http://www.collembola.org/taxa/willinae.htm - Last updated on 2019.08.09 by Frans Janssens
Checklist of the Collembola: Willowsiinae

Subfamilia Willowsiinae Yoshii R & Suhardjono YR, 1989:35 key  key  key  key

Genus Number
of species
Americabrya3
Desertia3
Drepanosira18
Epimetrura4
Hawinella2
IncertaeSedis4
Janetschekbrya2
Lepidobrya4
Lepidocyrtoides22
Lepidodens3
Lepidosinella1
Lepidosira59
Szeptyckiella3
Willowsia35
Willowsiinae163
Genera with number of species
  Syn.: Entomobryidae Willowsini(sic) Yoshii, R et Suhardjono, YR, 1989:35 ISS


  Syn.: Siraeformes Willowsini(sic) Yosii & Suhardjono, 1989 ISS
Willowsia buski from France
2007.06.09 © Lebeaux, P.
Willowsia platani from the UK
2007.06.30 © Cornwall, N.J.

  Syn.: Entomobryinae Willowsini(sic) Soto-Adames, FN, Barra, J-A, Christiansen, K et Jordana R, 2008:502 ISS
Remark by Janssens, F. 2016.12.26: Yoshii & Suhardjono, 1989 defined Willowsiini as a tribe of Entomobryidae. In this checklist, the polyphyletic scaled Entomobryinae sensu Zhang et al, 2016 are maintained at subfamily level mainly for convenience.





Genus IncertaeSedis Zhang, F, Chen, J-X et Deharveng, L, 2011:8

Species brahma (Imms, 1912) Zhang, F, Chen, J-X & Deharveng, L, 2011:8

             Syn.: Sira(sic) brahama(sic) Denis, JR, 1936:266 ISS
             Syn.: Sira(sic) brahama(sic) Yosii, R, 1942:490 ISS

Species crassicornis (Nicolet, H, 1847:370) Janssens, 2016

            Syn.: Sira(sic) intermedia Schött, 1921 =?


Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Species intermedia (Schött, 1921) Zhang, F, Chen, J-X & Deharveng, L, 2011:8

Species membranea (Nicolet, H, 1847:371) Janssens, 2016

            Syn.: Entomobrya multifasciata Handschin, 1924


Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

             Syn.: Degeeria Membranacea(sic) Kolenati, FA, 1858:132 ISS

Tribus Willowsiini Yoshii R & Suhardjono YR, 1989:35




Genus Willowsia Shoebotham, 1917 key  key  key

  Syn.: ... Seira Lubbock, J, 1873:143 partim
  Syn.: Collembola Sira(sic) Cassagnau, P, 1962:229 ISS


Willowsia platani from the UK
Apically pointed scales
2013.12.24 © Murray, A.
Willowsia nigromaculata from the USA
Apically pointed scales
2015.02.13 © Dean, T. & S.B.
Willowsia nr nigromaculata from Australia
Apically pointed scales
2015.10.01 © Anonymous.
Willowsia sp. from Ireland
Apically pointed scales with long basal ribs
2015.11.26 © Gaff, K.
Willowsia sp. from Ireland
Broad apically pointed scales with long uninterrupted apically raised ribs
2015.11.26 © Gaff, K.
Willowsia nigromaculata from the UK
Apically pointed backwards directed scales
2016.01.14 © Horton, M.

Species sp.1 Zhang, F, Chen, J-X et Deharveng, L, 2011:3

Species abrupta (Schött, H, 1917:30) Greenslade, P in Houston, WWK, 1994:102

Sira(sic) abrupta from Australia
After Schött H, 1917 Fig.36

Species baoshanensis Chai, R et Ma Y-T, 2017:2

Willowsia baoshanensis from China
After Chai R & Ma Y-T, 2017 Fig.1B
Willowsia baoshanensis from China
After Chai R & Ma Y-T, 2017 Fig.1C
Willowsia baoshanensis from China
Spinulate scale
After Chai R & Ma Y-T, 2017 Fig.3G

Species bartkei Stach, J, 1965:361

Willowsia bartkei from Vietnam
After Zhang F, 2015 Fig.26

Species buski (Lubbock, J, 1870:280) Schoebotham, 1917, B

            Syn.: Seira Buskii(sic) Lubbock, J, 1870:280 IOS
            Syn.: Degeeria pruni Nicolet, H, 1842:73
            Syn.: Degeeria cyanea Tullberg, 1871
            Syn.: Sira flava Ågren, 1903
            Syn.: Sira corticaloides Stach, 1922


             Syn.: Sira(sic) Buskii(sic) Tullberg, 1872 ISS
             Syn.: Seira Buskii(sic) Lubbock, J, 1873:145 ISS
Seira Buskii(sic) from the UK
After Lubbock J, 1873 Pl.22
Seira Buskii(sic) from the UK
Apically pointed scale with long basal ribs
After Beck J in Lubbock J, 1873 Pl.70

             Syn.: Seira Buskii(sic) Beck, J in Lubbock, J, 1873:253 ISS
             Syn.: Sira(sic) Buskii(sic) Reuter, OM, 1876:81 ISS
             Syn.: Sira(sic) buskii(sic) Tullberg, 1872 ISS
             Syn.: Seira Buski(sic) Lameere, A, 1895:593,B+ ISS
             Syn.: Sira(sic) Buski(sic) Carl, J, 1899:343 ISS
             Syn.: Sira(sic) pruni var. Buskii(sic) Börner, C, 1901:11 ISS
             Syn.: Sira(sic) busquii(sic) Krausbauer, 1902 ISS
             Syn.: Seira Buskii(sic) Schött, H, 1917:33 ISS
             Syn.: Sira(sic) Buski(sic) Stach, J, 1930:299 ISS
             Syn.: Willowsia buskii(sic) Salmon, JT, 1945:71 ISS
             Syn.: Seira buskii(sic) Kloet, GS et Hincks, WD, 1945:8 ISS
             Syn.: Willowsia buskii(sic) Palacios-Vargas & Gómez, 1991 ISS
             Syn.: Willowsia buskii(sic) Skidmore, 1995 ISS
Willowsia buskii(sic)
After Zhang, Palacios-Vargas & Chen, 2007 Fig.18a.

