mari Mutt, JA et Bellinger, PF, 1990:147 =?
            Syn.: Temeritas pulla da Silva Medeiros, G et Bellini, BC, 2019:551




            Syn.: Sminthurus macroceros(sic) Denis, J-R, 1933:309 IOS
            Syn.: Sminthurus macroceros(sic) Denis, J-R, 1933:315 IOS
            Syn.: Sminthurus macroceros(sic) Denis, J-R, 1933:320 IOS
            Syn.: Sminthurus macroceros(sic) Denis, J-R, 1933:322 IOS
            Syn.: Sminthurus macroceros(sic) Stach, 1956 IOS
            Syn.: Temeritas macroceros(sic) Delamare Deboutteville & Massoud, 1963 IOS
            Syn.: Temeritas macroceros(sic) Betsch, 1980 IOS
            Syn.: Temeritas macroceros(sic) Mari Mutt & Bellinger, 1990 IOS
            Syn.: Temeritas macroceros(sic) Palacios-Vargas, 1997 IOS
             Syn.: Temeritas macrocerus(sic) Palacios-Vargas, JG, Castaño-Meneses, G et Gómez-Anaya, JA, 1998:50 ISS
            Syn.: Temeritas macroceros(sic) Palacios-Vargas, JG, Castaño-Meneses, G et Gómez-Anaya, JA, 1998:51 IOS
            Syn.: Temeritas macroceros(sic) Palacios-Vargas, JG, 2000:33 IOS
            Syn.: Temeritas macroceros(sic) Bretfeld, G, 2005:47 IOS
            Syn.: Temeritas macroceros(sic) Mandal, GP, Suman, KK et Bhattacharya, KK, 2016:49,tt IOS
            Syn.: Temeritas macroceros(sic) da Silva Medeiros, G et Bellini, BC, 2019:550,gr IOS
            Syn.: Temeritas (Temeritas) macroceros(sic) da Silva Medeiros, G et Bellini, BC, 2019:550 IOS



Species mahorensis Betsch, 2000

Species mananteninensis Betsch, J-M, 1980:164

Temeritas mananteninensis from Madagascar
After Betsch J-M, 1980 Fig.52F.

Species mantasoensis Betsch, J-M, 1980:164

Species marojezensis Betsch, J-M, 1980:161

Species ormondae Arlé & Oliveira, 1977

Species paradoxalis da Silva Medeiros, G et Bellini, BC, 2019:544

Temeritas paradoxalis from Brazil
After da Silva Medeiros G & Bellini BC, 2019 Fig.2

Species perinetensis Betsch, J-M, 1980:162

Temeritas perinetensis from Madagascar
After Betsch J-M, 1980 Fig.52E.

Species rapoporti Najt, 1969

Species regalis (Womersley, H, 1939:258) Najt, 1968

Sminthurus regalis from Australia
After Womersley H, 1939 Fig.74Q
Temeritas regalis from Australia
After Betsch J-M in Greenslade P, 2013 Fig.1

Species sahafarensis Betsch, 2000

Species sinensis Dallai, R et Fanciulli, PP, 1985:157

Temeritas sinensis from China
After Dallai R & Fanciulli PP, 1985 Fig.2

Species soalalensis Betsch, J-M, 1980:156

Species summelongicornis (Uchida, H, 1965:90) Yosii, R, 1977:165

Species surinamensis Delamare Deboutteville, C et Massoud, Z, 1964:70

Temeritas surinamensis from Surinam
After Delamare Deboutteville C & Massoud Z, 1964 Planche I Fig.D

Species tamurai Itoh & Zhao, 2000

Species tsaratananensis Betsch, 2000

Species tucumanensis Delamare Deboutteville, C et Massoud, Z, 1963:278

Species venezuelensis Gómez Cova & Rapoport, 1972

Species womersleyi (Denis, 1948) Delamare Deboutteville & Massoud, 1963

             Syn.: Temeritas Womersleyi(sic) Delamare Deboutteville & Massoud, 1963 ISS
Temeritas womersleyi from the South Pacific
After Lawrence PN, 1968 Fig.10
Temeritas womersleyi from Asia
After Mari Mutt JA, 1978 Cover




Genus Keratosminthurus Zeppelini D in Zeppelini, D, Brito, RA, Zampaulo, R et de Lima, ECA, 2020:26

Species calamitosus Zeppelini D, Brito RA et Lima ECA in Zeppelini, D, Brito, RA, Zampaulo, R et de Lima, ECA, 2020:38

Keratosminthurus calamitosus from Brazil
After Zeppelini D & al, 2020 Fig.5C.right
Keratosminthurus calamitosus ♂ from Brazil
Sexually dimorph chaetae of ant.3 (*)
After Zeppelini D & al, 2020 Fig.20

Species tapiguus Zeppelini D, Brito RA et Lima ECA in Zeppelini, D, Brito, RA, Zampaulo, R et de Lima, ECA, 2020:30, t.t.

            Syn.: Keratosminthurus tapigu(sic) Zeppelini D, Brito RA et Lima ECA in Zeppelini, D, Brito, RA, Zampaulo, R et de Lima, ECA, 2020:30,tt IOS
Keratosminthurus tapigu(sic) from Brazil
After Zeppelini D & al, 2020 Fig.5C.left

            Syn.: Keratosminthurus tapigu(sic) Cipola, NG, 2022:8 IOS
Remark by Janssens, F. 2022.05.27: Due to the etymology : "Tapigü is the word for horn in Kuikúru Karib, the ancient language of the indigenous people Kuikúru from 'Alto Xingú' region in central Brazil" the new name refers to 'Keratosminthurus, the horn'. Although it is not explicitly stated by the etymology, it is clear that it was the intention of the authors to reflect in the new name that the new species has a horn, as in the 'Keratosminthurus with a horn'. The mismatch between the name and the intention of the authors is clear evidence of an inadvertent error that must be corrected (Art.32.5.1). In addition, an unmodified vernacular word should not be used as a scientific name (Art.11.3 Rec.11A). Given appropriate latinization is the preferred means of formation of names from vernacular words (Rec.11A) : we propose the following emendation of the name that matches the intention of the authors: 'tapiguus' = stem 'tapigu' + latin masculine adjective suffix 'us'.




Genus Allacma Börner, C, 1906:183

  Syn.: Sminthuridae Alacma(sic) Schliwa, W, 1965:447 ISS
Subgenus Allacma Börner C, 1906:183 s.s.
Allacma fusca from Germany
Epicuticular ultrastructure
2006 © Schulz, H.-J. & Schubert H.D.
Allacma sp. from France
2007.06.09 © Lebeaux, P.
Allacma sp. from France
2007.06.09 © Lebeaux, P.
Allacma sp. from Hungary
2008.06.27 © Keresztes, G.
Allacma fusca from the UK
Ant.4 with about 15 subdivisions
2010.08.10 © Robertson, A.
Allacma fusca from the UK
Post-antennal seta (PAO)
2012.06.30 © Nurcombe, E.
Allacma fusca juv. from the UK
Composed eye with 8 ocelli in perfect bilateral symmetric lay-out
2013.04.28 © Nurcombe, E.
Allacma fusca juv. from the UK
Bulby post-antennal seta (PAO)
2013.08.23 © Phillips, E.
Allacma fusca from the UK
Unguis with broad tunica
2013.11.18 © Nurcombe, E.
Allacma fusca from Germany
Slightly curved rough dorsal macroseta
2014.06.24 © Beck, K.
Allacma fusca from Germany
Composed eye with 8 pearly ocelli
2014.06.24 © Beck, K.
Allacma fusca from Germany
Ant.2 with long thick setae
2014.06.24 © Beck, K.
Allacma fusca from the UK
Ant.3 in basal half with long thick macrosetae
2014.07.14 © Nurcombe, E.




Species sp.1 Bernard, 2007

Allacma sp. nov. from the USA
2007.02.22 © Bernard, E.C.
Allacma sp. nov. from the USA
2007.02.22 © Bernard, E.C.
Allacma sp. nov. from the USA
2007.02.22 © Bernard, E.C.

Species cryptica Baquero E et Jordana R in Baquero, E, Jordana, R et Ortuño, VM, 2021:13

             Syn.: Allacma crypticae(sic) Baquero, E, Jordana, R et Ortuño, VM, 2021:19 ISS

Species dubia Börner, 1907

            Syn.: Corynephoria boerneri Salmon, JT, 1964:629

Species fusca (Linnæus, C, 1758:608) Börner, C, 1906:183, t.t., B

            Syn.: Podura globosa Linnæus, C, 1746:342
             Syn.: Smynthurus(sic) Buskii(sic) Lubbock, J, 1862:431 ISS
             Syn.: Smynthurus(sic) Buskii(sic) Lubbock, J, 1862:590 ISS
             Syn.: Smynthurus(sic) Buskii(sic) Lubbock, J, 1870:295 ISS
             Syn.: Podura fusco-nigra(sic) Tullberg, T, 1872:29 ISS
             Syn.: Podura fusco-nigra(sic) Guthrie, JE, 1903:53 ISS
             Syn.: Podura fusco-nigra(sic) Salmon, JT, 1964:606 ISS
            Syn.: Smynthurus(sic) plumbeus Lemoine, 1882
            Syn.: Sminthurus plumbeus Claypole, AM, 1898:70
            Syn.: Dicyrtoma fusca Parona, 1884
            Syn.: Sminthurus purpurascens Reuter, 1891
            Syn.: Sminthurus fuscus var. atra Scherbakov, AM, 1898:21
            Syn.: Sminthurus (Eusminthurus) fuscus var. albiceps Börner, C, 1901:119
Sminthurus fuscus var. albiceps
from Germany
After Börner C, 1901 Tafel I Fig.6
Allacma fusca var. albiceps from France
2007 © Benoit, R.
Allacma fusca var. albiceps ♀ from France
2008.11.03 © Lebeaux, P.
Allacma fusca var. albiceps ♀ from France
Female sub-anal appendages (*)
2008.11.03 © Lebeaux, P.
Allacma fusca var. albiceps from France
2008.11.03 © Lebeaux, P.
Allacma fusca var. albiceps from France
2008.11.03 © Lebeaux, P.
Allacma fusca var. albiceps from France
2008.11.03 © Lebeaux, P.
Allacma fusca var. albiceps from La Palma
2012.12.04 © Barton, T.
Allacma fusca var. albiceps from Portugal
2020.11.14 © Delgado, F.
Allacma fusca var. albiceps from Portugal
Eversed collophore vesicle
2020.12.08 © Almeida, J.
Allacma fusca var. albiceps from France
2021.03.09 © Nogues, Be.
Allacma fusca var. albiceps from France
2021.03.09 © Nogues, Be.


            Syn.: Sminthurus fuscus var. pustulatus Krausbauer, 1901
             Syn.: Sminthurus fuscus ssp. pustulatea(sic) Salmon, JT, 1964:609 ISS
Allacma fusca ♀ (= var. pustulata) from France
After Nayrolles P, 1994 Fig.2
Allacma fusca ♀ (= var. pustulata) from Sweden
2005 © Hall, K.
Allacma fusca ♀ (= var. pustulata)
2005 © Bellmann, H.
 
Allacma fusca ♀ (= var. pustulata) from France
2005 © Locasso, S.
Allacma fusca ♀ (= var. pustulata) from Denmark
2006 © Fjellberg, A.
Allacma fusca ♀ (= var. pustulata) from France
2007.06.09 © Lebeaux, P.
Allacma fusca ♀ (= var. pustulata) from France
2007.06.09 © Lebeaux, P.
Allacma fusca ♀ (= var. pustulata) from France
2007.06.09 © Lebeaux, P.
Allacma fusca ♀ (= var. pustulata) from France
2007.07.19 © Lebeaux, P.
Allacma fusca ♀ (= var. pustulata) from France
2007.07.20 © Lebeaux, P.
Allacma fusca ♀ (= var. pustulata) from France
2007.08.08 © Lebeaux, P.
Allacma fusca ♀ (= var. pustulata) from France
2007.08.08 © Lebeaux, P.
Allacma fusca ♀ (= var. pustulata) from France
2007.09.25 © Lebeaux, P.
Allacma fusca ♀ (= var. pustulata) from Germany
2008.07.30 © Pageler, J.
Allacma fusca ♀ (= var. pustulata) from Germany
2008.07.30 © Pageler, J.
Allacma fusca ♀ (= var. pustulata) from the UK
2008.08.23 © Barton, T.
Allacma fusca ♀ (= var. pustulata) from Ukraine
2009.06.14 © Loboda., B
Allacma fusca ♀ (= var. pustulata) from Sweden
2011.09.25 © Hall, K.
Allacma fusca ♀ (= var. pustulata) from the UK
2013.08.31 © Nurcombe, E.
Allacma fusca ♀ (= var. pustulata) from the UK
2013.09.29 © Nurcombe, E.
Allacma fusca ♀ (= var. pustulata) from Germany
2014.08.10 © Beck, K.
Allacma fusca ♀ (= var. pustulata) from Germany
2014.08.17 © Beck, K.
Allacma fusca ♀ (= var. pustulata) from the UK
2015.01.10 © Nurcombe, E.
Allacma fusca ♀ (= var. pustulata) from the UK
2015.10.07 © Ridgen, D.
Allacma fusca ♀ (= var. pustulata) from Canada
2015.10.20 © de Repentigny, L.-G.
Allacma fusca ♀ (= var. pustulata) from Belgium
2016.08.31 © Huskens, M.-L.
Allacma fusca ♀ (= var. pustulata) from the UK
2017.07.09 © Gardener, A.
Allacma fusca ♀ (= var. pustulata) from Belgium
2017.07.16 © Huskens, M.-L.
Allacma fusca ♀ (= var. pustulata) from Ukraine
2018.03.09 © L'dov, S.
Allacma fusca ♀ (= var. pustulata) from Ukraine
Female genital papilla and sub-anal appendage
2018.03.09 © L'dov, S.
Allacma fusca ♀ (= var. pustulata) from Belgium
2020.09.05 © San Martin, G.


            Syn.: Allacma fusca var. picea Latzel, R, 1917:251
Allacma fusca f. picea from Sweden
2006 © Hall, K.
Allacma fusca f. picea from Holland
2016.10.03 © Belgers, D.