             Syn.: Willowsia buskii(sic) Fiera, C, 2007:27 ISS
             Syn.: Willowsia buskii(sic) Zhang, F, Palacios-Vargas, JG et Chen, J-X, 2007:36 ISS
Willowsia buski from Germany
2006 © Kohl, F.
Willowsia buski from the USA
2006 © McClarin, J.
Willowsia buski from Germany
2006.12.27 © Kohl, F.
Willowsia buski from Germany
2006.12.30 © Kohl, F.
Willowsia buski from the USA
2007 © McClarin, J.
Willowsia buski juvenile from the USA
2007.02.25 © Murray, T.
Willowsia buski juvenile from the USA
2007.02.25 © Murray, T.
Willowsia buski subadult from the USA
2007.02.25 © Murray, T.
Willowsia buski adult from the USA
2007.02.25 © Murray, T.
Willowsia buski from Canada
2007.04.18 © Luk, S.
Willowsia buski from the USA
2007.04.22 © Reed, D.
Willowsia buski from the USA
2007.04.22 © Reed, D.
Willowsia buski from the USA
2007.04.22 © Reed, D.
Willowsia buski from Hungary
2008.10.23 © Keresztes, G.
Willowsia buski from the USA
2009.04.24 © Zungri, D.
Willowsia buski from Holland
2010.01.09 © van Duinen, J.
Willowsia buski from Czechia
2010.08.02 © Deml, M.
Willowsia buski from Czechia
2010.08.05 © Deml, M.
Willowsia buski from the USA
2011.06.07 © Nanz, S.
Willowsia buski juv. from the UK
Descaled specimen
2012.08.06 © Murray, A.
Willowsia buski juv. from the UK
2012.08.25 © Murray, A.

             Syn.: Willowsia buskii(sic) Soto-Adames, FN et Taylor, SJ, 2013:63 ISS
Willowsia buski from the USA
2014.11.10 © Cristofono, P.

             Syn.: Willowsia buskii(sic) Pan, Z et Zhang, F, 2016:11 ISS
             Syn.: Willowsia buskii(sic) Pan, Z et Zhang, F, 2016:15 ISS
             Syn.: Willowsia buskii(sic) Pan, Z et Zhang, F, 2016:23 ISS
             Syn.: Willowsia buskii(sic) Soto-Adames, FN, 2016:3 ISS
Willowsia buski from the USA
After Katz AD, 2017 Fig.1
Willowsia buski from the USA
Scales with interrupted ribs
After Katz AD, 2017 Fig.12
Willowsia buski from the USA
2017.01.04 © Dobbs, Br.
Willowsia buski juv. from the USA
2019.01.22 © Duncan, W.L.
Willowsia buski juv. from the USA
Ant.1 yellowish; Ant.2-4 bluish
2019.01.22 © Duncan, W.L.





Species cassagnaui Zhang, F, 2015:490

Willowsia cassagnaui from Nepal
After Zhang F, 2015 Fig.1

Species fascia Zhang F et Pan Z in Pan, Z et Zhang, F, 2016:3

Willowsia fascia from China
After Pan Z & Zhang F, 2016 Fig.1A
Willowsia fascia from China
Body scale with long basal ribs
After Pan Z & Zhang F, 2016 Fig.1B

Species formosana (Denis, 1929) Yosii, R, 1956:73

            Syn.: Sira(sic) gibbosa Uchida, 1958


Species fuscana (Uchida, 1944)

Species guangdongensis Zhang, F, Xu, G-L et Chen, J-X, 2007:?

Species guangxiensis Shi & Chen, 2004

Species hyalina (Handschin, 1928) Yosii, 1966

Species ieti Yosii, 1971

Species jacobsoni (Börner, 1913) Stach, 1965

            Syn.: Willowsia jacobsoni var. lipostropha Börner, 1913
            Syn.: Sira(sic) tricincta Schött, H, 1917:31
Sira(sic) tricincta from Australia
After Schött H, 1917 Fig.37


            Syn.: Sira(sic) jacobsoni var. indica Handschin, E, 1929:240
            Syn.: Sira(sic) parajacobsoni Denis, 1929
            Syn.: Sira(sic) abrupta Womersley, 1934
            Syn.: Sira(sic) abrupta Womersley, 1937
            Syn.: Sira(sic) abrupta Womersley, 1942
            Syn.: Sira(sic) jacobsoni var. handschini Uchida, 1944
            Syn.: Sira(sic) jacobsoni var. africana Delamare Deboutteville, 1951
            Syn.: Willowsia mesothoraxa Nguyen, 2001



             Syn.: Sira(sic) Jacobsoni(sic) Denis, JR, 1936:266 ISS
Willowsia jacobsoni
After Zhang, Palacios-Vargas & Chen, 2007 Fig.18c.
Willowsia jacobsoni ♂ from Puerto Rico
After Katz AD, 2017 Fig.3
Willowsia jacobsoni ♀ from Puerto Rico
After Katz AD, 2017 Fig.4
Willowsia jacobsoni from Puerto Rico
Scales with interrupted ribs
After Katz AD, 2017 Fig.11



Species japonica (Folsom, JW, 1898:55) Yosii, 1955

Seira japonica from Japan
After Folsom JW, 1898 Fig.15
            Syn.: Ptenura bimaculata Börner, C, 1909:121
             Syn.: Sira(sic) binoculata(sic) Denis, JR, 1936:265 ISS
            Syn.: Sira(sic) bimaculata Yosii, R, 1942:488
Sira(sic) bimaculata from Japan
After Yosii R, 1942 Abb.11.1
Sira(sic) bimaculata from Japan
After Yosii R, 1942 Abb.11.2

            Syn.: Sira(sic) (Sira(sic)) gibbosa Gisin, 1944 =?
            Syn.: Willowsia kahlertae Christiansen, K et Bellinger, P, 1992:254
            Syn.: Willowsia qui Zhang, F, Chen, J-X et Deharveng, L, 2011:9 partim
            Syn.: Willowsia qui Pan, Z et Zhang, F, 2016:23



Sira(sic) japonica from Japan
After Yosii R, 1942 Abb.18.1
Willowsia japonica ♀ from Japan
After Yoshii, 1992
Willowsia japonica ♂ from Japan
After Yoshii, 1992
Willowsia japonica from Japan
2005.05.12 © Keresztes, G.
Willowsia japonica from Japan
2005.05.12 © Keresztes, G.
Willowsia japonica ♀ from China
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.10A
Willowsia japonica ♀ from China
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.10B
Willowsia japonica ♀ from China
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.10C
Willowsia japonica ♀ from Japan
2013.03.12 © Hara, Y.

Species kalatopensis Baquero E et Jordana R in Baquero, E, Mandal, G et Jordana, R, 2015:35

            Syn.: Willowsia kalatopii(sic) Baquero E et Jordana R in Baquero, E, Mandal, G et Jordana, R, 2015:35 IOS
Willowsia kalatopii(sic) from India
After Baquero E & al, 2015 Fig.22



Species mekila Christiansen, K et Bellinger, P, 1992:256

Species mexicana Zhang, F, Palacios-Vargas, JG et Chen, J-X, 2007:37

            Syn.: Willowsia nigromaculata Palacios-Vargas, 1997


Willowsia mexicana from Mexico
After Zhang, Palacios-Vargas & Chen, 2007 Fig.1
Willowsia mexicana from Mexico
Scales with uninterrupted ribs
After Katz AD, 2017 Fig.8-10

Species neocaledonica Zhang, F, Bedos, A et Deharveng, L, 2014:382

Willowsia neocaledonica from New Caledonia
After Zhang F & al, 2014 Fig.1
             Syn.: Willowsia neocaledonia(sic) Katz, AD, 2017:549 ISS
             Syn.: Willowsia neocaledonia(sic) Katz, AD, 2017:553 ISS
             Syn.: Willowsia neocaledonia(sic) Katz, AD, 2017:556 ISS
             Syn.: Willowsia neocaledonia(sic) Katz, AD, 2017:560 ISS


Species nigra Zhang, F, Bedos, A et Deharveng, L, 2014:384

Species nigromaculata (Lubbock, J, 1873:146) Shoebotham, 1917, t.t., B

            Syn.: Sira(sic) elongata Nicolet
            Syn.: Seira mimica Harvey, 1894
            Syn.: Lepidocyrtus americanus Marlatt, 1896
Lepidocyrtus americanus from North America
Dorsal aspect
After Howard LO, 1901 Fig.246 left
Lepidocyrtus americanus from North America
Ventral aspect
After Howard LO, 1901 Fig.246 right
Lepidocyrtus americanus from North America
Lateral aspect
After Howard LO, 1901 Fig.247

            Syn.: Lepidocyrtus americanus Christiansen, K et Bellinger, P, 1980:920 =?
            Syn.: Lepidocyrtus americanus Christiansen, K et Bellinger, P, 1998:1020 =?
            Syn.: Degeeria platani Goto, HE et Lawrence, PN in Kloet, GS et Hincks, WD, 1964:9
            Syn.: Willowsia bimaculata Yosii, R, 1977:161 =?