            Syn.: Allacma fusca var. pardalina Latzel, R, 1917:251
            Syn.: Allacma fusca ab. alba Stach, J, 1956:223
             Syn.: Allacma fusca ab. u-signata(sic) Stach, J, 1956:223 ISS
Allacma ab. usignata fusca from France
2005 © Locasso, S.
Allacma fusca ab. usignata from the USA
2006 © Murray, T.
Allacma fusca ab. usignata from Germany
2006.09.16 © Kohl, F.
Allacma fusca ab. usignata from Belgium
2007.06.30 © Janssens, F.
Allacma fusca ab. usignata from Germany
2007.07.18 © Peters, J.
Allacma fusca ab. usignata from Germany
2007.07.18 © Peters, J.
Allacma fusca ab. usignata from the UK
2007.08.03 © Botting, J.
Allacma fusca ab. usignata from Czechia
2007.08.11 © Krásenský P.
Allacma fusca ab. usignata from the UK
2007.08.25 © Kilford., B
Allacma fusca ab. usignata from Czechia
2007.09.09 © Krásenský P.
Allacma fusca ab. usignata from Czechia
2007.09.09 © Krásenský P.
Allacma fusca ab. usignata from Czechia
2007.09.09 © Krásenský P.
Allacma fusca ab. usignata from the UK
2007.10.12 © Kilford., B
Allacma fusca ab. usignata from Holland
2007.10.13 © Baas, A.H.
Allacma fusca ab. usignata from the UK
2008.02.09 © Campbell, A.
Allacma fusca ab. usignata from the UK
2008.07.10 © Robertson, A.
Allacma fusca ab. usignata from the USA
Pale form
2008.08.09 © Murray, T.
Allacma fusca ab. usignata ♀ from the USA
2008.08.10 © Murray, T.
Allacma fusca ab. usignata from Portugal
2008.09.23 © Almeida, J.
Allacma fusca ab. usignatafrom Holland
2008.10.21 © Baas, A.H.
Allacma fusca ab. usignata from the USA
2009.06.24 © Murray, T.
Allacma fusca ab. usignata from Belgium
2009.09.25 © Mattelart, O.
Allacma fusca ab. usignata ♀ from Sweden
2009.12.23 © Halmen, J.
Allacma fusca ab. usignata ♀ from Sweden
Female sub-anal appendages (*)
2009.12.23 © Halmen, J.
Allacma fusca ab. usignatafrom Holland
2013.08.08 © van Duinen, J.
Allacma fusca ♀ ab. usignata from Canada
2015.10.20 © de Repentigny, L.-G.
Allacma fusca ab. usignata ♀ from the UK
Genital papilla (*) and sub-anal appendage (saa)
2019.10.09 © David W.
Allacma fusca ab. usignata from the UK
Ready to moult
2020.07.29 © Hodgson, J.
Allacma fusca ab. usignata from the UK
Ready to moult
Polygonal reticulate epidermal pattern
2020.07.29 © Hodgson, J.
Allacma fusca ab. usignata from the UK
Ready to moult
Polygonal epidermal cells
2020.07.29 © Hodgson, J.


            Syn.: Sphyrotheca stachi Jeannenot, 1957



Podura fusca from Sweden
After de Geer, C., 1743 Tab.VII fig.2.
Podura fusca from Sweden
After de Geer, C., 1751 Tab.VII fig.2.
Smynthurus(sic) fuscus from Switzerland
After Nicolet H, 1842 Pl.9 Fig.10
Smynthurus(sic) fuscus from the UK
After Lubbock J, 1873 Pl.2
Allacma fusca juv. from France
After Nayrolles P, 1994 Fig.1
Allacma fusca from Russia
2006 © Macroclub.ru.
Allacma fusca from the UK
2007.05.14 © Cornwall, N.J.
Allacma fusca from the UK
2007.05.14 © Cornwall, N.J.
Allacma fusca from the UK
2007.05.14 © Cornwall, N.J.
Allacma fusca from Czechia
2007.06.12 © Krásenský, P.
Allacma fusca from Czechia
2007.06.12 © Krásenský, P.
Allacma fusca from the UK
2007.07.01 © Cornwall, N.J.
Allacma fusca juv. from France
2008.03.24 © Lebeaux, P.
Allacma fusca from the UK
Colony of grazing specimens
2008.06.25 © Robertson, A.
Allacma fusca from the UK
2008.06.25 © Robertson, A.
Allacma fusca from Czechia
2008.07.01 © Krásenský, P.
Allacma fusca from Czechia
2008.07.01 © Krásenský, P.
Allacma fusca from Germany
2008.07.22 © Pageler, J.
Allacma fusca from the UK
2008.07.27 © Michaelson, J.
Allacma fusca from the UK
2008.08.07 © Farrell, S.
Allacma fusca from the UK
2008.09.26 © Campbell, A.
Allacma fusca from the Netherlands
2008.10.21 © Baas, A.H.
Allacma fusca from Italy
After Dallai R, 2009 Fig.1B
Allacma fusca from the USA
2009.06.24 © Murray, T.
Allacma fusca from Belgium
On Podarcis muralis
2009.11.26 © Devalez, J.
Allacma fusca from Belgium
2009.12.13 © D'Hert, D.
Allacma fusca juv. from Holland
2011.04.10 © van Duinen, J.
Allacma fusca from Holland
2011.07.11 © van Duinen, J.
Allacma fusca from Sweden
Bulby post-antennal organ (PAO)
2011.09.21 © Hall, K.
Allacma fusca from Sweden
Thoracic epicuticular superstructure
2011.09.21 © Hall, K.
Allacma fusca from Sweden
Mucro
2011.09.21 © Hall, K.
Allacma fusca from the UK
Facial chaetotaxy
2011.11.10 © Nurcombe, E.
Allacma fusca juv. from the UK
2012.04.19 © Nurcombe, E.
Allacma fusca juv. from the UK
2012.04.20 © Nurcombe, E.
Allacma fusca juv. from Holland
2012.04.26 © van Duinen, J.
Allacma fusca juv. from the UK
2012.04.30 © Nurcombe, E.
Allacma fusca from the UK
2012.06.18 © Nurcombe, E.
Allacma fusca from the UK
Post-antennal seta (*)
2012.06.18 © Nurcombe, E.
Allacma fusca from the UK
Composed eye with 8 ocelli
2012.09.05 © Nurcombe, E.
Allacma fusca juv. from the UK
Annulated 4th antennal segment
2013.04.21 © Murray, A.
Allacma fusca juv. from the UK
Bent dorsal macrosetae
2013.05.13 © Murray, A.
Allacma fusca from the UK
Facial chaetotaxy
2013.08.20 © Phillips, E.
Allacma fusca from the UK
Facial and antennal chaetotaxy
2013.08.20 © Phillips, E.
Allacma fusca from Germany
Short bulby post-antennal seta (PAO)
2014.08.21 © Kluck, A.
Allacma fusca from the UK
2013.08.27 © Nurcombe, E.
Allacma fusca from the UK
Spiky macrosetae
2013.09.14 © Nurcombe, E.
Allacma fusca juv. from Germany
2014.04.06 © Beck, K.
Allacma fusca from Germany
With phoretic Acari nymphs
2014.06.21 © Beck, K.
Allacma fusca ♀ from Germany
Genital papilla (*) and sub-anal appendages (**)
2014.08.07 © Kluck, A.
Allacma fusca ♀ from Germany
Genital papilla and sub-anal appendages (*)
2014.08.07 © Kluck, A.
Allacma fusca from Holland
With phoretic protonymph Acari
2014.08.17 © van Duinen, J.
Allacma fusca ♂ from Germany
Raised genital papilla (*)
2014.09.04 © Kluck, A.
Allacma fusca from the UK
Short bulby post-antennal seta (PAO)
2015.02.03 © Nurcombe, E.
Allacma fusca ♀ from Denmark
SEM Mucro
2016.08.01 © Godeiro, N.N.
Allacma fusca ♀ from Denmark
SEM Unguis with tunica + pseudonychia
2016.08.01 © Godeiro, N.N.
Allacma fusca ♀ from Denmark
SEM Cephalic macrosetae with microscales
2016.08.01 © Godeiro, N.N.
Allacma fusca ♀ from Denmark
SEM Epicuticular ultrastructure
2016.08.08 © Godeiro, N.N.
Allacma fusca ♀ from Denmark
SEM Abdominal dorsal macrosetae + bothriotrichum
2016.08.08 © Godeiro, N.N.
Allacma fusca ♀ from Denmark
SEM Subanal appendages
2016.08.08 © Godeiro, N.N.
Allacma fusca from Belgium
With everted warted vesicle of collophore
2016.08.30 © Huskens, M.-L.
Allacma fusca from Belgium
2017.07.16 © Huskens, M.-L.
Allacma fusca from Belgium
Ant.2 and ant.3 with strong macrosetae (*)
2017.07.16 © Huskens, M.-L.
Allacma fusca ♀ from Germany
2020.07.16 © Kluck, U.
Allacma fusca ♀ from Germany
Sub-anal appendage on papilla (*)
2020.07.16 © Kluck, U.

Species gallica (Carl, 1899) Palissa, A in Brohmer, P, Ehrmann, P & Ulmer, G, 1964:275


            Syn.: Smynthurus(sic) punctatus Lucas, H, 1846:255
Smynthurus(sic) punctatus from Algeria
After Lucas H, 1849 Pl.2 Fig.4
Smynthurus(sic) punctatus from Algeria
After Lucas H, 1849 Pl.2 Fig.4e
Smynthurus(sic) punctatus from Algeria
Ocelli A-H; D smaller
Modified after Lucas H, 1849 Pl.2 Fig.4b
Smynthurus(sic) punctatus from Algeria
Ant.4 with long basal + 9 short apical subdivisions
Modified after Lucas H, 1849 Pl.2 Fig.4c
Smynthurus(sic) punctatus from Algeria
Furca
After Lucas H, 1849 Pl.2 Fig.4d


             Syn.: Sminthurus punctulatus(sic) Grigoreva, 1965 ISS
            Syn.: Sminthurus punctatus Bretfeld, G, 1999:177
            Syn.: Sminthurus punctatus Bretfeld, G, 1999:178 = juv.?
            Syn.: Sminthurus punctatus Bretfeld, G, 2001:130 = juv.
Remark by Janssens, F. 2019.02.03: Ant.4 subdivision and ocelli do not fit Allacma but Bourletiellidae.




             Syn.: Sminthurus gallica(sic) Salmon, 1964 ISS
             Syn.: Sminthurus gallica(sic) Richards, 1968 ISS
Allacma gallica from France
2006 © Lavoué., B
Allacma gallica from France
2007.06.09 © Lebeaux, P.
Allacma gallica ♂ from France
2007.06.09 © Lebeaux, P.
Allacma gallica from France
2007.06.09 © Lebeaux, P.
Allacma gallica from France
2007.06.09 © Lebeaux, P.
Allacma gallica from France
2008.mm.dd © Lebeaux, P.
Allacma gallica from France
2008.mm.dd © Lebeaux, P.
Allacma gallica from Germany
In terrarium
2014.09.13 © Jäschke, T.
Allacma gallica from Germany
In terrarium
2014.09.13 © Jäschke, T.
Allacma gallica from France
Face
2018.11.04 © Picard, J.
Allacma gallica from France
Face
2018.11.04 © Picard, J.
Allacma gallica from France
Unguis with broad tunica (*)
2018.11.04 © Picard, J.
Allacma gallica from France
Central ocelli D smallest
2018.11.04 © Picard, J.
Allacma gallica from France
Post-antennal organ (PAO)
2018.11.04 © Picard, J.
Allacma gallica from France
2018.11.27 © Picard, J.
Allacma gallica from France
2018.11.27 © Picard, J.
Allacma gallica from France
2019.08.31 © Dejoux, P.
Allacma gallica from France
2019.11.27 © Dejoux, P.
Allacma gallica ♂ from France
Male genital papilla
2019.12.07 © Dejoux, P.



Species koreana Yosii, R et Lee, C-E, 1963:14

Allacma koreana from Korea
After Yosii R & Lee C-E, 1963 Fig.8A

            Syn.: Allacma koreana f. leucocephala Yosii, R et Lee, C-E, 1963:15
Allacma koreana from Korea
After Yosii R & Lee C-E, 1963 Fig.8B




Allacma koreana? from Taiwan
2021.11.22 © Cheng, H.-J.
Allacma koreana? from Taiwan
2021.11.22 © Cheng, H.-J.
Allacma koreana? from Taiwan
2021.11.23 © Cheng, H.-J.
Allacma koreana? from Taiwan
2021.12.06 © Cheng, H.-J.
Allacma koreana? from Taiwan
2021.12.06 © Cheng, H.-J.
Allacma koreana? from Taiwan
2021.12.08 © Cheng, H.-J.
Allacma koreana? from Taiwan
2021.12.08 © Cheng, H.-J.

Species plumosa Handschin, 1926, s.d., -

Species plumosetosa Handschin, 1926, s.d., -

Species setosa (von Olfers, 1907 -) Handschin, 1926, s.d., -


Subgenus Trochopilosa Christiansen, K et Bellinger, P, 1998:1353
Species purpurescens (MacGillivray, AD, 1894:109) Christiansen & Bellinger, 1980, t.t.

            Syn.: Sminthurus fraternus Banks, 1899
            Syn.: Sminthurus nigripes Banks, 1899


             Syn.: Sminthurus purpurascens(sic) Yosii, R et Lee, C-E, 1963:15 ISS
Sminthurus (Allacma) pupurescens from the USA
After Snider RJ & Calandrino FJ, 1987 Fig.49
Sminthurus (Allacma) pupurescens from the USA
After Snider RJ & Calandrino FJ, 1987 Fig.50

             Syn.: Allacma purpurascens(sic) Christiansen, K et Bellinger, P, 1998:1354 ISS



Allacma (Trochopilosa) purpurescens
from the USA
2005.08.19 © Murray, T.
Allacma (Trochopilosa) purpurescens
from the USA
2007.07.20 © Murray, T.
Allacma (Trochopilosa) purpurescens
from the USA
2007.09.18 © Murray, T.
Allacma (Trochopilosa) purpurescens
from the USA
2008.06.18 © Cowen, R.
Allacma (Trochopilosa) purpurescens
from the USA; everted ventral tube vesicle
2008.06.20 © Cowen, R.


Genus Sminthurus Latreille in Sonnini, [1802] key

  Syn.: ... Smynthurus(sic) Latreille in Sonnini, [1802] IOS
  Syn.: Seconde famille Podures de Geer
  Syn.: ... Podura Linnæus, 1746, partim
  Syn.: Aptera Podura Linnæus, 1758
  Syn.: Sminthurides(sic) Podura Elditt, HL, 1854:42 partim
  Syn.: Sminthurus Setosi Reuter, OM in Lindberg, SO, 1881:203
  Syn.: ... Eusminthurus Börner, 1900, partim
  Syn.: Sminthurinus Krausbaueri(sic) Börner, C, 1901:708 = 1st instar Sminthurus IOS
  Syn.: Sminthurinus Krausbaueri(sic) Denis, J-R, 1933:283 ISS
  Syn.: Sminthurinus Krausbaueri(sic) Denis, J-R, 1933:289 ISS
  Syn.: Sminthurinus Krausbaueri(sic) Denis, J-R, 1933:291 ISS
  Syn.: P.(sic) Krausbaueri(sic) Denis, J-R, 1933:293 ISS
  Syn.: Sminthurus multifasciatus Ionesco, 1913
  Syn.: Collembola Papyrius(sic) Salmon, JT, 1964:597 partim ISS
  Syn.: Pararrhopalites neomexicanus Scott, 1964 partim = juv. Sminthurus
  Syn.: Sminthurus sp. Betsch, 1977 = juv
  Syn.: Sminthurus sp. Bretfeld, G, 1999:108 = 1st instar



  Syn.: Podurelles(sic) Smynthurus(sic) Nicolet, H in Desor, E, 1841:384 IOS
  Syn.: Podurelles(sic) (Corps globuleux) Smynthurus(sic) Nicolet, H, 1842:80 IOS
  Syn.: Thysanoures(sic) Smynthurus(sic) Lucas, H in Desmarest, E, 1846:110 IOS
  Syn.: Smynthurelles(sic) Smynthurus(sic) Nicolet, H, 1847:357 IOS
  Syn.: Smynthurelles(sic) Smynthurus(sic) Lucas, H, 1849:381 IOS
  Syn.: Podurellen(sic) Smynthurus(sic) Elditt, HL, 1854:38 IOS
  Syn.: Podurellen(sic) Smynthurus(sic) Elditt, HL, 1854:39 IOS
  Syn.: Podurellen(sic) Smynthurus(sic) Elditt, HL, 1854:40 IOS
  Syn.: Sminthurides(sic) Smynthurus(sic) Elditt, HL, 1854:41 IOS
  Syn.: Sminthurides(sic) Smynthurus(sic) Elditt, HL, 1854:42 IOS
  Syn.: Podurelles(sic) Smynthurus(sic) Elditt, HL, 1854:43 IOS
  Syn.: Podurida(sic) Smynthurus(sic) Kolenati, FA, 1858:130 IOS
  Syn.: Podurida(sic) Smynthurus(sic) Kolenati, FA, 1858:134 IOS
  Syn.: Poduridæ(sic) Smynthurus(sic) Packard, AS, 1869:624 IOS
  Syn.: Poduridæ(sic) Smynthurus(sic) Packard, AS, 1872:624 IOS
  Syn.: Thysanura Smynthurus(sic) Lubbock, J, 1873:100 IOS
  Syn.: Collembola Smynthurus(sic) Lubbock in 3089:14 IOS
  Syn.: Thysanura Smynthurus(sic) MacGillivray, AD, 1893:127 partim IOS
  Syn.: Smynthuridæ(sic) Smynthurus(sic) MacGillivray, AD, 1894:108 partim IOS
  Syn.: Collembola Smynthurus(sic) Dallas, WS in Duncan, PM, 1896:149 IOS
  Syn.: Smynthuri(sic) Smynthurus(sic) Folsom, JW, 1896:449 partim IOS
  Syn.: Smynthurinae(sic) Smynthurus(sic) Kloet, GS et Hincks, WD, 1945:9 IOS
  Syn.: Sminthuridae Smindiurus(sic) Castaño-Meneses, G, Mejía-Recamier, BE, Catellanos-Vargas, I, Estrada, DA, Varela-Gómez, M et Vences-Blanco, MO, 2001:21 sp ISS
Sminthurus sp. from Russia
2006 © Macroclub.ru.