Willowsia nigromaculata from Europe
After Gisin H, 1960 Abb.433
Willowsia nigromaculata from the USA
2006 © Bernard, E.C.
Willowsia nigromaculata from Germany
2006.03.27 © Rodeland, J.
Willowsia nigromaculata from Germany
2006.08.24 © Kohl, F.
Willowsia nigromaculata from Germany
2006.08.24 © Kohl, F.
Willowsia nigromaculata from the USA
2007.03.23 © Bradford, A.
Willowsia nigromaculata from Belgium
2007.05.10 © Janssens, F.
Willowsia nigromaculata from the UK, juvenile
2007.06.09 © Barton, T.
Willowsia nigromaculata from the UK, juvenile
2007.06.09 © Barton, T.
Willowsia nigromaculata from the UK, juvenile
2007.06.09 © Barton, T.
Willowsia nigromaculata from the USA
caught by an ant...
2007.06.13 © DiTerlizzi, T.
Willowsia nigromaculata from the UK
2007.08.22 © Valentine., B
Willowsia nigromaculata from the UK
2007.09.17 © Valentine., B
Willowsia nigromaculata from Hungary
2008.01.12 © Keresztes, G.
Willowsia nigromaculata from Hungary
2008.03.04 © Keresztes, G.
Willowsia nigromaculata from Hungary
2008.03.16 © Keresztes, G.
Willowsia nigromaculata from the USA
2008.03.23 © Tortuga, I.
Willowsia nigromaculata from the USA
2008.04.05 © Justis, P.S.
Willowsia nigromaculata from the USA
2008.09.11 © Bradford, A.
Willowsia nigromaculata from Hungary
2008.10.19 © Keresztes, G.
Willowsia nigromaculata from Russia
2008.10.21 © Macroclub.ru
Willowsia nigromaculata from the USA
2008.12.17 © Bradford, A.
Willowsia nigromaculata from Canada
2009.01.07 © Luk, S.
Willowsia nigromaculata from the USA
2009.05.04 © Murray, T.
Willowsia nigromaculata from the USA
2009.07.23 © Bradford, A.
Willowsia nigromaculata from Czechia
2009.09.14 © Deml, M.
Willowsia nigromaculata from Belgium
2009.12.16 © De Bruyn, L.
Willowsia nigromaculata
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.11B
Willowsia nigromaculata
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.11C
Willowsia nr nigromaculata from Australia
2011.04.21 © Flyman.
Willowsia nigromaculata from the UK
2011.10.05 © Shaw, P.
Willowsia nigromaculata from the USA
2011.10.13 © Schmidt, M.L.
Willowsia nigromaculata juv. from the USA
2012.01.08 © Allen, K.
Willowsia nigromaculata from the USA
2013.04.12 © Eiseman, C.
Det. Soto-Adames, F.N., 2017.12.28
Willowsia nr nigromaculata from Australia
2014.12.08 © Flyman.
Willowsia nigromaculata from the USA
2014.12.17 © Snyder, S.L.
Det. Soto-Adames, F.N., 2017.12.28
Willowsia nigromaculata from the USA
2014.12.19 © Snyder, S.L.
Det. Soto-Adames, F.N., 2017.12.28
Willowsia nigromaculata from the USA
Descaled
2014.12.19 © Snyder, S.L.
Det. Soto-Adames, F.N., 2017.12.28
Willowsia nigromaculata from the USA
Descaled
2014.12.19 © Snyder, S.L.
Det. Soto-Adames, F.N., 2017.12.28
Willowsia nigromaculata from the USA
Banded antennae
2014.12.19 © Snyder, S.L.
Det. Soto-Adames, F.N., 2017.12.28
Willowsia nr nigromaculata from Australia
Dorsal aspect
2015.10.01 © Anonymous.
Willowsia nr nigromaculata from Australia
Lateral aspect
2015.10.01 © Anonymous.
Willowsia nr nigromaculata from Australia
Ventral aspect
2015.10.01 © Anonymous.
Willowsia nr nigromaculata from Australia
Head
2015.10.01 © Anonymous.
Willowsia nigromaculata from the USA
2015.03.04 © Snyder, S.L.
Det. Soto-Adames, F.N., 2017.12.28
Willowsia nigromaculata from the UK
2016.01.14 © Horton, M.
Willowsia nigromaculata from the UK
Caudal aspect cephalic mimicry
2016.01.14 © Horton, M.
Willowsia nigromaculata from the USA
After Katz AD, 2017 Fig.2
Willowsia nigromaculata from the USA
Scales with interrupted ribs
After Katz AD, 2017 Fig.13
Willowsia nigromaculata from the USA
2017.01.04 © Dobbs, Br.
Willowsia nigromaculata from the USA
2017.01.13 © Snyder, S.L.
Det. Soto-Adames, F.N., 2017.12.28
Willowsia nigromaculata from the USA
2017.01.13 © Snyder, S.L.
Det. Soto-Adames, F.N., 2017.12.28
Willowsia nigromaculata from Belgium
2017.07.16 © Huskens, M.-L.
Willowsia nigromaculata from Belgium
Apically pointed scales
2017.07.16 © Huskens, M.-L.

Species nivalis Yosii, 1971

            Syn.: Willowsia brahmides Yosii, 1966


Species platani (Nicolet, H, 1842:72) Palissa, 1959, B

Degeeria platani from Switzerland
After Nicolet H, 1842 Pl.8 Fig.4
            Syn.: Podura argenteo cincta Bourlet, 1842
            Syn.: Degeeria platani var. var. Nicolet, H, 1842:73
            Syn.: Sira(sic) elongata Tullberg, 1871, nec Nicolet, 1842
            Syn.: Seira mimica Harvey, 1894
            Syn.: Sira(sic) flava Ågren, 1903
            Syn.: Sira(sic) corticaloides Stach, 1922
            Syn.: Sira(sic) platani ab. corticalis Stach, J, 1930:300
            Syn.: Lepidocyrtus americanus Mills, HB, 1934:71
            Syn.: Lepidocyrtus americanus Denis, 1941
            Syn.: Lepidocyrtus americanus Salmon, JT, 1964:487
            Syn.: Sira(sic) nigromaculata Mills, HB, 1934:71
            Syn.: Willowsia japonica Yosii, R, 1977:161 =?



             Syn.: Degeeria Platani(sic) Lubbock, J, 1862:595 ISS
             Syn.: Sira(sic) plantani(sic) Bryk, F, 1955:1823 ISS
             Syn.: Willowsia planati(sic) Gisin, H, 1960:234 ISS
Willowsia platani from Europe
Modified after Gisin H, 1960 Abb.434
Willowsia platani from Europe
Modified after Gisin H, 1960 Abb.434