Remark by ICZN. 2007:759: Placed on the Official List of Names in Zoology. Opinion 291.

Sminthurus sp. from France
2008.05.01 © Lebeaux, P.
Sminthurus viridis from Lithuania
With parasitic larva of Acari:Erythraeoidea
2009.07.05 © Joanitis, L.
Sminthurus sp. from the USA
2011.05.17 © Loser, I.
Sminthurus sp. from Greece
2011.10.31 © Cladoniophile., B
Sminthurus sp. from the USA
2011.12.12 © McCulloch, J.
Sminthurus sp.
Fossil specimen in amber
2012.02.13 © Hendericks, H.
Sminthurus sp. from Alaska
Depositing eggs
2012.08.10 © Sikes, D.
Sminthurus hispanicus from Ibiza
Long and annulated 4th antennal segment
2012.11.11 © Beer, R.
Sminthurus sp. from La Palma
2012.12.03 © Barton, T.
Sminthurus sp. from La Palma
2012.12.03 © Barton, T.
Sminthurus sp. from La Palma
2012.12.04 © Barton, T.
Sminthurus sp. from La Palma
2012.12.04 © Barton, T.
Sminthurus sp. from La Palma
2012.12.06 © Barton, T.


  Syn.: Sminthuridae Smithurus(sic) Cruz-Leal, JI, Escalante-Jiménez, AL, Salgado-Ortiz, J et Palacios-Vargas, JG, 2013:1667 ISS
Sminthurus sp. from France
2016.11.28 © Cassin, M.R.
Sminthurus sp. ♂ from Israel
2018.03.03 © Lerman, D.
Sminthurus sp. ♂ from Israel
Male genital papilla (*)
2018.03.03 © Lerman, D.

  Syn.: Sminthurus plus(sic) Leo, C, Carapelli, A, Cicconardi, F, Frati, F et Nardi, F, 2019:12 lapsus
Sminthurus? sp. from Chile
2019.08.19 © Guzmán, V.V.
Sminthurus? sp. from Chile
2019.08.19 © Guzmán, V.V.
Sminthurus? sp. from Chile
2019.08.19 © Guzmán, V.V.
Sminthurus? sp. from Chile
2019.08.19 © Guzmán, V.V.
Sminthurus sp. from Taiwan
2022.01.06 © Cheng, H.-J.
Sminthurus sp. ♂ from Taiwan
Depositing spermatophore
2022.02.07 © Cheng, H.-J.






Species sp.1 Simón-Benito, Ortuño & Espantaleón, 2002 -

Species sp.2 Janssens F & Cowen R, 2007

Sminthurus sp. nov. from the USA
2007.04.30 © Cowen, R.
Sminthurus sp. nov. from the USA
2007.04.30 © Cowen, R.
Sminthurus sp. nov. from the USA
2007.04.30 © Cowen, R.
Sminthurus sp. nov. from the USA
2007.05.03 © Cowen, R.
Sminthurus sp. nov. from the USA
2007.05.04 © Cowen, R.
Sminthurus sp. nov. from the USA
2007.05.08 © Cowen, R.
Sminthurus sp. nov. from the USA
2007.05.10 © Cowen, R.

Species sp.3 Janssens F & Cowen R, 2009

Sminthurus sp. nov. from the USA
With parasite larval erythraeid Leptus sp.
2009.05.23 © Cowen, R.
Sminthurus sp. nov. from the USA
2009.05.23 © Cowen, R.

Species sp.4 Cruz-Leal, JI, Escalante-Jiménez, AL, Salgado-Ortiz, J et Palacios-Vargas, JG, 2013:1667

Species sp.5 Janssens F & Young D, 2015

Sminthurus sp. nov. from the USA
2015.03.30 © Young, D.
Sminthurus sp. nov. from the USA
2015.03.30 © Young, D.

Species sp.6 Janssens F & Murray A, 2015

Sminthurus sp. nov. from Canada
2015.07.07 © Murray, A.
Sminthurus sp. nov. from Canada
2015.07.07 © Murray, A.
Sminthurus sp. nov. from Canada
2015.07.07 © Murray, A.

Species sp.7 Janssens F & Fleming KR, 2017

Sminthurus sp. nov. from the USA
2012.02.21 © Engel, V.
Sminthurus sp. nov. juv. from the USA
2012.06.14 © Loser, I.
Sminthurus sp. nov. from the USA
2013.01.29 © Fleming, K.
Sminthurus sp. nov. from the USA
2013.01.29 © Fleming, K.
Sminthurus sp. nov. from the USA
2014.01.18 © Klockers, W.B.
Sminthurus sp. nov. from the USA
2014.01.18 © Klockers, W.B.
Sminthurus sp. nov. from the USA
2017.02.03 © Fleming, K.
Sminthurus sp. nov. from the USA
Everted tubelike vesicle of collophore
2017.02.03 © Fleming, K.
Sminthurus sp. nov. from the USA
2018.01.07 © Abbott, J.C.
Sminthurus sp. nov. from the USA
Ant.4 with, 20 subdivisions
2018.01.07 © Abbott, J.C.
Sminthurus sp. nov. juv. from the USA
2020.07.09 © Cam, C.
Sminthurus sp. nov. juv. from the USA
Pale form
2020.07.09 © Cam, C.
Sminthurus sp. nov. from the USA
2020.07.10 © Cam, C.
Sminthurus sp. nov. juv. from the USA
Feeding instar
2020.07.12 © Cam, C.
Sminthurus sp. nov. from the USA
2020.07.20 © Cam, C.
Sminthurus sp. nov. from the USA
2020.07.20 © Cam, C.

Species sp.9 Janssens F & Colombo M, 2021

Sminthurus sp. nov. from Italy
2021.02.03 © Colombo, M.
Sminthurus sp. nov. from Italy
2021.02.03 © Colombo, M.
Sminthurus sp. nov. from Italy
2021.02.03 © Colombo, M.

Species abei Yoshii, 1992

Species adamsi Wray, 1967

Species adirondackus Maynard, EA, 1951:270

Sminthurus adirondackus from the USA
After Maynard, E.A., 1951 Pl.XXIV Fig.417.
Sminthurus adirondackus from the USA
After Maynard, E.A., 1951 Pl.XXIV Fig.416.
Sminthurus adirondackus from the USA
After Maynard, E.A., 1951 Pl.XXIV Fig.415.

Species baulastinus Harvey, 1848

Species betae Westwood, 1849

Species bicolor Parona, 1884

Species bivittatus Snider, RJ, 1985:575

Sminthurus bivittatus from the USA
After Snider RJ, 1985 Fig.1
Sminthurus bivittatus from the USA
After Snider RJ, 1985 Fig.2
Sminthurus bivittatus from the USA
2006.02.02 © Schimming, L.
Sminthurus bivittatus from the USA
2007.02.26 © Montgomery, G.
Sminthurus bivittatus from the USA
2008.11.27 © Justis, S.
Sminthurus bivittatus from the USA
2008.11.27 © Justis, S.
Sminthurus bivittatus from the USA
2008.11.27 © Justis, S.
Sminthurus bivittatus from the USA
2008.11.27 © Justis, S.
Sminthurus bivittatus from the USA
2008.11.27 © Justis, S.
Sminthurus bivittatus from the USA
2008.12.10 © Kermit, J.
Sminthurus bivittatus from the USA
2008.12.27 © Justis, S.
Sminthurus bivittatus from the USA
2008.12.27 © Justis, S.
Sminthurus bivittatus from the USA
2009.01.16 © Kermit, J.
Sminthurus bivittatus juv. from the USA
2009.01.24 © Justis, S.
Sminthurus bivittatus juv. from the USA
2009.01.24 © Justis, S.
Sminthurus bivittatus from the USA
2009.05.01 © Taft, M.
Sminthurus bivittatus from the USA
2009.05.01 © Taft, M.
Sminthurus bivittatus from the USA
2009.05.01 © Taft, M.
Sminthurus bivittatus from the USA
2012.12.19 © Zimlich, R.L.
Sminthurus bivittatus from the USA
2012.12.19 © Zimlich, R.L.

Species bourgeoisi Nayrolles, 1995:221

Species bozoulensis Nayrolles, P, 1995:222

Sminthurus bozoulensis from France
After Nayrolles P, 1995 Fig.9

Species brevicornis Koch, CL et Berendt, GC in Berendt, GC, 1854:121, -

Smynthurus(sic) brevicornis from Baltic amber
After Koch CL & Berendt GC, 1845 Tab.XV Fig.131

Species brunneus Harvey, 1900

Species butcheri Snider, RJ, 1969:366

Sminthurus butcheri from the USA
After Snider RJ, 1969 Fig.43

            Syn.: Sminthurus longicornis Schött, 1896
            Syn.: Smynthurus(sic) dorsalis Banks, 1899
Sminthurus dorsalis from the USA
After Maynard EA, 1951 PlateXXXIII Fig.610
Sminthurus butcheri f. dorsalis from the USA
2018.06.28 © Johnson, S.T.
Sminthurus butcheri f. dorsalis from the USA
2021.05.07 © Pavy, R.
Sminthurus butcheri f. dorsalis from the USA
2021.05.16 © Pavy, R.
Sminthurus butcheri f. dorsalis from the USA
2021.05.16 © Pavy, R.

            Syn.: Sminthurus spinulosus Snider, RJ, 1969:372
Sminthurus spinulosus from the USA
After Snider RJ, 1969 Fig.63
Sminthurus butcheri f. spinulosus from the USA
Colouration of immature specimen
2012.06.10 © Fort Lewis Zoology.
Sminthurus butcheri f. spinulosus from the USA
Ventral aspect
2012.06.10 © Fort Lewis Zoology.
Sminthurus butcheri f. spinulosus from the USA
4th antennal segment with 15 annulations
2012.06.10 © Fort Lewis Zoology.
Sminthurus butcheri f. spinulosus from the USA
2017.08.09 © Gross, J.
Sminthurus butcheri f. spinulosus from the USA
2017.08.09 © Gross, J.
Sminthurus butcheri f. spinulosus from the USA
2017.08.09 © Gross, J.
Sminthurus butcheri f. spinulosus from the USA
2018.06.11 © Lee, C.
Sminthurus butcheri f. ?spinulosus? ♀ from the USA
2018.07.20 © Bollich, C.
Sminthurus butcheri f. ?spinulosus? ♀ from the USA
2018.07.20 © Bollich, C.
Sminthurus butcheri f. ?spinulosus? ♀ from the USA
Ventral aspect
2018.07.20 © Bollich, C.

            Syn.: Sminthurus (Sminthurus) banksi Christiansen, K et Bellinger, P, 1981:1197
            Syn.: Sminthurus (Sminthurus) banksi Christiansen, K et Bellinger, P, 1981:1198 =?
            Syn.: Sminthurus (Sminthurus) banksi Christiansen, K et Bellinger, P, 1998:1361



             Syn.: Sminthurus bucheri(sic) Palacios-Vargas, JG, Mejía-Recamier, BE et Cutz-Pool, LQ, 2009:206 ISS
Sminthurus butcheri from the USA
2009.02.11 © Justis, S.
Sminthurus butcheri from the USA
2009.02.11 © Justis, S.
Sminthurus butcheri from the USA
2012.02.06 © Zimlich, R.L.
Sminthurus butcheri from the USA
2018.02.11 © Doby, J



Species capucinus Koch, 1840

Species carolinensis Snider, RJ, 1981:418

Sminthurus carolinensis from the USA
After Snider RJ, 1981 Fig.1
Sminthurus carolinensis from the USA
After Snider RJ, 1981 Fig.2
Sminthurus carolinensis from the USA
2009.01.16 © Fleming, K.
Sminthurus carolinensis from the USA
2012.01.12 © Champagne, C.

Species coeruleus Strebel, 1938

Species cogsonzavi Betsch, J-M, 1977:72

Species denisi Womersley, 1934

            Syn.: Sminthurus longicornis Womersley, 1932, nec Joseph, 1882


Species denticulatus Bretfeld, 2000

Species discolor Stach, 1956

Species doderoii Parona, 1888

             Syn.: Sminthurus doderoi(sic) Stach, 1956,is ISS

Species eisenii Schött, H, 1891:7

            Syn.: Sminthurus (Setosi) Eisenii(sic) Schött, H, 1891:7 IOS
Sminthurus Eisenii(sic) from the USA
After Schött H, 1891 Taf.I Fig.1
Sminthurus Eisenii(sic) from the USA
After Schött H, 1891 Taf.I Fig.2

            Syn.: Sminthurus anomalus Betsch, J-M, 1964:542
            Syn.: Sminthurus medialis Betsch, J-M, 1964:533
            Syn.: Pararrhopalites neomexicanus Scott, 1964 partim =? juvenile



             Syn.: Sminthurus Eisenii(sic) Schött, 1893 ISS
             Syn.: Sminthurus Eisenii(sic) Bertkau, P, 1894:58 ISS
             Syn.: Sminthurus Eiseni(sic) Denis, J-R, 1933:305 ISS
             Syn.: Sminthurus Eiseni(sic) Denis, J-R, 1933:308 ISS
             Syn.: Sminthurus (Sminthurus) eiseni(sic) Christiansen, K et Bellinger, P, 1981:1200 ISS
             Syn.: Sminthurus eisneni(sic) Snider, RJ et Loring, SJ, 1984:37 ISS
             Syn.: Sminthurus eiseni(sic) Snider, RJ et Loring, SJ, 1984:38 ISS
             Syn.: Sminthurus eiseni(sic) Bellinger, PF, 1985:80 ISS
             Syn.: Sminthurus eiseni(sic) Nayrolles, P, 1995:217 ISS
             Syn.: Sminthurus eiseni(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS
Sminthurus ?eisenii? from the USA
2010.06.24 © Boyd, I.
Sminthurus ?eisenii? from the USA
2014.06.24 © Rosenfeld, J.
Sminthurus eisenii from the USA
2020.02.27 © Schneider, K.
Sminthurus eisenii from the USA
2020.02.27 © Schneider, K.
Sminthurus eisenii from the USA
2020.02.27 © Schneider, K.
Sminthurus eisenii from the USA
2020.02.27 © Schneider, K.
Sminthurus eisenii from the USA
2021.12.30 © Stempien, C.