             Syn.: Sira(sic) platani Cassagnau, P, 1962:229 ISS
Willowsia platani from the UK
Leaf-shaped scales
2005 © Hopkin, S.P.
Willowsia platani from Germany
2006 © Kohl, F.
Willowsia platani from Germany, juvenile
2007 © Peters, J.
Willowsia platani from Germany, juvenile
2007.03.14 © Peters, J.
Willowsia platani from Czechia
2007.07.22 © Krásenský, P.
Willowsia platani from Germany, juvenile
2008.01.11 © Peters, J.
Willowsia platani from the UK, juvenile
2008.04.23 © Barton, T.
Willowsia platani from the UK
2008.05.24 © Barton, T.
Willowsia platani from France
2008.07.09 © Lavoué., B
Willowsia platani from the UK
2009.07.04 © Talbot, M.E.
Willowsia cf. platani from the USA
2009.07.12 © Zungri, D.
Willowsia platani from the UK
2010.03.24 © Barton, T.
Willowsia platani from Europe
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.11A
Willowsia platani from the UK
2011.06.30 © Shaw, P.
Willowsia platani from the UK
2011.06.30 © Shaw, P.
Willowsia platani from France
2011.10.02 © Hugues, C.
Willowsia platani juv. from the UK
2011.12.11 © Barton, T.
Willowsia platani from Portugal
2012.04.20 © Coelho, J.
Willowsia platani from Spain
2012.09.23 © Campo., B
Willowsia platani juv. from the UK
2012.12.15 © Barton, T.
Willowsia platani from Spain
2013.04.25 © Baquero, E.
Willowsia platani from Holland
2014.01.24 © van Duinen, J.
Willowsia platani from Holland
2014.01.24 © van Duinen, J.
Willowsia platani from Czechia
2014.05.01 © Miroslav, D.
Willowsia platani from the UK
2015.02.09 © Barton, T.
Willowsia platani from Holland
2015.02.27 © Baas, A.H.
Willowsia platani from Holland
2015.02.27 © Baas, A.H.
Willowsia platani from Holland
2015.02.27 © Baas, A.H.
Willowsia platani juv. from Holland
2015.02.27 © Baas, A.H.
Willowsia platani from Holland
2017.09.15 © Wieringa, P.
Willowsia nr platani juv. from France
2017.09.29 © Garcelon, P.



Species potapovi Zhang, F, Chen, J-X et Deharveng, L, 2011:13

Willowsia potapovi from Azerbaijan
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.8A
Willowsia potapovi from Azerbaijan
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.8B

Species pseudobuski Pan, Z et Zhang, F, 2016:11

            Syn.: Willowsia pseudobuskii(sic) Pan, Z et Zhang, F, 2016:11 IOS
Willowsia pseudobuskii(sic) from China
After Pan Z & Zhang F, 2016 Fig.7A
Willowsia pseudobuskii(sic) from China
Body scale with long basal ribs
After Pan Z & Zhang F, 2016 Fig.7E

             Syn.: Willowsia pseudobuskii(sic) Pan, Z et Zhang, F, 2016:23 ISS
             Syn.: Willowsia pseudobuskii(sic) Chai, R et Ma Y-T, 2017:10 ISS




Species pseudoplatani Zhang F et Pan Z in Pan, Z et Zhang, F, 2016:7

Willowsia pseudoplatani from China
After Pan Z & Zhang F, 2016 Fig.4A
Willowsia pseudoplatani from China
Body scales with long basal ribs
After Pan Z & Zhang F, 2016 Fig.4B

Species pseudosocia Stach, J, 1965:363

Species pyrrhopygia Katz, AD, 2017:554

Willowsia pyrrhopygia from the USA
After Katz AD, 2017 Fig.20
Willowsia pyrrhopygia from the USA
After Katz AD, 2017 Fig.21
Willowsia pyrrhopygia from the USA
Scales with uninterrupted ribs
After Katz AD, 2017 Fig.5-7
            Syn.: Willowsia sp.nov.1 Katz, AD, Giordano, R et Soto-Adames, FN, 2015:8
            Syn.: Willowsia sp.n.1 Katz, AD, Giordano, R et Soto-Adames, F, 2015:7


Species qui Zhang, F, Chen, J-X et Deharveng, L, 2011:9 partim

Willowsia qui from China
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.6A
Willowsia qui from China
After Pan Z & Zhang F, 2016 Fig.13A
Willowsia qui from China
Body scale with long basal ribs
After Pan Z & Zhang F, 2016 Fig.13B

Species samarkandica Martynova, 1972

Species shiae Pan, J-L, Zhang, F et Chen, J-X, 2006:261

Willowsia shiae from China
After Pan J-L, Zhang F & Chen J-X, 2006 Fig.1

             Syn.: Willowsia shi(sic) Chai, R et Ma Y-T, 2017:3 ISS
             Syn.: Willowsia shi(sic) Chai, R et Ma Y-T, 2017:6 ISS
             Syn.: Willowsia shi(sic) Chai, R et Ma Y-T, 2017:11 ISS


Species similis Pan, Z et Zhang, F, 2016:15

Willowsia similis from China
After Pan Z & Zhang F, 2016 Fig.10A
Willowsia similis from China
Body scale with long basal ribs
After Pan Z & Zhang F, 2016 Fig.10D

Species yamashitai Uchida, 1969

Species yiningensis Zhang, F, Chen, J-X et Deharveng, L, 2011:12

Species zhaotongensis Chai, R et Ma Y-T, 2017:6

Willowsia zhaotongensis from China
After Chai R & Ma Y-T, 2017 Fig.4B
Willowsia zhaotongensis from China
Spinulate scale
After Chai R & Ma Y-T, 2017 Fig.6G


Genus Lepidodens Zhang F et Pan Z-X in Zhang, F, Pan, Z-X, Wu, J, Ding, Y-H, Yu, D-Y et Wang, B-X, 2016:600

Species hainanicus Zhang, F, Pan, Z-X, Wu, J, Ding, Y-H, Yu, D-Y et Wang, B-X, 2016:610

Lepidodens hainanicus from South China
After Zhang F & Pan Z-X in Zhang F & al, 2016 Fig.3C

             Syn.: Lepidodens hananicus(sic) Zhang, F, Pan, Z-X, Wu, J, Ding, Y-H, Yu, D-Y et Wang, B-X, 2016:602 ISS


Species nigrofasciatus Zhang F et Pan Z-X in Zhang, F, Pan, Z-X, Wu, J, Ding, Y-H, Yu, D-Y et Wang, B-X, 2016:602, t.t.

Lepidodens nigrofasciatus from South China
After Zhang F & Pan Z-X in Zhang F & al, 2016 Fig.3A

Species similis Zhang F et Pan Z-X in Zhang, F, Pan, Z-X, Wu, J, Ding, Y-H, Yu, D-Y et Wang, B-X, 2016:607

Lepidodens similis from South China
After Zhang F & Pan Z-X in Zhang F & al, 2016 Fig.3B


Genus Janetschekbrya Yosii, 1971 key

  Syn.: Xenyllodes pallescens Scott, HG, 1960:186 partim
  Syn.: Entomobryidae Janetschekbra(sic) Mari Mutt, JA et Palacios-Vargas, JG, 1987:100 ISS

Species brahamides (Denis, JR, 1936:265) Yosii, 1971

Sira(sic) brahamides from India
After Denis JR, 1936 Fig.13

Species himalica Yosii, 1971, t.t.


Genus Americabrya Mari Mutt, JA et Palacios-Vargas, JG, 1987:99 key

  Syn.: Entomobryidae Janetschekbrya Mari Mutt & Palacios-Vargas, 1987 partim


Species arida (Christiansen, K & Bellinger, P, 1980:918) Mari Mutt, JA & Palacios-Vargas, JG, 1987:100, t.t.

            Syn.: Janetschekbrya sp. Palacios-Vargas, 1979
            Syn.: Janetschekbrya sp. Palacios-Vargas, 1981


Americabrya arida from the USA
2015.03.17 © Landerman, J.
Americabrya arida from the USA
2015.03.17 © Landerman, J.
Americabrya arida from the USA
2015.03.23 © Christopherson, S.
Americabrya arida from Mexico
Narrow scales with uninterrupted ribs
After Katz AD, 2017 Fig.14

Species epiphyta (Loring, 1984) Mari Mutt, JA & Palacios-Vargas, JG, 1987:107

Species matthewsi (Snider, RJ, 1981:39) Mari Mutt, JA & Palacios-Vargas, JG, 1987:107


Genus Drepanosira Bonet, F, 1942:57

Species albescens Stach, J, 1960:538

Species altimontana Stach, 1963

Species frigida (Imms, 1912)

Species gisini Nosek, 1964

Species grisescens Stach, J, 1960:532

Species hussi Neuherz, 1976

Drepanosira hussi from India
After Baquero E & al, 2015 Fig.17

Species lineata Stach, J, 1960:533

Species ombrophila Stach, J, 1960:535

Species ornata (Bonet, 1930) Bonet, F, 1942:58, t.t.

            Syn.: Parasira subornata Denis, 1936


Species ornata (Stach, 1963 nec Bonet, 1930) Baquero, E, Mandal, G & Jordana, R, 2015:34

Species pallida (Brown, JM, 1926:209)

Species pulchra Stach, J, 1960:534

Species rara Stach, J, 1960:537

Species raviensis Baquero E et Jordana R in Baquero, E, Mandal, G et Jordana, R, 2015:32

            Syn.: Drepanosira ravi(sic) Baquero E et Jordana R in Baquero, E, Mandal, G et Jordana, R, 2015:32 IOS
Drepanosira ravi(sic) from India
After Baquero E & al, 2015 Fig.18
Drepanosira ravi(sic) from India
After Baquero E & al, 2015 Fig.19



Species shimlaensis Baquero E et Jordana R in Baquero, E, Mandal, G et Jordana, R, 2015:34

            Syn.: Drepanosira shimlai(sic) Baquero E et Jordana R in Baquero, E, Mandal, G et Jordana, R, 2015:34 IOS
Drepanosira shimlai(sic) from India
After Baquero E & al, 2015 Fig.20
Drepanosira shimlai(sic) from India
After Baquero E & al, 2015 Fig.21



Species stachi Nosek, 1964

Species subornata (Denis, JR, 1936:267) Bonet, F, 1942:58

Parasira subornata from India
After Denis JR, 1936 Fig.21
Parasira subornata from India
After Denis JR, 1936 Fig.22

Species villosa (Börner, 1903)


Genus Desertia Tshelnokov, 1979

Species kaplini Tshelnokov, 1979, t.t.

Species pulchra (Handschin, 1928)

             Syn.: Seira pulcher(sic) Mari Mutt, JA et Bellinger, PF, 1990:106 ISS
             Syn.: Seira pulcher(sic) Culik, MP et Zeppelini, DF, 2003:1129 ISS

Species semicolorata (Handschin, 1928)


Genus Hawinella Bellinger & Christiansen, 1974

Species kuaola Christiansen, K et Bellinger, P, 1992:260

Species lava Bellinger, PF et Christiansen, KA, 1974:36, t.t.


Genus Szeptyckiella Zhang, F, Bedos, A et Deharveng, L, 2014:2 key

Species boulouparica Zhang, F, Bedos, A et Deharveng, L, 2014:4, t.t.

Szeptyckiella boulouparica from New Caledonia
After Zhang F, Bedos A & Deharveng L, 2014 Fig.1A

Species lii Zhang, F, Bedos, A et Deharveng, L, 2014:12

Szeptyckiella lii from South China
After Zhang F, Bedos A & Deharveng L, 2014 Fig.7A

Species sinelloides Zhang, F, Bedos, A et Deharveng, L, 2014:8

Szeptyckiella sinelloides from New Caledonia
After Zhang F, Bedos A & Deharveng L, 2014 Fig.4A


Genus Lepidosinella Handschin, 1920

Species armata Handschin, 1920, t.t.