Species fischeri Snider, RJ, 1982:322

Sminthurus fischeri from the USA
After Snider RJ, 1982 Fig.1
Sminthurus fischeri from the USA
After Snider RJ, 1982 Fig.2
Sminthurus fischeri? from the USA
2017.01.28 © Abela, A.
Sminthurus fischeri? from the USA
2017.01.28 © Abela, A.
Sminthurus fischeri from Canada
2021.05.21 © Ellyne.

Species fitchi Folsom, JW, 1896:449

            Syn.: Smynthurus(sic) fitchii(sic) Folsom, JW, 1896:449 IOS
            Syn.: Sminthurus albescens Banks, 1899
            Syn.: Sminthurus argenteornatus Banks, 1899
Sminthurus argenteornatus from the USA
After Maynard EA, 1951 Pl.XXIV Fig.421

            Syn.: Sminthurus trilineatus Banks, 1903
            Syn.: Sminthurus obscurus Mills, HB, 1934:104



             Syn.: Sminthurus Fitchi(sic) Denis, J-R, 1933:305 ISS
Sminthurus fitchi from the USA
After Maynard, E.A., 1951 Pl. XXIV Fig.431
Sminthurus fitchi from the USA
2007.10.03 © Murray, T.
Sminthurus fitchi from the USA
2007.10.03 © Murray, T.
Sminthurus fitchi from the USA
2007.10.03 © Murray, T.
Sminthurus fitchi from the USA
2007.10.03 © Murray, T.
Sminthurus fitchi from the USA
2007.10.03 © Murray, T.
Sminthurus fitchi from the USA
2007.10.03 © Murray, T.
Sminthurus fitchi from the USA
2007.10.03 © Murray, T.
Sminthurus fitchi from the USA
2007.10.28 © Murray, T.
Sminthurus fitchi from the USA
2007.11.02 © Murray, T.
Sminthurus fitchi from the USA
2007.11.04 © Murray, T.
Sminthurus fitchi from the USA
2007.11.04 © Murray, T.
Sminthurus fitchi from the USA
2007.11.04 © Murray, T.
Sminthurus fitchi from the USA
2007.11.04 © Murray, T.
Sminthurus fitchi from the USA
2007.11.04 © Murray, T.
Sminthurus fitchi from the USA
2012.10.05 © Murray, T.
Sminthurus fitchi from the USA
2019.12.23 © Woo, Br.





Species floridanus MacGillivray, AD, 1893:127

             Syn.: Smynthurus(sic) floridana(sic) MacGillivray, AD, 1894:109 ISS

Species ghilarovi Stebaeva, SK, 1966:9

Sminthurus ghilarovi from Siberia
After Stebaeva SK, 1966 Fig.2.1

Species giantensis Baijal & Kohli, 1972

Species hamtaensis Baijal, HN, 1958:360

Sminthurus hamtaensis from the Himalaya
After Baijal HN, 1958 Fig.13A.

Species hispanicus Nayrolles, P, 1995:231

Sminthurus hispanicus from Spain
After Nayrolles P, 1995 Fig.18
Sminthurus hispanicus from Ibiza
2009.03.03 © Beer, R.
Sminthurus hispanicus from Ibiza
2009.03.03 © Beer, R.
Sminthurus hispanicus from Ibiza
2009.03.08 © Beer, R.
Sminthurus hispanicus from Ibiza
2009.04.08 © Beer, R.
Sminthurus hispanicus from Ibiza
2009.11.05 © Beer, R.
Sminthurus hispanicus from Italy
2009.11.08 © Segreto, J.
Sminthurus hispanicus from Ibiza
Reproductive male with raised genital papilla
2011.11.10 © Beer, R.
Sminthurus hispanicus from Ibiza
2011.11.12 © Beer, R.
 
Sminthurus hispanicus juv. from Ibiza
2011.12.16 © Beer, R.
Sminthurus hispanicus juv. from Ibiza
2011.12.30 © Beer, R.
Sminthurus hispanicus juv. from Ibiza
2011.12.30 © Beer, R.
Sminthurus hispanicus juv. from Ibiza
2012.01.08 © Beer, R.
Sminthurus hispanicus juv. from Ibiza
2012.01.25 © Beer, R.
Sminthurus hispanicus from Ibiza
2012.11.04 © Beer, R.
Sminthurus hispanicus from Ibiza
Grooming: cleaning antennae with drop of saliva on claw of frontleg
2013.02.13 © Beer, R.
Sminthurus hispanicus juv. & adult from Ibiza
2013.09.18 © Beer, R.
Sminthurus hispanicus juv. from Ibiza
2013.09.18 © Beer, R.
Sminthurus hispanicus from Ibiza
2013.09.27 © Beer, R.
Sminthurus hispanicus from Ibiza
2013.09.28 © Beer, R.
Sminthurus hispanicus juv. from Ibiza
2013.12.24 © Beer, R.
Sminthurus hispanicus from Ibiza
2014.01.13 © Beer, R.
Sminthurus hispanicus from Ibiza
Pollen feeding
2014.01.20 © Beer, R.
Sminthurus hispanicus juv. from Ibiza
Pollen feeding
2014.02.01 © Beer, R.
Sminthurus hispanicus from Ibiza
Chewing away upper epidermis layer of leaf
2014.10.06 © Beer, R.
Sminthurus hispanicus from Ibiza
Ingesting pollen in a dew drop
2014.10.11 © Beer, R.
Sminthurus hispanicus ♀ & ♂ from Ibiza
♀ upper; ♂ lower
2014.11.02 © Beer, R.
Sminthurus hispanicus from Ibiza
2014.11.16 © Beer, R.
Sminthurus hispanicus juv. from Ibiza
First instar
2014.11.24 © Beer, R.
Sminthurus hispanicus juv. from Ibiza
First instar
2014.12.04 © Beer, R.
Sminthurus hispanicus juv. from Ibiza
First instar
2015.02.02 © Beer, R.
Sminthurus hispanicus from Ibiza
2015.02.05 © Beer, R.
Sminthurus hispanicus from Ibiza
Feeding: damage to leaf surface
2015.04.01 © Beer, R.
Sminthurus hispanicus juv. from Ibiza
2015.10.22 © Beer, R.
Sminthurus hispanicus from Ibiza
Pale coulour form
2015.11.21 © Beer, R.
Sminthurus hispanicus from Ibiza
Greenish coulour form
2015.12.08 © Beer, R.
Sminthurus hispanicus from Ibiza
Grooming body with long tube-like vesicle of collophore
2015.12.08 © Beer, R.
Sminthurus hispanicus from Ibiza
2015.12.16 © Beer, R.
Sminthurus hispanicus from Italy
2016.02.18 © Scozzari, K.
Sminthurus hispanicus from France
2017.02.05 © Tiky, S.
Sminthurus hispanicus from France
2017.02.09 © Tiky, S.
Sminthurus hispanicus from France
2018.01.10 © Garcelon, P.
Sminthurus hispanicus from France
Caudal aspect
2018.05.23 © Picard, J.
Sminthurus hispanicus from France
2019.11.09 © Tcheng, E.
Sminthurus hispanicus from Spain
Caudal aspect
2019.12.06 © Zeeders, M.
Sminthurus hispanicus ♂ from France
2019.11.09 © Joachim, J.
Sminthurus hispanicus ♂ from France
Genital papilla (*)
2019.12.15 © Joachim, J.

Species incisus Snider, 1978

Sminthurus incisus? from the USA
1999 © Van Dyk, J.
Sminthurus incisus? from the USA
1999 © Van Dyk, J.
Sminthurus incisus? from the USA
2017.01.28 © Abela, A.
Sminthurus incisus from the USA
2006 © Murray, T.
Sminthurus incisus from the USA
2007.09.05 © Murray, T.
Sminthurus incisus from the USA
2007.09.05 © Murray, T.
Sminthurus incisus from the USA
2011.07.29 © Bowser, M.
Sminthurus incisus ♂ from the USA
2016.11.24 © Dobbs, Br.
Sminthurus incisus ♂ from the USA
2016.11.24 © Dobbs, Br.
Sminthurus incisus from the USA
2017.09.07 © Solomon.
Sminthurus incisus from the USA
2018.01.23 © Fleming, K.

Species incognitus Snider, RJ et Loring, SJ, 1982:216

Sminthurus incognitus from the USA
After Snider RJ & Loring SJ, 1982 Fig.1
Sminthurus incognitus from the USA
After Snider RJ & Loring SJ, 1982 Fig.2

Species laterculus Koch, 1840

Species leucomelanus Nayrolles, P, 1995:226

Sminthurus leucomelanus from France
After Nayrolles P, 1995 Fig.16
Sminthurus leucomelanus from Czechia
2015.08.02 © Krásenský, P.
Sminthurus leucomelanus from Czechia
2015.08.02 © Krásenský, P.
Sminthurus leucomelanus from Czechia
Macrosetae in black sockets
2015.08.02 © Krásenský, P.
Sminthurus leucomelanus from Czechia
2015.08.21 © Krásenský, P.
Sminthurus leucomelanus from Holland
2016.05.08 © Beck, K.
Sminthurus leucomelanus from Holland
2017.06.13 © van Duinen, J.
Sminthurus leucomelanus from Holland
2017.06.13 © Wieringa, P.
Sminthurus leucomelanus ♂ from France
2018.05.20 © Picard, J.
Sminthurus leucomelanus from France
2018.05.20 © Picard, J.
Sminthurus leucomelanus ♀ from France
Female genital papilla (*)
2019.06.19 © Picard, J.
Sminthurus leucomelanus from Poland
2020.08.09 © Antosik-Zoladek, J.
Sminthurus leucomelanus from Poland
2020.10.01 © Antosik-Zoladek, J.

Species longicornis Koch, CL et Berendt, GC in Berendt, GC, 1854:121, -

Smynthurus(sic) longicornis from Baltic amber
After Koch CL & Berendt GC, 1845 Tab.XV Fig.130

Species longipes Börner, C, 1906:184

Species lusseri Nicolet, H, 1847:360

            Syn.: Smynthurus(sic) Lusseri(sic) Nicolet, H, 1847:360 IOS
Smynthurus Lusseri(sic) from Switzerland
After Nicolet H, 1847 Pl.6 Fig.5




             Syn.: Smynthurus(sic) Lusseri(sic) Lubbock, J, 1873:115 ISS



Species maculatus Tömösváry, 1883

             Syn.: Sminthurus viridis var. maculata(sic) Stach, 1920 ISS
             Syn.: Smynthurus(sic) viridis ssp. maculata(sic) Stach, 1922 ISS
             Syn.: Sminthurus viridis f. maculata(sic) Handschin, E in Dahl, F, 1929:121 ISS
             Syn.: Smynthurus(sic) viridis ssp. maculata(sic) Szent-Ivany, 1941 ISS
             Syn.: Sminthurus viridis ssp. maculata(sic) Schaller, 1951 ISS
             Syn.: Sminthurus viridis f. maculata(sic) Pistor, D, 1955:525 ISS
Sminthurus viridis f. maculata(sic) from Germany
After Pistor D, 1955 Abb.15

Sminthurus maculatus from Poland
After Stach J, 1956 PlateXXXIII Fig.6
Sminthurus nr maculatus from Slovakia
2007.08.29 © Stefanovie, P.
Sminthurus nr maculatus from Hungary
2008.11.08 © Keresztes, G.
Sminthurus nr maculatus juv. from Hungary
2009.09.11 © Keresztes, G.
Sminthurus nr maculatus from Hungary
2009.09.21 © Keresztes, G.
Sminthurus nr maculatus from Hungary
2009.09.21 © Keresztes, G.
Sminthurus maculatus from Hungary
2010.08.12 © Pfliegler, W.
Sminthurus maculatus from Hungary
2013.10.13 © Rahmé, N.
Sminthurus nr maculatus from Spain
2017.01.04 © Schulz, K.
Sminthurus nr maculatus from France
2017.03.25 © Tiky, S.
Sminthurus maculatus from Hungary
2017.10.17 © de Haas, M.
Sminthurus nr maculatus from France
2017.12.07 © Tiky, S.
Sminthurus nr maculatus from France
2017.12.07 © Tiky, S.
Sminthurus nr maculatus from France
2017.12.07 © Tiky, S.
Sminthurus nr maculatus from France
2017.12.07 © Tiky, S.



Species maglicianus Cvijovic, 1986

Species marmorellus Latzel, R, 1917:250

            Syn.: Sminthurus sp. Stach, J, 1956:239


Species medialis Mills, HB, 1934:103

            Syn.: Sminthurus dorsalis Mills, 1930


Sminthurus medialis from the USA
2012.12.07 © Zimlich, R.L.
Sminthurus medialis from the USA
2012.12.18 © Montgomery, G.
Sminthurus medialis from the USA
2012.12.19 © Zimlich, R.L.
Sminthurus medialis from the USA
2015.04.21 © Gallagher, J.
Sminthurus medialis from the USA
2020.09.11 © Ekker, S.
Sminthurus medialis from the USA
2020.12.21 © Lampkin, J.
Sminthurus medialis from the USA
2020.12.21 © Lampkin, J.
Sminthurus medialis from the USA
2020.12.21 © Lampkin, J.
Sminthurus medialis from the USA
2021.01.18 © Ekker, S.
Sminthurus medialis from the USA
2021.01.18 © Ekker, S.
Sminthurus medialis from the USA
2021.01.18 © Ekker, S.
Sminthurus medialis from the USA
2021.01.18 © Ekker, S.
Sminthurus medialis from the USA
2021.01.26 © Ekker, S.
Sminthurus medialis from the USA
2021.03.13 © Ekker, S.
Sminthurus medialis from the USA
2021.03.13 © Ekker, S.

Species melanonotus Uchida, 1938

             Syn.: Sminthurus melanotus(sic) Stach, J, 1956:231 ISS

Species mencenbergae Snider, RJ, 1983:115

Sminthurus mencenbergae from Canada
After Snider RJ, 1983 Fig.1
Sminthurus mencenbergae from Canada
After Snider RJ, 1983 Fig.2
Sminthurus mencenbergae from the USA
After Snider RJ, 1983 Fig.3
Sminthurus mencenbergae from the USA
After Snider RJ, 1983 Fig.4
Sminthurus mencenbergae from the USA
2007.06.13 © Montgomery, G.
Sminthurus mencenbergae from the USA
2008.01.22 © Montgomery, G.
Sminthurus mencenbergae from the USA
2009.05.23 © Trammel, E.
Sminthurus mencenbergae from the USA
2009.05.23 © Trammel, E.
Sminthurus mencenbergae from the USA
2011.03.05 © Montgomery, G.
Sminthurus mencenbergae from the USA
2011.06.15 © Hoyer, R.
Sminthurus mencenbergae from the USA
2011.06.15 © Hoyer, R.
Sminthurus mencenbergae from the USA
2011.06.15 © Hoyer, R.
Sminthurus mencenbergae from the USA
2012.04.14 © Kouri, J.
Sminthurus mencenbergae from the USA
2012.04.14 © Kouri, J.
Sminthurus mencenbergae from the USA
2012.04.14 © Kouri, J.
Sminthurus mencenbergae from the USA
2012.04.23 © Kouri, J.
Sminthurus mencenbergae from the USA
2012.04.23 © Kouri, J.
Sminthurus mencenbergae from the USA
2012.04.23 © Kouri, J.
Sminthurus mencenbergae from the USA
2013.12.26 © Schneider, J.
Sminthurus mencenbergae from the USA
2014.01.11 © Burishkin, J.
Sminthurus mencenbergae from the USA
2015.02.08 © ak.
Sminthurus mencenbergae from the USA
2015.03.31 © Montgomery, G.
Sminthurus mencenbergae from the USA
2017.10.12 © Betros, Be.
Sminthurus mencenbergae from the USA
2017.12.12 © Pugh, D.E.
Sminthurus mencenbergae from the USA
2019.10.18 © Betros, Be.
Sminthurus mencenbergae from the USA
2020.12.11 © Fleming, K.
Sminthurus mencenbergae from the USA
2021.03.15 © Foreman, S.
Sminthurus mencenbergae from the USA
2021.04.16 © Pavy, R.

Species millsi Betsch, J-M, 1965:439

Species montanus Bretfeld, G, 2010:307

Species multipunctatus Schäffer, 1896, B

            Syn.: Sminthurus (Eusminthurus) variegatus Börner, C, 1901:114
            Syn.: Sminthurus ammophilus Börner, 1903
            Syn.: Sminthurus multipunctatus f. albopunctata(sic) Stach, 1922 IOS
             Syn.: Sminthurus multipunctatus f. albopunctata(sic) Handschin, E in Dahl, F, 1929:120 ISS
             Syn.: Sminthurus multipunctatus var. albopunctata(sic) Bretfeld, G, 1999:198 ISS
Sminthurus multipunctatus f. albopunctatus
from Israel
2007.12.26 © Nir, A.

            Syn.: Sminthurus multipunctatus var. viridipunctata(sic) Stach, 1922 IOS
             Syn.: Sminthurus multipunctatus f. viridipunctata(sic) Handschin, E in Dahl, F, 1929:120 ISS
             Syn.: Sminthurus multipunctarus var. viridipunctata(sic) Stach, J, 1956:264 ISS
             Syn.: Sminthurus multipunctatus var. viridipunctata(sic) Bretfeld, G, 1999:198 ISS
            Syn.: Sminthurus multipunctatus var. tristrigata(sic) Stach, J, 1956:265 IOS
Sminthurus multipunctatus var. tristrigata(sic) from Poland
After Stach J, 1956 PlateXXXII Fig.1

             Syn.: Sminthurus multipunctatus var. tristrigata(sic) Bretfeld, G, 1999:198 ISS
Sminthurus multipunctatus var. tristrigatus from France
2019.02.09 © Tiky, S.
Sminthurus multipunctatus var. tristrigatus from France
2019.02.09 © Tiky, S.
Sminthurus multipunctatus var. tristrigatus from France
2019.02.09 © Tiky, S.
Sminthurus multipunctatus var. tristrigatus from France
2019.02.09 © Tiky, S.
Sminthurus multipunctatus var. tristrigatus from France
2019.10.26 © Dejoux, P.
Sminthurus multipunctatus var. tristrigatus from France
2019.10.26 © Dejoux, P.
Sminthurus multipunctatus var. tristrigatus from France
2019.10.26 © Dejoux, P.
Sminthurus multipunctatus var. tristrigatus from France
2019.10.26 © Dejoux, P.




Sminthurus multipunctatus from Poland
After Stach J, 1956 PlateXXXIII Fig.1
Sminthurus multipunctatus from France
After Nayrolles P, 1995 Fig.17
Sminthurus multipunctatus from France
2010.03.09 © Chevaillot, F.
Sminthurus multipunctatus from France
2010.03.09 © Chevaillot, F.
Sminthurus multipunctatus? from France
Grooming using extended tubular vesicle of collophore
2012.10.28 © Lebeaux, P.
Sminthurus multipunctatus? from France
Grooming using a drop of secretion
2012.10.28 © Lebeaux, P.
Sminthurus multipunctatus from France
2020.01.12 © Tiky, S.
Sminthurus multipunctatus from France
2020.01.12 © Tiky, S.
Sminthurus multipunctatus from France
2020.01.12 © Tiky, S.
Sminthurus multipunctatus from France
2020.01.12 © Tiky, S.
Sminthurus multipunctatus from France
Caudal aspect
2020.01.12 © Tiky, S.
Sminthurus multipunctatus ♂ from France
Male genital papilla
2020.01.12 © Tiky, S.
Sminthurus multipunctatus from France
2020.01.12 © Tiky, S.



Species muscicolus Betsch, J-M, 1977:66

Species nigrinus Bretfeld, 2000

Species ochraceus Koch, 1840

Species orientalis Bretfeld, 2000

Species osmeryzskensis Bretfeld, 2000

Species ovaculatus Koch & Berendt, 1854 -

Species packardi Folsom, JW, 1896:385

            Syn.: Smynthurus packardii(sic) Folsom, JW, 1896:385 IOS
            Syn.: Papirius texensis Packard, 1873 partim


             Syn.: Sminthurus Packardi(sic) Denis, J-R, 1933:305 ISS
             Syn.: Sminthurus Packardi(sic) Denis, J-R, 1933:308 ISS
             Syn.: Sminthurus packardii(sic) Folsom, 1934 ISS
Sminthurus (Sminthurus) packardi from the USA
After Christiansen K & Bellinger P, 1981 Fig.1009A

             Syn.: Sminthurus (Sminthurus) packardii(sic) Christiansen, K et Bellinger, P, 1998:1369 ISS
Sminthurus packardi from the USA
2015.04.30 © Frish, L.





Species pardalinus Betsch, J-M, 1977:69

Sminthurus pardalinus from Mongolia
After Betsch J-M, 1977 Fig.2B

Species parvulus Ritter, 1911

Species pinetorum Koch, 1840

Species primorskiyensis Bretfeld, 2002

Species pseudofuscus Schött, 1901

Species pseudoviolaceus Ritter, 1911

Species rosai Arlé, 1939

Species rubidipunctatus Bretfeld, 2000

Species sagittus Loring, SJ et Snider, RJ, 1983:185

            Syn.: Sminthurus sagitta(sic) Loring, SJ et Snider, RJ, 1983:185 IOS
Sminthurus sagitta(sic) from the USA
After Loring SJ & Snider RJ, 1983 Fig.1
Sminthurus sagitta(sic) from the USA
After Loring SJ & Snider RJ, 1983 Fig.2


            Syn.: Sminthurus sagitta(sic) Poole, RW in Poole, RW et Gentili, P, 1997:56 IOS
Sminthurus sagittus? from the USA
2017.01.28 © Abela, A.
Sminthurus sagittus? from the USA
2017.01.28 © Abela, A.
Sminthurus sagittus? from the USA
2017.01.28 © Abela, A.



Species schoetti Salmon, JT, 1964:598

            Syn.: Sminthurus longicornis Schött, 1896, nec Joseph, 1882


Species serratomucronatus Grinbergs, A, 1962:65

Species serrulatus Börner, C, 1909:127

            Syn.: Sminthurus serrulatus var. trimaculata Börner, 1909


Sminthurus serrulatus from Japan
After Yosii R, 1970 Fig.17A
Sminthurus serrulatus from Japan
2018.10.12 © Hara, Y.
Sminthurus serrulatus from Japan
2018.10.12 © Hara, Y.
Sminthurus serrulatus from Japan
2018.10.12 © Hara, Y.
Sminthurus serrulatus from Japan
2018.10.12 © Hara, Y.
Sminthurus serrulatus from Japan
2018.10.12 © Hara, Y.
Sminthurus ?serrulatus? from Taiwan
2022.02.09 © Cheng, H.-J.
Sminthurus ?serrulatus? from Taiwan
2022.02.09 © Cheng, H.-J.

Species sexpunctatus Becker, 1905

Species stachi Betsch, J-M, 1977:75

Sminthurus stachi from Mongolia
After Betsch J-M, 1977 Fig.2C

Species succineus Stach, J, 1922:55, -

Sminthurus succineus from Baltic amber
After Stach J, 1922 Taf.3 Fig.1
Sminthurus cf. succineus from Baltic Amber
2012.01.23 © Henderickx, H.

Species tristrigatus Stach, 1956

Species variegatus Tullberg, T, 1877:29

Sminthurus variegatus from the Arctics
After Tullberg, T., 1877 Tafl.8 fig.1.
Sminthurus variegatatus from the Arctics
After Schött H, 1893 Taf.I Fig.6

Species virginidari Wray, 1948

Species viridis (Linnæus, C, 1758:608) Latreille in Sonnini, [1802-03], t.t., B

            Syn.: Smynthurus(sic) lupulinæ Bourlet, 1843
            Syn.: Sminthurus viridis var. nigro-maculatus(sic) Tullberg, 1871 IOS
            Syn.: Sminthurus viridis Schött, 1893 =?
            Syn.: Sminthurus viridis Hammer, 1953 =?
            Syn.: Sminthurus viridis var. nigro-maculatus(sic) Tullberg, T, 1872:30 IOS
            Syn.: Sminthurus viridis ssp. nigro-maculatus(sic) Parona, 1884 IOS
             Syn.: Sminthurus viridis ssp. nigromaculata(sic) Schäffer, 1897 ISS
             Syn.: Sminthurus viridis ssp. nigromaculata(sic) Scherbakow, 1898 ISS
            Syn.: Sminthurus viridis ssp. nigro-maculatus(sic) Börner, 1900 IOS
             Syn.: Sminthurus viridis ssp. nigromaculata(sic) Axelson, 1904 ISS
            Syn.: Sminthurus viridis ssp. nigro-maculatus(sic) Axelson, 1905 IOS
             Syn.: Sminthurus viridis ssp. nigromaculata(sic) Schille, 1908 ISS
             Syn.: Sminthurus viridis ssp. nigromaculata(sic) Schille, 1912 ISS
             Syn.: Sminthurus viridis ssp. nigromaculata(sic) Stach, 1920 ISS
             Syn.: Sminthurus viridis f. nigromaculata(sic) Handschin, E in Dahl, F, 1929:121 ISS
             Syn.: Sminthurus viridis var. nigro-maculata(sic) Buitendijk, AM, 1930:74 ISS
            Syn.: Sminthurus viridis ssp. nigro-maculatus(sic) Stach, 1930 IOS
             Syn.: Sminthurus viridis ssp. nigromaculata(sic) Schubert, 1933 ISS
             Syn.: Sminthurus viridis ssp. nigromaculata(sic) Strebel, 1938 ISS
             Syn.: Sminthurus viridis ssp. nigromaculata(sic) Buitendijk, 1941 ISS
             Syn.: Sminthurus viridis var. nigromaculata(sic) Stach, J, 1956:268 ISS
Sminthurus viridis from Poland
After Stach J, 1956 PlateXXXII Fig.3

             Syn.: Sminthurus viridis ssp. nigromaculata(sic) Stevanovic, 1956 ISS
             Syn.: Sminthurus viridis ssp. nigromaculata(sic) Grinbergs, 1958 ISS
             Syn.: Sminthurus viridis ssp. nigromaculata(sic) Grinbergs, 1958 ISS
             Syn.: Sminthurus viridis ssp. nigromaculata(sic) Grinbergs, 1960 ISS
            Syn.: Sminthurus viridis ssp. nigro-maculatus(sic) Salmon, JT, 1964:605 IOS
Sminthurus nigromaculatus ♀ from Rhodos
Only 1 spot on abd.6
After Ellis WN, 1974 Fig.6a
Sminthurus nigromaculatus from Rhodos
Only 1 spot on abd.6
After Ellis WN, 1974 Fig.6b
Sminthurus nigromaculatus from Holland
2 spots on abd.6
After Ellis WN, 1974 Fig.6c
Sminthurus (Sminthurus) nigromaculatus from the USA
After Snider RJ & Calandrino FJ, 1987 Fig.47
Sminthurus (Sminthurus) nigromaculatus from the USA
After Snider RJ & Calandrino FJ, 1987 Fig.48
Sminthurus nigromaculatus from the USA
After Snider R in Christiansen K & Bellinger P, 1998 Fig.1010.6A
Sminthurus nigromaculatus from the USA
After Snider R in Christiansen K & Bellinger P, 1998 Fig.1010.6B
Sminthurus viridis from Sweden
Ecovariant nigromaculatus
2007.09.03 © Hall, K.
Sminthurus viridis from the UK
Ecovariant nigromaculatus
2007.03.04 © Cornwall, N.J.
Sminthurus viridis from the UK
Ecovariant nigromaculatus
2008.09.11 © Wood, A.
Sminthurus viridis from the UK
Ecovariant nigromaculatus
2008.09.15 © Wood, A.
Sminthurus viridis from Czechia
Ecovariant nigromaculatus
2010.11.06 © Fiala, M.
Sminthurus viridis from Czechia
Ecovariant nigromaculatus
2010.11.12 © Krásenský, P.
Sminthurus viridis from Czechia
Ecovariant nigromaculatus
Grooming
2010.11.12 © Krásenský, P.
Sminthurus viridis from Holland
Ecovariant nigromaculatus
2011.12.02 © van Duinen, J.
Sminthurus viridis from Holland
Ecovariant nigromaculatus
2015.09.23 © van Duinen, J.
Sminthurus viridis from Holland
Ecovariant nigromaculatus
2015.09.23 © van Duinen, J.
Sminthurus viridis from Holland
Ecovariant nigromaculatus
2016.11.05 © van Leeuwen, J.

             Syn.: Sminthurus nigromaculata(sic) Mayvan, MM, Sadeghi-Namaghi, H et Shayanmehr, M, 2021:9 ISS
Sminthurus viridis ecovariant nigromaculatus ♂ from Belgium
Small abdomen with 3 black spots
2021.08.04 © Huskens, M.-L.
Sminthurus viridis ecovariant nigromaculatus ♂ from Belgium
2021.08.04 © Huskens, M.-L.
Sminthurus viridis ecovariant nigromaculatus ♂ from Belgium
Genital papilla
Apical circumvental setae
2021.08.04 © Huskens, M.-L.
Sminthurus viridis ecovariant nigromaculatus ♂ from Belgium
Bothriotricha ABC
In posteriad open obtuse angle
2021.08.04 © Huskens, M.-L.
Sminthurus viridis ecovariant nigromaculatus ♂ from Belgium
2021.08.04 © Huskens, M.-L.