Tribus Lepidosirini Yoshii R & Suhardjono YR, 1989:36 sensu Zhang F & al, 2017:534


Genus Lepidobrya Womersley, 1937

Lepidobrya? sp. from Tasmania
2015.09.26 © Tony.
Lepidobrya? sp. from Tasmania
Apically pointed scales
2015.09.26 © Tony.
Lepidobrya? sp. from Tasmania
2016.09.10 © Tony.
Lepidobrya? sp. from Tasmania
Apically pointed scales
2016.09.10 © Tony.

Species mawsoni (Tillyard, 1920) Greenslade P, 1990, t.t.

            Syn.: Lepidobrya aurantiaca Salmon, JT, 1949:46
             Syn.: Lepidobrya aurantica(sic) Greenslade P, 1990 ISS
             Syn.: Lepidobrya aurantica(sic) Greenslade, P in Houston, WWK, 1994:93 ISS


Lepidobrya mawsoni from Antarctica
After Davis, G. in Greenslade, P., 2006 Fig.76.
Lepidobrya mawsoni from Macquarie Island
After Zhang F, Greenslade P & Stevens MI, 2017 Fig.1
Lepidobrya mawsoni from Macquarie Island
After Zhang F, Greenslade P & Stevens MI, 2017 Fig.2
Lepidobrya mawsoni from Macquarie Island
Narrow pointed body scales
After Zhang F, Greenslade P & Stevens MI, 2017 Fig.5

Species mitchelli (Womersley, 1934) Greenslade, P in Houston, WWK, 1994:93

Entomobrya mitchelli from Australia
After Womersley H, 1939 Fig.58M

Species thalassarchia Salmon, JT, 1949:44

Species violacea Salmon, JT, 1949:46


Genus Lepidosira Schött, 1925

  Syn.: Entomobryini Urewera Salmon, JT, 1938:349
  Syn.: Entomobryinae Urewara(sic) Womersley, H, 1939:322 ISS


Lepidosira sp. from Australia
2008.12.10 © Bert
Lepidosira? sp. from Australia
2015.02.28 © Murray, A.

Species sp.1 Yosii, R, 1940:116

Species sp.2 Yayuk, 1989:10

            Syn.: Lepidocyrtus variabilis Oudemans, JT in Weber, M, 1890:74

Species alba (Nguyen, TT, 2005:8) Zhang, F, Chen, J-X & Deharveng, L, 2011:5

Species angulata (Schött, H, 1917:46) Howard, GW, 1969:189

Lepidocyrtoides angulatus from Australia
After Schött H, 1917 Taf.3 Fig.5




             Syn.: Lepidocyrtus (Lepidocyrtoides) angustatus(sic) Schött, 1925 ISS

Species anomala Salmon, JT, 1944:164

Species arborea Salmon, JT, 1944:163

Species australica (Schött, H, 1917:40) Greenslade P & Majer in Dindal, 1980

Lepidocyrtus australicus from Australia
After Schött H, 1917 Taf.3 Fig.1

  Subspecies tamburinensis Schött, H, 1917:41

Lepidocyrtus australicus var. tamburinensis from Australia
After Schött H, 1917 Taf.3 Fig.2

             Syn.: Lepidosira tambourinensis(sic) Yoshii, R et Greenslade, P, 1994:18 ISS


Species bidentata Salmon, JT, 1938:358

Species bifasciata (Salmon, JT, 1944:157)

Species bisecta (Salmon, JT, 1944:162) Greenslade, 2012

Species brunnea (Womersley, H, 1935:217) Greenslade, P in Houston, WWK, 1994:98

Species calolepis (Börner, 1913) Yayuk, 1989:10

            Syn.: Lepidocyrtus variabilis Oudemans, 1890 =?
            Syn.: Lepidocyrtus setosus Folsom, 1924
            Syn.: Lepidosira montana Schött, 1925
            Syn.: Sira nilgiri Denis, 1936 =?


Species congoia Salmon, 1956

Species dorsalis (Salmon, JT, 1941:383) Greenslade, 2012

Urewera flava
subsp. dorsalis from New Zealand
After Salmon JT, 1941 Pl.64 Fig.401

Species faaroana (Carpenter, 1934) Greenslade, 2012

Species fallaciosa Yoshii, 1989

Species flava (Salmon, JT, 1938:353) Greenslade, 2012

Urewera flava from New Zealand
After Salmon JT, 1938 Pl.37 Fig.38
Urewera flava from New Zealand
After Salmon JT, 1941 Pl.64 Fig.400

Species fuchsiata (Salmon, JT, 1938:350) Greenslade, 2012, t.t.

Urewera fuchsiata from New Zealand
After Salmon JT, 1938 Pl.35 Fig.1
Urewera fuchsiata from New Zealand
After Salmon JT, 1941 Pl.63 Fig.390

Species fuscata Womersley, 1930

Species gigantea (Börner, C, 1909:122) Yosii, R, 1956:73




Species glebosa Salmon, JT, 1941:394

Species gupta Howard, GW, 1969:189

Lepidosira gupta from Australia
After Howard GW, 1969 Fig.1

Species ianthina (Salmon, JT, 1941:378) Greenslade, 2012

Species inconstans (Salmon, JT, 1938:353) Greenslade, 2012

Urewera inconstans from New Zealand
After Salmon JT, 1941 Pl.64 Fig.394

Species indistincta Salmon, JT, 1938:359

Species javana (Börner, 1913) Yayuk, RS (sic), 1989:10

Species laboriosa Greenslade, P in Houston, WWK, 1994:98

            Syn.: Lepidocyrtus (Lepidocyrtoides) cinctus Schött, H, 1917:45 nec Schäffer, 1898