            Syn.: Sminthurus viridis var. trimaculata Reuter, 1891
            Syn.: Sminthurus nigromaculatus var. infuscatus Schött, H, 1893:22
            Syn.: Sminthurus viridis var. dorsovittatus Schött, H, 1893:23
Sminthurus viridis var. dorsovittatus from Sweden
After Schött H, 1893 Taf.I Fig.3

             Syn.: Sminthurus viridis ssp. dorsovittata(sic) Axelson, 1907 ISS
             Syn.: Sminthurus viridis ssp. dorsovittata(sic) Axelson, 1912 ISS
             Syn.: Sminthurus viridis ssp. dorso-vittata(sic) Ionesco, 1915 ISS
             Syn.: Sminthurus viridis ssp. dorsovittata(sic) Ionesco, 1916 ISS
             Syn.: Sminthurus viridis ssp. dorsovittata(sic) Stach, 1920 ISS
             Syn.: Sminthurus viridis ssp. dorsovittata(sic) Stach, 1922 ISS
             Syn.: Sminthurus viridis ssp. dorsovittata(sic) Handschin, 1924 ISS
             Syn.: Sminthurus viridis f. dorsovittata(sic) Handschin, E in Dahl, F, 1929:121 ISS
             Syn.: Sminthurus viridis ssp. dorsovittata(sic) Kseneman, 1937 ISS
             Syn.: Sminthurus (Sminthurus) viridis ssp. dorsovittata(sic) Grinbergs, 1960 ISS
            Syn.: Sminthurus viridis var. speciosus Schött, H, 1893:23
Sminthurus viridis var. speciosus from Sweden
After Schött H, 1893 Taf.I Fig.4

             Syn.: Sminthurus viridis ssp. speciosa(sic) Lie-Pettersen, 1896 ISS
             Syn.: Sminthurus viridis ssp. speciosa(sic) Scherbakow, 1898 ISS
             Syn.: Sminthurus viridis ssp. speciosa(sic) Krausbauer, 1901 ISS
             Syn.: Sminthurus viridis ssp. speciosa(sic) Ågren, 1903 ISS
             Syn.: Sminthurus viridis ssp. speciosa(sic) Axelson, 1905 ISS
             Syn.: Sminthurus viridis ssp. speciosa(sic) Wahlgren, 1906 ISS
             Syn.: Sminthurus viridis ssp. speciosa(sic) Axelson, 1907 ISS
             Syn.: Sminthurus viridis ssp. speciosa(sic) Axelson, 1912 ISS
             Syn.: Sminthurus viridis ssp. speciosa(sic) Schille, 1912 ISS
             Syn.: Sminthurus viridis f. speciosa(sic) Handschin, E in Dahl, F, 1929:121 ISS
             Syn.: Sminthurus viridis ssp. speciosa(sic) Kseneman, 1938 ISS
             Syn.: Sminthurus viridis var. speciosa(sic) Stach, J, 1956:269 ISS
             Syn.: Sminthurus viridis ssp. speciosa(sic) Grinbergs, 1958 ISS
             Syn.: Sminthurus viridis ssp. speciosa(sic) Grinbergs, 1958 ISS
             Syn.: Sminthurus (Sminthurus) viridis ssp. speciosa(sic) Grinbergs, 1960 ISS
             Syn.: Sminthurus viridis var. speciosa(sic) Walters, MC, 1967:18 ISS
            Syn.: Sminthurus viridis ssp. irroratus Reuter, 1900
             Syn.: Sminthurus viridis ssp. irrorata(sic) Axelson, 1907 ISS
             Syn.: Sminthurus viridis ssp. irrorata(sic) Stach, 1922 ISS
             Syn.: Sminthurus viridis ssp. irrorata(sic) Handschin, 1924 ISS
             Syn.: Sminthurus viridis ssp. irrorata(sic) Handschin, 1924 ISS
             Syn.: Sminthurus viridis f. irrorata(sic) Handschin, E in Dahl, F, 1929:121 ISS
             Syn.: Sminthurus viridis ssp. irrorata(sic) Strebel, 1938 ISS
             Syn.: Sminthurus viridis var. irrorata(sic) Stach, J, 1956:269 ISS
             Syn.: Sminthurus (Sminthurus) viridis ssp. irrorata(sic) Grinbergs, 1960 ISS
            Syn.: Sminthurus viridis var. variegatus Latzel, R, 1917:250
            Syn.: Sminthurus viridis var. medicaginis Schött, H, 1917:58
Sminthurus viridis var. medicaginis from Australia
After Schött H, 1917 Taf.4 Fig.11
Sminthurus viridis var. medicaginis from Australia
Abdomen ventral aspect
After Schött H, 1917 Taf.4 Fig.12


            Syn.: Sminthurus viridis var. marmorata(sic) Stach, J, 1924:121 IOS
Sminthurus viridis var. marmorata(sic) from Tunis
After Stach J, 1924 Taf.II Fig.8


             Syn.: Sminthurus viridis var. marmorata(sic) Stach, J, 1956:269 ISS
            Syn.: Arrhopalites adelaidica Womersley, 1933 partim
            Syn.: Sminthurus multidentata(sic) Salmon, JT, 1943:10 IOS
Sminthurus multidentata(sic) from New Zealand
After Salmon J.T., 1943 Pl.1 Fig.1


            Syn.: Sminthurus multipunctatus Bretfeld, G, 2001:130 =?



            Syn.: Pulex viridis plantarum Linnaeus, 1736 i
            Syn.: Podura viridis Hill, 1752 i
Smynthurus(sic) viridis from Ireland
After Templeton, R, 1836 Pl.XII Fig.7
Smynthurus(sic) viridis from Switzerland
After Nicolet H, 1842 Pl.9 Fig.9
Smynthurus(sic) viridis from the UK
After Lubbock J, 1870 Pl.XXI Fig.1
Smynthurus(sic) viridis from the UK
Anterior foot
After Lubbock J, 1870 Pl.XXI Fig.2
Smynthurus(sic) viridis from the UK
Saltatory appendage
After Lubbock J, 1870 Pl.XXI Fig.3
Smynthurus(sic) viridis from the UK
After Lubbock J, 1873 Pl.1


             Syn.: Sminthurus Sviridis(sic) Schött, H, 1893:4 ISS
             Syn.: Sminthurus (Setosi) virides(sic) Schött, H, 1891:9 ISS
Sminthurus viridis f. principalis from Sweden
After Schött H, 1893 Taf.I Fig.1
Sminthurus viridus(sic) from Germany
Capitulum of maxilla
After Anonymous Fig.A
Sminthurus viridis from Tasmania
After Womersley, 1931 Pl.I Fig.7
Sminthurus viridis from Australia
After Womersley H, 1939 Pl.1 Fig.3.
Sminthurus viridis from Malta
Colour variations
After Stach J, 1967 Pl.XXXIV


             Syn.: Sminthuro(sic) viridi(sic) Keller, S et Steenberg, T, 1997:22 ISS
Neozygites sminthuri from Denmark. Bar=10µm
Conidia infecting Sminthurus viridis
After Dromph & al, 2001 Fig.1
Sminthurus viridis (juv.) from the UK
2007 © Cornwall, N.J.
Sminthurus viridis (juv.) from Spain
2007.03.09 © Portela, J.
Sminthurus viridis (juv.) from Belgium
2007.04.25 © Lindsley, J.
Sminthurus viridis (juv.) from Belgium
2007.07.13 © Haentjes, R.
Sminthurus viridis (juv.) from Belgium
2007.07.25 © Vuijlsteke, M.
Sminthurus viridis (juv.) from Slovakia
2007.08.27 © Stefanovie, P.
Sminthurus viridis (juv.) from the UK
2007.09.08 © Botting, J.
Sminthurus viridis (juv.) from France
2008.04.14 © Liger, D.
Sminthurus viridis (juv.) from the UK
2008.04.25 © Valentine., B
Sminthurus viridis from Sweden
2006 © Hall, K.
Sminthurus viridis from the UK
2006 © Cornwall, N.J.
Sminthurus viridis from France
2006 © Lavoué., B
Sminthurus viridis from France
2006 © Silversword31.
Sminthurus viridis from France
2006 © Silversword31.
Sminthurus viridis from France
2006 © Silversword31.
Sminthurus viridis from the UK
2006 © Cornwall, N.J.
Sminthurus viridis from the UK
2007.03.18 © Cornwall, N.J.
Sminthurus viridis from Sweden
2007.07.25 © Hall, K.
Sminthurus viridis from the UK
2007.10.16 © Campbell, A.
Sminthurus viridis from the UK
2007.10.29 © Campbell, A.
Sminthurus viridis from the UK
2007.11.03 © Campbell, A.
Sminthurus viridis from the UK
2008.03.30 © Botting, J.
Sminthurus viridis from Belgium
2008.04.17 © San Martin, G.
Sminthurus viridis from Belgium
2008.07.23 © Haentjens, R.
Sminthurus viridis from France
2008.19.12 © Lebeaux, P.
Sminthurus viridis from the UK
2008.11.12 © Thompson, S.
Sminthurus viridis from the UK
2008.11.12 © Thompson, S.
Sminthurus viridis from Australia
2009.09.07 © Hort, J.
Sminthurus viridis from Australia
2009.09.07 © Hort, J.
Sminthurus viridis from Ibiza
2011.11.03 © Beer, R.
Sminthurus viridis from the UK
Pale colour form
2012.03.29 © Murray, A.
Sminthurus viridis from Italy
2012.05.09 © Fardels, J.
Sminthurus viridis from the UK
Annulated 4th antennal segment
2012.06.01 © Robertson, A.
Sminthurus viridis from Greece
2012.12.29 © Markopoulos, S.
Sminthurus viridis from the UK
Eyepatch
2015.09.20 © Murray, A.
Sminthurus viridis from France
2015.12.18 © Gros, P.
Sminthurus viridis from France
2015.12.18 © Gros, P.
Sminthurus viridis from France
2015.12.18 © Gros, P.
Sminthurus viridis ♂ from Belgium
Depositing spermatophore
2017.08.23 © Huskens, M.-L.
Sminthurus viridis from France
2019.12.08 © Tcheng, E.
Sminthurus viridis from France
Eye with 8 ocelli A-H
2020.03.26 © Garcelon, P.
Sminthurus viridis juv. from Spain
2022.01.16 © Zeeders, M.

  Subspecies annulatus Folsom, JW, 1899:270

Sminthurus viridis var. annulatus from Japan
After Folsom JW, 1899 Pl.3 Fig.41

             Syn.: Smynthurus(sic) viridis f. cinereo-viridis(sic) Folsom, JW, 1899:272 ISS



  Subspecies caucasicus Karsch, 1893

  Subspecies cinereoviridis Tullberg, 1871


            Syn.: Smynthurus(sic) viridis f. cinereo-viridis(sic) Tullberg, 1871 IOS
            Syn.: Smynthurus(sic) viridis Nicolet, H, 1842:82 partim


             Syn.: Sminthurus viridis var. cinereo-viridis(sic) Tullberg, T, 1872:30 ISS
             Syn.: Sminthurus viridis var. cinereo-viridis(sic) Reuter, OM, 1876:79 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Parona, 1884 ISS
             Syn.: Smynthurus(sic) fuscus var. cinereo-viridis(sic) von Dalla Torre, 1888 ISS
             Syn.: Smynthurus(sic) viridis var. cinereo-viridis(sic) Parona, 1888 ISS
             Syn.: Sminthurus viridis var. cinereo-viridis(sic) Reuter, 1891 ISS
             Syn.: Sminthurus viridis var. cinereo-viridis(sic) Schött, H, 1893:22 ISS
Sminthurus viridis var. cinereo-viridis(sic) from Sweden
After Schött H, 1893 Taf.I Fig.2

             Syn.: Sminthurus viridis var. cinereo-viridis(sic) Reuter, OM, 1895:9 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Schäffer, 1897 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Börner, 1900 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Schäffer, 1900 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Schäffer, 1900 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Carl, 1901 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Krausbauer, 1901 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Ågren, 1903 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Axelson, 1905 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Axelson, 1906 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Axelson, 1907 ISS
             Syn.: Sminthurus viridis var. cinereo-viridis(sic) Linnaniemi, WM, 1912:313 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Schille, 1912 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Ionesco, 1915 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Ionesco, 1916 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Ionesco, 1920 ISS
             Syn.: Sminthurus viridis f. cinereo-viridis(sic) Handschin, E in Dahl, F, 1929:121 ISS
             Syn.: Sminthurus viridis var. cinereo-viridis(sic) Buitendijk, AM, 1930:73 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Stach, 1930 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Schubert, 1933 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Durkop, 1934 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Buitendijk, 1941 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Strenzke, 1949 ISS
             Syn.: Sminthurus viridis var. cinereo-viridis(sic) Stach, J, 1956:269 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Grinbergs, 1958 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Grinbergs, 1958 ISS
             Syn.: Sminthurus viridis ssp. cinereo-viridis(sic) Salmon, JT, 1964:603 ISS
Sminthurus ssp. cinereoviridis from France
2006 © Silversword31.
Sminthurus ssp. cinereoviridis from France
2006 © Silversword31.
Sminthurus ssp. cinereoviridis from France
2008.02.14 © Lavoué, Br.
Sminthurus viridis ssp. cinereoviridis from the UK
2008.11.26 © Thompson, S.
Sminthurus ssp. cinereoviridis from Portugal
2011.02.25 © Almeida, J.
Sminthurus ssp. cinereoviridis from Portugal
2011.02.25 © Almeida, J.
Sminthurus ssp. cinereoviridis from Portugal
Annulated 4th antennal segment
2011.02.25 © Almeida, J.
Sminthurus ssp. cinereoviridis from Portugal
Small abdomen
2011.02.25 © Almeida, J.
Sminthurus ssp. cinereoviridis from Portugal
2011.02.25 © Almeida, J.
Sminthurus ssp. cinereoviridis juv. from Portugal
2011.02.25 © Almeida, J.
Sminthurus ssp. cinereoviridis from Holland
Drinking from dew drops on body using long eversed vesicle of collophore
2011.11.20 © van Duinen, J.
Sminthurus ssp. cinereoviridis from France
2013.02.02 © Sandra.
Sminthurus ssp. cinereoviridis from Italy
2013.02.19 © Consolo, M.
Sminthurus viridis ssp. cinereoviridis from Germany
2014.03.12 © Haselböck,A.
Sminthurus viridis ssp. cinereoviridis from Germany
2014.03.12 © Haselböck,A.
Sminthurus viridis ssp. cinereoviridis from Holland
With parasitic larva of Acari:Erythraeoidea
2015.07.26 © van Duinen, J.
Sminthurus viridis ssp. cinereoviridis from Holland
With 2 black spots on small abdomen
2015.07.26 © van Duinen, J.
Sminthurus viridis ssp. cinereoviridis from Holland
Small abdomen with 2 black dots (*)
2015.09.11 © van Duinen, J.
Sminthurus viridis ssp. cinereoviridis from Holland
Ready to moult
2015.09.16 © van Duinen, J.
Sminthurus viridis ssp. cinereoviridis from Holland
2015.09.16 © van Duinen, J.
Sminthurus viridis ssp. cinereoviridis from Holland
Eyepatch with large ocelli A & B (*)
2015.09.18 © van Duinen, J.
Sminthurus viridis ssp. cinereoviridis from Sweden
2015.09.20 © Neergaard, R.
Sminthurus ssp. cinereoviridis from Slovakia
2015.11.02 © Sersen, J.
Sminthurus ssp. cinereoviridis from Slovakia
2015.11.05 © Sersen, J.
Sminthurus ssp. cinereoviridis from Slovakia
2015.11.05 © Sersen, J.
Sminthurus viridis ssp. cinereoviridis from Holland
2016.09.03 © van Duinen, J.
Sminthurus viridis ssp. cinereoviridis from Belgium
2016.12.13 © Huskens, M.-L.
Sminthurus viridis ssp. cinereoviridis from Belgium
2016.12.13 © Huskens, M.-L.
Sminthurus viridis ssp. cinereoviridis from Belgium
Two dots on abd.6
2016.12.13 © Huskens, M.-L.
Sminthurus viridis ssp. cinereoviridis from Italy
2017.10.04 © Regina, E.
Sminthurus viridis ssp. cinereoviridis juv. from France
2017.10.09 © Picard, J.
Sminthurus viridis ssp. cinereoviridis from France
Caudal aspect
2017.10.09 © Picard, J.
Sminthurus viridis ssp. cinereoviridis from France
2018.01.10 © Garcelon, P.
Sminthurus viridis ssp. cinereoviridis from France
2019.01.26 © Joachim, J.
Sminthurus viridis ssp. cinereoviridis from Belgium
Distinctly larger ocelli A & B
2019.06.18 © Huskens, M.-L.
Sminthurus viridis ssp. cinereoviridis from Belgium
Two dots on abd.6
2019.06.18 © Huskens, M.-L.
Sminthurus viridis ssp. cinereoviridis from Germany
2020.02.14 © Gutekunst, V.
Sminthurus viridis ssp. cinereoviridis from Germany
Dorsal anal valve with 2 black spots
2020.02.14 © Gutekunst, V.
Sminthurus viridis ssp. cinereoviridis from Holland
Small abdomen
2020.08.29 © Kamsteeg, G.
Sminthurus viridis ssp. cinereoviridis from Belgium
Grooming
With extended collophore vesicle
2020.10.13 © Huskens, M.-L.
Sminthurus viridis ssp. cinereoviridis from Belgium
With extended collophore vesicle
2020.10.13 © Huskens, M.-L.
Sminthurus viridis ssp. cinereoviridis from Belgium
With extended collophore vesicles
2020.10.13 © Huskens, M.-L.
Sminthurus viridis ssp. cinereoviridis juv. from Belgium
2020.12.16 © Huskens, M.-L.
Sminthurus viridis ssp. cinereoviridis ♂ from Belgium
With parasitic Acari protonymph
2021.06.17 © Huskens, M.-L.
Sminthurus viridis ssp. cinereoviridis from Belgium
2022.01.18 © Huskens, M.-L.
Sminthurus viridis ssp. cinereoviridis from Belgium
Abd.5 without spot
Abd.6 with 2 spots
2022.01.18 © Huskens, M.-L.

  Subspecies elegans Ionesco, 1915

  Subspecies infuscatus Reuter, OM, 1895:10

            Syn.: Sminthurus nigromaculatus var. infuscatus Reuter, OM, 1892:232 nn
             Syn.: Sminthurus viridis var. infuscata(sic) Stach, J, 1956:269 ISS
Sminthurus viridis ssp. infuscatus from the UK
2012.05.11 © Metal, P.

  Subspecies lineatus Scherbakov, AM, 1898:22

Sminthurus viridis var. lineata from Russia
After Scherbakov AM, 1898 Table III Fig.29

             Syn.: Sminthurus viridis ssp. lineata(sic) Ionesco, 1916 ISS
             Syn.: Sminthurus viridis var. lineata(sic) Stach, J, 1956:269 ISS
  Subspecies pictus Skorikov, 1899


             Syn.: Sminthurus viridis ssp. picta(sic) Axelson, 1919 ISS
             Syn.: Sminthurus viridis var. picta(sic) Stach, J, 1956:269 ISS
             Syn.: Sminthurus (Sminthurus) viridis ssp. picta(sic) Grinbergs, 1960 ISS
  Subspecies sabulosus Skorikow, 1899


            Syn.: Sminthurus viridis ssp. sabulosa(sic) Skorikow, 1899 IOS
             Syn.: Sminthurus viridis ssp. sabulosa(sic) Grinbergs, 1960 ISS
  Subspecies stscherbakowi Skorikow, 1899

  Subspecies tripunctatus Reuter, 1891

Sminthurus viridis var. tripunctatus from Sweden
After Schött H, 1893 Taf.I Fig.5

             Syn.: Sminthurus viridis ssp. tripunctata(sic) Schäffer, 1900 ISS
             Syn.: Sminthurus viridis var. tripunctata(sic) Stach, J, 1956:269 ISS
             Syn.: Sminthurus viridis ssp. tripunctata(sic) Grinbergs, 1960 ISS
  Subspecies viridiflavus Parona, 1884


             Syn.: Sminthurus viridi-flavus(sic) Stach, J, 1924:122 ISS
  Subspecies xerophilus Skorikov, 1899


            Syn.: Sminthurus viridis ssp. xerophila(sic) Skorikov, 1899 IOS
             Syn.: Sminthurus viridis var. xerophila(sic) Stach, J, 1956:269 ISS



Species wahlgreni Stach, 1919

             Syn.: Sminthurus Wahlgreni(sic) Stach, J, 1930:310 ISS
             Syn.: Sminthurus Wahlgreni(sic) Denis, J-R, 1933:305 ISS
             Syn.: Sminthurus Wahlgreni(sic) Denis, J-R, 1933:309 ISS
Sminthurus wahlgreni from Poland
After Stach J, 1956 PlateXXVIII Fig.3


             Syn.: Sminthurus walgrenii(sic) Gruia, 1966 ISS
             Syn.: Sminthurus walgreni(sic) Fiera, C, 2007:32 ISS
Sminthurus wahlgreni? from France
2018.05.22 © Picard, J.
Sminthurus wahlgreni? from France
2018.05.22 © Picard, J.
Sminthurus wahlgreni? from France
Ant.4 with twenty annulations
2018.05.22 © Picard, J.



Species wutaii Uchida, 1948

Species yemenensis Bretfeld, 2000

Species yonahlosee Wray, 1948

             Syn.: Sminthurus yonahlossee(sic) Poole, RW in Poole, RW et Gentili, P, 1997:56 ISS



Genus Austrosminthurus 0082:274

Species mirabilis Delamare Deboutteville, C et Massoud, Z, 1963:274, t.t.


Genus Novokatianna Salmon, JT, 1944:173

  Syn.: Katiannini Novakatianna(sic) Richards, WR, 1968:22 ISS
  Syn.: Sminthuridae Novakatianna(sic) Zeppelini Filho, D, 1996:17 ISS
  Syn.: Sminthuridae Novakatianna(sic) Greenslade, P, 2015:3 ISS
  Syn.: Sminthurinae Novakatianna(sic) Zeppelini, D, Brito, RA, Zampaulo, R et de Lima, ECA, 2020:26 ISS
  Syn.: Sminthurinae Novakatianna(sic) Zeppelini, D, Brito, RA, Zampaulo, R et de Lima, ECA, 2020:27 ISS
  Syn.: Sminthurinae Novakatianna(sic) Zeppelini, D, Brito, RA, Zampaulo, R et de Lima, ECA, 2020:44 ISS

Species sp.1 Janssens F & Murray A, 2014

Novokatianna sp. from New Zealand
2014.02.24 © Murray, A.
Novokatianna sp. from New Zealand
2014.02.24 © Murray, A.
Novokatianna sp. from New Zealand
2016.02.11 © Murray, A.
Novokatianna sp. from New Zealand
2016.02.20 © Murray, A.

Species cummyxa Salmon, JT, 1944:174, t.t.

Novokatianna cummyxa from New Zealand
2014 © TePapa Museum
Novokatianna cummyxa from New Zealand
2014.08.06 © Murray, A.

             Syn.: Novakatianna(sic) cunnaxy(sic) Greenslade, P, 2015:3 ISS
Novakatianna(sic) cunnaxy(sic) from New Zealand
2015 © Murray A in Greenslade P, 2015 Fig.1



Species radiata Salmon, JT, 1946:36

Novokatianna radiata from New Zealand
2014 © TePapa Museum

Species venusta (Salmon, JT, 1943:4) Salmon, JT, 1944:175

Katianna venusta from New Zealand
After Salmon JT, 1943 Pl.1 Fig.2
Katianna venusta from New Zealand
After Salmon JT, 1943 Pl.1 Fig.3
Novokatianna venusta from New Zealand
2014 © TePapa Museum
Novokatianna venusta from New Zealand
2016.02.08 © Murray, A.


Genus Richardsitas Betsch, J-M, 1975:480 key

  Syn.: Collembola Richarsitas(sic) Thibaud, J-M, 2013:241 ISS

Species griveaudi Betsch, 1977

Species najtae Betsch, J-M, 1975:482, t.t.

Species subferoleum Medeiros, GDS, Greenslade, P et Bellini, BC, 2020:3

Richardsitas subferoleum ♂ from Australia
With short candle-shaped dorsal setae
After Medeiros GDS & al, 2020 Fig.5E
Richardsitas subferoleum ♀ from Australia
Large abdomen
After Medeiros GDS & al, 2020 Fig.5F


Genus Spatulosminthurus Betsch & Betsch-Pinot, 1983

  Syn.: Collembola Smithurus(sic) Stach, J, 1967:394 ISS
  Syn.: ... Sminthurus Betsch & Betsch-Pinot, 1984 partim


  Syn.: Collembola Spatulosminthurus Betsch & Betsch-Pinot, 1982 u
Spatulosminthurus betschi ♂ from France
Unguis with tunica (*)
Footcomplex with knobbed tenent macrosetae (**)
2020.01.01 © Tiky, S.



Species betschi Nayrolles, P, 1990:65

Spatulosminthurus betschi ♂ from Spain
After Nayrolles P, 1990 Fig.1
Spatulosminthurus betschi ♀ juv. from Spain
After Nayrolles P, 1990 Fig.2
Spatulosminthurus betschi ♀ from Spain
After Nayrolles P, 1990 Fig.3
Spatulosminthurus ?betschi? from Spain
2014.11.08 © Martinez, I.
Spatulosminthurus betschi from France
2019.12.25 © Tiky, S.
Spatulosminthurus betschi from France
Caudal aspect
2019.12.25 © Tiky, S.
Spatulosminthurus betschi juv. from France
2019.12.25 © Tiky, S.
Spatulosminthurus betschi from France
Caudal aspect
2019.12.25 © Tiky, S.
Spatulosminthurus betschi from France
Middorsal pattern
Caudal aspect
2019.12.25 © Tiky, S.
Spatulosminthurus betschi from France
2019.12.25 © Tiky, S.
Spatulosminthurus betschi from France
2019.12.25 © Tiky, S.
Spatulosminthurus betschi from France
2019.12.25 © Tiky, S.
Spatulosminthurus betschi from France
2019.12.25 © Tiky, S.
Spatulosminthurus betschi from France
2019.12.28 © Tiky, S.
Spatulosminthurus betschi from France
Ant.4 with 21 subdivisions
2019.12.28 © Tiky, S.
Spatulosminthurus betschi ♂ from France
2020.01.01 © Tiky, S.
Spatulosminthurus betschi ♂ from France
Caudal aspect
2020.01.01 © Tiky, S.
Spatulosminthurus betschi from France
Face
2020.01.01 © Tiky, S.
Spatulosminthurus betschi from France
2020.11.14 © Tcheng, E.
Spatulosminthurus betschi from France
Median dorsal anterior pattern
2020.11.14 © Tcheng, E.
Spatulosminthurus betschi from France
Median dorsal posterior pattern
2020.11.14 © Tcheng, E.

Species flaviceps (Tullberg, 1871) Bretfeld, G, 1999:206

            Syn.: Sminthurus flaviceps var. fennicus Reuter, OM, 1876:83
             Syn.: Sminthurus flaviceps f. fennica(sic) Handschin, E in Dahl, F, 1929:118 ISS
             Syn.: Spatulosminthurus flaviceps var. fennica(sic) Bretfeld, G, 1999:206 ISS
            Syn.: Sminthurus fuscus var. ornatus Uzel, 1891
Sminthurus flaviceps var. ornata from Poland
After Stach J, 1956 PlateXXX Fig.2
Spatulosminthurus ?flaviceps var. ornata? from France
2019.06.12 © Picard, J.

            Syn.: Sminthurus flaviceps var. media Stach, 1922
            Syn.: Sminthurus flaviceps f. principalis Stach, J, 1956:258
            Syn.: Sminthurus flaviceps var. bifasciata(sic) Stach, J, 1956:260
Sminthurus flaviceps var. bifasciata(sic) from Poland
After Stach J, 1956 PlateXXXII Fig.2

            Syn.: Sminthurus flaviceps var. discolor Stach, J, 1956:260
Sminthurus flaviceps var. discolor from Poland
After Stach J, 1956 PlateXXX Fig.3

            Syn.: Sminthurus bulgaricus Rusek, 1965
             Syn.: Spatulosminthurus bulgarius(sic) Nayrolles, P, 1990:65 ISS



Sminthurus flaviceps from Sweden
After Schött H, 1893 Taf.I Fig.7
             Syn.: Sminthurus flavicipite(sic) Reuter, OM, 1895:9 ISS
Sminthurus flaviceps from Poland
After Stach J, 1956 PlateXXX Fig.1



Species flaviceps2 (Buitendijk, 1941) Ellis, 1976

Species gattoi (Stach, J, 1967:413) Betsch & Betsch-Pinot, 1983

Sminthurus gattoi from Malta
After Stach J, 1967 Pl.XXXV Fig.1
Spatulosminthurus gattoi from Italy
2017.01.28 © Rinaldo, R.
Spatulosminthurus gattoi from Italy
2017.01.28 © Rinaldo, R.
Spatulosminthurus gattoi from Italy
2017.01.28 © Rinaldo, R.
Spatulosminthurus gattoi from Italy
Ant.4 with about, 20 subdivisions
2021.02.24 © Colombo, M.
Spatulosminthurus gattoi from Italy
2021.02.24 © Colombo, M.

Species guthriei (Stach, 1919) Betsch, J-M & Betsch-Pinot, M-C, 1984:71

            Syn.: Sminthurus guthriei var. aurantiaca Stach, 1921


             Syn.: Sminthurus Guthriei(sic) Denis, J-R, 1933:305 ISS
             Syn.: Sminthurus Guthriei(sic) Denis, J-R, 1933:308 ISS
  Subspecies dimorphus Stach, J, 1956:251

Sminthurus guthriei var. dimorphus from Poland
After Stach J, 1956 PlateXXVIII Fig.1
Sminthurus guthriei var. dimorphus from Poland
After Stach J, 1956 PlateXXVIII Fig.2
Spatulosminthurus guthriei ssp. dimorphus from Hungary
2018.05.06 © Ujvári, Z.
Spatulosminthurus guthriei ssp. dimorphus from Hungary
2018.05.06 © Ujvári, Z.
Spatulosminthurus guthriei ssp. dimorphus from Hungary
2018.05.06 © Ujvári, Z.

  Subspecies sibiricus Bretfeld, 1996



Species lesnei (Carl in Denis, 1925) Betsch & Betsch-Pinot, 1983, t.t.

            Syn.: Sminthurus lesnei f. obscura(sic) Carl in Denis, 1925 IOS
            Syn.: Spatulosminthurus lesnei ssp. obscura(sic) Betsch, J-M et Betsch-Pinot, M-C, 1984:75 IOS
            Syn.: Spatulosminthurus lesnei f. obscura(sic) Bretfeld, G, 1999:210 IOS
            Syn.: Sminthurus lesnei f. obscura(sic) Thibaud, J-M, 2013:234 IOS
            Syn.: Sminthurus lesnei f. maculata(sic) Denis, 1925 IOS
            Syn.: Sminthurus lesnei f. maculata(sic) Dallai, 1967 IOS
            Syn.: Spatulosminthurus lesnei f. maculata(sic) Bretfeld, G, 1999:210 IOS
            Syn.: Spatulosminthurus betschi f. maculata(sic) Bretfeld, G, 1999:210 =? IOS
            Syn.: Sminthurus lesnei f. maculata(sic) Thibaud, J-M, 2013:234 IOS
Spatulosminthurus lesnei f. maculatus from Ibiza
2009.01.25 © Beer, R.
Spatulosminthurus lesnei f. maculatus from Ibiza
2009.12.02 © Beer, R.
Spatulosminthurus lesnei f. maculatus from Ibiza
2009.12.02 © Beer, R.
Spatulosminthurus lesnei f. maculatus from Ibiza
2009.12.03 © Beer, R.
Spatulosminthurus lesnei f. maculatus from Ibiza
2009.12.05 © Beer, R.
Spatulosminthurus lesnei f. maculatus from Ibiza
2009.12.05 © Beer, R.
Spatulosminthurus lesnei f. maculatus from Ibiza
Spatulate tenent hair
2009.12.06 © Beer, R.
Spatulosminthurus lesnei f. maculatus from Ibiza
Antenna
2009.12.06 © Beer, R.
Spatulosminthurus lesnei f. maculatus from Ibiza
2009.12.06 © Beer, R.
 
Spatulosminthurus lesnei f. maculatus from Ibiza
2011.12.04 © Beer, R.
Spatulosminthurus lesnei f. maculatus from Portugal
2012.10.16 © Murray, A.
Spatulosminthurus lesnei f. maculatus from Ibiza
2014.12.04 © Beer, R.
Spatulosminthurus lesnei f. maculatus from Ibiza
2015.11.28 © Beer, R.
Spatulosminthurus lesnei f. maculatus from Portugal
2020.11.21 © Almeida, J.

            Syn.: Sminthurus lesnei var. remaudierei Delamare Deboutteville, 1945
Spatulosminthurus lesnei f. remaudierei from France
2019.12.27 © Tiky, S.
Spatulosminthurus lesnei f. remaudierei ♂ from France
2020.01.01 © Tiky, S.
Spatulosminthurus lesnei f. remaudierei ♂ from France
2020.01.01 © Tiky, S.




             Syn.: Sminthurus Lesnei(sic) Denis, J-R, 1933:305 ISS
             Syn.: Sminthurus Lesnei(sic) Denis, J-R, 1933:308 ISS
            Syn.: Spatulosminthurus lesnei Betsch & Betsch-Pinot, 1982 u
Spatulosminthurus lesnei from France
After Denis, 1925 in Bretfeld G, 1999 Fig.453
Spatulosminthurus lesnei from Spain
2013.02.10 © Frias, J.R.
Spatulosminthurus lesnei juv. from Ibiza
2014.12.04 © Beer, R.
Spatulosminthurus lesnei from France
With black head and broad black dorsal longitudinal stripe
2017.10.18 © Tiky, S.
Spatulosminthurus lesnei from France
With pale narrow thoracic stripe
2017.10.18 © Tiky, S.
Spatulosminthurus lesnei from France
Caudal aspect
2017.10.18 © Tiky, S.
Spatulosminthurus lesnei from France
Ant.4 with more than twenty annulations
2017.10.18 © Tiky, S.
Spatulosminthurus lesnei ♂ from France
Raised genital papilla (*)
2017.11.01 © Tiky, S.
Spatulosminthurus lesnei ♂ from France
Caudal aspect
2017.11.01 © Tiky, S.
Spatulosminthurus lesnei ♂ from France
2017.11.01 © Tiky, S.
Spatulosminthurus lesnei ♂ from France
Small abdomen
2017.11.01 © Tiky, S.
Spatulosminthurus lesnei from France
Facial chaetotaxy
2017.11.01 © Tiky, S.
Spatulosminthurus lesnei ♀ from France
Genital gonopore (*)
2017.11.01 © Tiky, S.
Spatulosminthurus lesnei ♂ from France
Long narrow mucro
2017.11.01 © Tiky, S.
Spatulosminthurus lesnei from France
The movie
2017.11.04 © Tiky, S.
Spatulosminthurus lesnei from France
2019.06.12 © Picard, J.
Spatulosminthurus lesnei ♂ from France
2019.06.12 © Picard, J.
Spatulosminthurus lesnei from France
2019.06.12 © Picard, J.
Spatulosminthurus lesnei from Spain
Caudal aspect
2019.12.16 © Zeeders, M.






Genus Janusius Bretfeld, G, 2010:313

  Syn.: Symphypleona Sminthuris(sic) Murray, T, 2012:11 ISS


Species annulatus (Kang J-S & Lee B-H in Kang, J-S, Lee, B-H & Park, K-H, 2005:238) Dányi & Park, 2016

Sminthurus annulatus from Korea
After Kang J-S, Lee B-H & Park K-H, 2005 Fig.3A

Species daisetsuzanus (Uchida, H, 1957:25) Bretfeld, G, 2010:313

Sminthurus daisetsuzanus from Japan
After Uchida H, 1957 Fig.2A

Species geumgangensis Dányi & Park, 2016

Species sensibilis (Börner, C, 1909:126) Bretfeld, G, 2010:313

            Syn.: Sminthurus arborealis Itoh, 1985


Spatulosminthurus sensibilis from Japan
After Bretfeld G, 1999 Fig.455
Janusius sensibilis from Japan
2010.02.20 © Tsukiji, T.
Janusius sensibilis from Japan
2014.01.05 © Hara, Y.
Janusius sensibilis from Japan
2014.01.05 © Hara, Y.
Janusius sensibilis from Japan
2014.01.05 © Hara, Y.

Species sylvestris (Banks, 1899 tt) Bretfeld, G, 2010:313

            Syn.: Sminthurus facialis Banks, 1903
Sminthurus facialis from the USA
After Maynard, E.A., 1951
Painting by Carolyn Fallon.

            Syn.: Sminthurus variegatus Axelson, 1903, nec Tullberg



Sminthurus sylvestris from the USA
2006 © Murray, T.
Sminthurus sylvestris from the USA
2007.05.30 © Murray, T.
Sminthurus sylvestris from the USA
2009.05.16 © Murray, T.
Sminthurus sylvestris from the USA
2009.06.07 © Murray, T.
Sminthurus sylvestris from the USA
2010.05.04 © Eiseman, C.
Sminthurus sylvestris from the USA
2010.06.07 © Murray, T.
Janusius sylvestris from the USA
2011.01.20 © Montgomery, G.
Janusius sylvestris from the USA
2011.06.04 © Murray, T.
Janusius sylvestris from the USA
2012.01.27 © Lapp, J.
Janusius sylvestris from the USA
2012.01.27 © Lapp, J.
Janusius sylvestris from the USA
2013.04.27 © Bradford, A.
Janusius sylvestris from the USA
2014.05.04 © B., L.
Janusius sylvestris from the USA
2014.05.04 © B., L.
Janusius sylvestris from the USA
White eyepatch with black ocelli
2015.05.28 © Eiseman, C.


Genus Caprainea Dallai, 1970

  Syn.: Symphypléones(sic) Sminthurus Delamare Deboutteville, C et Bassot, J-M, 1957:82
  Syn.: ... Sphyrotheca(sic) Thibaud & Massoud, 1980 lapsus
  Syn.: Sminthuridae Cordobaia Simón, JC, Bach, C et Gaju, M, 1987:303
  Syn.: Sminthurinae Cordobaia Bretfeld, G, 1999:178



Caprainea marginata f. cinerea from Bulgaria
Macrosetae stout equally thick apically suddenly pointed
2016.06.25 © van Duinen, J.
Caprainea marginata f. cinerea from Bulgaria
Ratio ant.3/2=1.6; ant.4/3=2.8
2016.06.25 © van Duinen, J.
Caprainea marginata f. cinerea from Bulgaria
Distinctly smaller ocelli C+D
2016.06.25 © van Duinen, J.

Species bremondi (Delamare Deboutteville, C & Bassot, J-M, 1957:80) Dallai, 1972, t.t.

Sminthurus Bremondi(sic) from Madeira
After Delamare Deboutteville C & Bassot J-M, 1957 Fig.3A
Sminthurus Bremondi(sic) from Madeira
After Delamare Deboutteville C & Bassot J-M, 1957 Fig.3B

            Syn.: Sminthurus sp. da Gama, 1959
            Syn.: Sminthurus sp. da Gama, 1961
            Syn.: Cordobaia bembezarensis Simón, JC, Bach, C et Gaju, M, 1987:305
Cordobaia bembezarensis from Spain
Abdominal chaetotaxy
After Simón JC & al, 1987 Fig.13
Cordobaia bembezarensis from Spain
Dorsal macrosetae baseball-bat shaped
After Simón JC & al, 1987 Fig.13
Cordobaia bembezarensis from Spain
Cephalic chaetotaxy
After Simón JC & al, 1987 Fig.14

            Syn.: Cordobaia bembezarensis Bretfeld, G, 1999:179



             Syn.: Sminthurus Bremondi(sic) Delamare Deboutteville, C et Bassot, J-M, 1957:81 ISS
             Syn.: Sminthurus Bremondi(sic) Delamare Deboutteville, C et Bassot, J-M, 1957:83 ISS
             Syn.: Sminthurus Bremondi(sic) Delamare Deboutteville, C et Bassot, J-M, 1957:85 ISS
             Syn.: Sminthurus Bremondi(sic) Delamare Deboutteville, C et Bassot, J-M, 1957:86 ISS
Caprainea bremondi from Madeira
After Delamare & Bassot in Gisin H, 1960 Abb.546

            Syn.: Sphyrotheca(lapsus) bremondi Thibaud & Massoud, 1980
Caprainea bremondi from Spain
After Nayrolles P, 1993 Fig.18
Caprainea bremondi from Romania
After Nitzu E in Popa I, 2012 Fig.1A



Species marginata (Schött, H, 1893:25) Dallai, 1972, B

            Syn.: Sminthurus erinaceus Börner, 1903
            Syn.: Sminthurus marginatus Denis, 1924, nec Schött, 1893
            Syn.: Sminthurus echinatus Stach, J, 1930:72,tt
Sminthurus echinatus from Spain
After Stach J, 1930 Abb.XV.8
Sminthurus echinatus from Spain
After Stach J, 1930 Abb.XV.9

Sminthurus echinatus from Poland
After Stach J, 1956 PlateXXXI Fig.5
Sminthurus echinatus from Poland
After Stach J, 1956 PlateXXXI Fig.6
Caprainea marginata from Italy
After Dallai R & al, 2000 Fig.7
Caprainea ?marginata f. echinata?
2006 © Boldsystems.org
Caprainea marginata f. echinata from Portugal
2012.10.16 © Murray, A.
Caprainea marginata f. echinata from Spain
2012.11.29 © Campo., B
Caprainea marginata f. echinata from France
Dark colour form
2016.12.31 © Garcelon, P.
Caprainea marginata f. echinata from France
Pale colour form
2018.11.11 © Garcelon, P.
Caprainea marginata f. echinata from France
Dark specimen
2019.01.26 © Joachim, J.
Caprainea marginata f. echinata from France
Dark specimen
2019.11.09 © Joachim, J.

            Syn.: Sminthurus marginatus var. cinereus Stach, J, 1956:235
             Syn.: Sminthurus marginatus var. cineraceus(sic) Stach, J, 1956:254 ISS
Sminthurus marginatus var. cineraceus(sic) from Poland
After Stach J, 1956 PlateXXXI Fig.9

Caprainea marginata f. cinerea from Bulgaria
2016.06.25 © van Duinen, J.
Caprainea marginata f. cinerea from Bulgaria
2016.06.25 © van Duinen, J.
Caprainea marginata f. cinerea from Holland
2016.10.26 © van Duinen, J.
Caprainea marginata f. cinerea from Holland
Dorsal macrosetae distinctly straight and apically blunt
2016.10.26 © van Duinen, J.
Caprainea marginata f. cinerea from Holland
Distinctly smaller ocelli C+D
2016.10.26 © van Duinen, J.
Caprainea marginata f. cinerea from Holland
Pale face
2016.10.26 © van Duinen, J.
Caprainea marginata f. cinerea from France
2016.12.11 © Garcelon, P.
Caprainea marginata f. cinerea from Belgium
2017.08.31 © Huskens, M.-L.
Caprainea marginata f. cinerea from Belgium
2017.08.31 © Huskens, M.-L.
Caprainea marginata f. cinerea from Holland
2017.09.21 © Wieringa, P.
Caprainea marginata f. cinerea ♂ from France
Mushroom shaped male genital papilla
2017.10.08 © Picard, J.
Caprainea marginata f. cinerea from France
2017.10.09 © Picard, J.
Caprainea marginata f. cinerea from France
2018.10.17 © Picard, J.
Caprainea marginata f. cinerea ♂ from France
2018.10.28 © Tiky, S.
Caprainea marginata f. cinerea from France
2019.11.09 © Joachim, J.
Caprainea marginata f. cinerea from France
2019.12.26 © Reinach, Br.
Caprainea marginata f. cinerea from France
2019.12.26 © Reinach, Br.

            Syn.: Caprainea marginata f. bremondialis Janssens F & Colombo M, 2020
Caprainea marginata f. bremondialis from Italy
2020.10.28 © Colombo, M.
Caprainea marginata f. bremondialis from Italy
Dorsal pattern
2020.10.28 © Colombo, M.
Caprainea marginata f. bremondialis from Italy
Antenna
2020.10.28 © Colombo, M.
Caprainea marginata f. bremondialis from Italy
Eye with ocelli C+D distinctly smaller
2020.10.28 © Colombo, M.
Caprainea marginata f. bremondialis from Italy
2020.10.28 © Colombo, M.
Caprainea marginata f. bremondialis from Italy
Spiky dorsal macrosetae
2020.10.28 © Colombo, M.
Caprainea marginata f. bremondialis from Italy
2020.11.04 © Colombo, M.
Caprainea marginata f. bremondialis from Italy
Well pigmented
2020.11.04 © Colombo, M.
Caprainea marginata f. bremondialis from Italy
Ocelli C+D distinctly smaller
2020.11.04 © Colombo, M.
Caprainea marginata f. bremondialis from Italy
Ant.4 with 15 whorls of setae
2020.11.04 © Colombo, M.
Caprainea marginata f. bremondialis from Italy
2020.11.07 © Colombo, M.
Caprainea marginata f. bremondialis from Italy
Spiky dorsal macrosetae
2020.11.07 © Colombo, M.
Caprainea marginata f. bremondialis juv. from Italy
2020.11.21 © Colombo, M.




Sminthurus marginatus from Sweden
After Schött H, 1893 Taf.I Fig.11
Sminthurus marginatus from Corsica
After Denis J-R, 1947 Fig.4a
Sminthurus marginatus from Poland
After Stach J, 1956 PlateXXXI Fig.1
Caprainea marginata from Hungary
2007.mm.dd © Pfliegler, W.
Caprainea marginata from France
2008.11.03 © Lebeaux, P.
Caprainea marginata from France
2008.11.03 © Lebeaux, P.
Caprainea marginata from France
2008.11.03 © Lebeaux, P.
Caprainea marginata from France
2010.12.10 © Nicolosi., B
Caprainea marginata from the UK
2013.08.22 © Murray, A.
Caprainea marginata from the UK
2013.08.22 © Murray, A.
Caprainea marginata from France
2017.10.14 © Picard, J.
Caprainea marginata from France
Spiky abdominal macrosetae
2017.10.14 © Picard, J.
Caprainea marginata from France
2018.03.05 © Tiky, S.
Caprainea marginata from France
2018.10.29 © Picard, J.
Caprainea marginata from France
2020.01.26 © Nogues, Be.
Caprainea marginata from France
2020.02.13 © Garcelon, P.
Caprainea marginata from France
Distinctly smaller ocelli C and D
2020.02.13 © Garcelon, P.
Caprainea marginata from France
Spiky abdominal macrosetae
2020.02.13 © Garcelon, P.
Caprainea marginata from France
2021.09.25 © Joachim, J.
Caprainea marginata from France
Bothriotricha ABC in straight line
2021.11.10 © Fiacre, R.
Caprainea marginata from France
Eye with distinctly smaller ocelli C+D
2021.11.10 © Fiacre, R.



Genus Galeriella Curcic BPM et Lucic LR in Curcic, BPM, Lucic, LR, Tomic, VT, Makarov, SE et Karaman, IM, 2007:63P

Species liciniana Curcic BPM et Lucic LR in Curcic, BPM, Lucic, LR, Tomic, VT, Makarov, SE et Karaman, IM, 2007:64P, t.t.

Galeriella liciniana from Serbia
After Curcic BPM et al., 2007 Fig.1
Galeriella liciniana Croatia
After Lukic M, 2019 Fig.2C

References