Species lichenata (Salmon, JT, 1938:352) Greenslade, 2012

Species longicornis (Schött, H, 1917:38) Yoshii, R & Greenslade, P, 1994:14

Lepidocyrtoides longicornis from Australia
After Schött H, 1917 Taf.3 Fig.6

Species magna (Salmon, JT, 1937:355) Greenslade, 2012

Pseudosinella magna from New Zealand
After Salmon JT, 1937 Pl.50 Fig.e
Urewera tridentifera from New Zealand
After Salmon JT, 1938 Pl.35 Fig.8
Urewera magna from New Zealand
After Salmon JT, 1941 Pl.64 Fig.397

Species minima Salmon, JT, 1938:356

Lepidosira minima from New Zealand
After Salmon JT, 1938 Pl.36 Fig.15
Lepidosira minima from New Zealand
After Salmon JT, 1941 Pl.65 Fig.416

Species minuta Salmon, JT, 1938:356

Lepidosira minuta from New Zealand
After Salmon JT, 1938 Pl.36 Fig.22

Species neotropicalis Nunes RC et Bellini BC in Nunes, RC, Godeiro, NN, Pacheco, G, Liu, S, Gilbert, MTP, Alvarez-Valin, F, Zhang, F, et Bellini, BC, 2019:1

Lepidosira neotropicalis from Brazil
After Nunes RC & Bellini BC in Nunes RC & al, 2019 Fig.S1

Species nigrocephala (Womersley, H, 1936:480) Yosii, 1960

Mesira nigrocephala from Australia
After Womersley H, 1936 Fig.4a

Species nigropunctata (Nguyen, TT, 2005:8) Zhang, F, Chen, J-X & Deharveng, L, 2011:5

Species nilgiri (Denis, JR, 1936:266) Yosii, 1955

Sira(sic) nilgiri from India
After Denis JR, 1936 Fig.17
             Syn.: Lepidosira nilghiri(sic) Yosii, 1955 ISS


Species obscura (Salmon, JT, 1944:160) Greenslade, 2012

Species okarita Salmon, JT, 1938:360

Species omniofusca Salmon, JT, 1941:391

Species pallida (Ritter, 1911) Salmon, JT, 1964:507

Species parva (Salmon, JT, 1941:379) Greenslade, 2012

Species pigmenta Salmon, JT, 1944:164

Species punctata Yosii, 1960

Species purpurea (Salmon, JT, 1938:354) Greenslade, 2012

Species quadradentata (Salmon, JT, 1941:382) Greenslade, 2012

Urewera quadradentata from New Zealand
After Salmon JT, 1941 Pl.63 Fig.387

Species reducta (Salmon, JT, 1938:355) Greenslade, 2012

Species rotorua Salmon, JT, 1938:357

Lepidosira rotorua from New Zealand
After Salmon, J.T., 1938 Pl.36 Fig.24.

Species sagmaria (Schött, H, 1917:43) Schött, 1925

Lepidocyrtoides sagmarius from Australia
After Schött H, 1917 Fig.45
             Syn.: Lepidosira (Lepidocyrtoides) sagmarius(sic) Womersley, 1930 ISS
             Syn.: Lepidosira sagmarius(sic) Womersley, H, 1936:328 ISS
             Syn.: Lepidosira sagmarius(sic) Salmon, JT, 1941:390 ISS
Lepidosira sagmarius(sic) from New Zealand
After Salmon JT, 1941 Pl.66 Fig.430



Species sexmacula Salmon, JT, 1938:358

Lepidosira sexmacula from New Zealand
After Salmon JT, 1938 Pl.37 Fig.29
Lepidosira sexmacula from New Zealand
After Salmon JT, 1941 Pl.65 Fig.412
             Syn.: Lepidosira sexmaculata(sic) Nunes, RC, Godeiro, NN, Pacheco, G, Liu, S, Gilbert, MTP, Alvarez-Valin, F, Zhang, F, et Bellini, BC, 2019:6 ISS


Species splendida (Salmon, JT, 1941:380) Greenslade, 2012

Urewera splendida from New Zealand
After Salmon JT, 1941 Pl.63 Fig.382

Species sundana Yoshii, R et Suhardjono, YR, 1989:38

Species terraereginae (Ellis, WN & Bellinger, PF, 1973:28) Howard, GW, 1969:189, t.t.

            Syn.: Lepidocyrtus (Lepidocyrtoides) coeruleus Schött, H, 1917:45 nec Ritter, 1911
             Syn.: Lepidosira coerulus(sic) Howard, GW, 1969:189 ISS


Species unguserrata Salmon, 1970

Species variabilis (Oudemans, 1890)

Species variabilis (Schäffer, 1897) Christiansen & Bellinger, 2000:42, nec Oudemans, 1890

  Varietas albella Börner, 1906

  Varietas albiceps Börner, 1906

  Varietas bimaculata Börner, 1906

Species vicina Yoshii, 1989

Species violacea (Salmon, JT, 1938:352) Greenslade, 2012

Species violaceapallipes (Denis, 1931) Christiansen, K & Bellinger, P, 2000:42

            Syn.: Lepidocyrtinus domesticus f. violacea-pallipes(sic) Denis, 1931 IOS

Species womersleyi Greenslade P in Houston, 1994:99


Genus Epimetrura Schött, 1925

Species caudata (Carpenter, 1917) Denis, 1948

Lepidocyrtus caudatus from India
After Mandal GP & Hazra AK, 2009 Fig.31a

Species mirabilis Schött, 1925, t.t.

Species rostrata Greenslade, P et Sutrisno, 1994:53

Species tonkinensis Denis, 1948


Genus Lepidocyrtoides Schött, 1917 key  key

  Syn.: Entomobryinae Promesira Womersley, H, 1942:29
  Syn.: Lepidosira Najtsira Yoshii, 1989


  Syn.: Entomobryidae Sepidocyrtoides(sic) Zhang, F, Bedos, A et Deharveng, L, 2016:19 ISS

Species bicolor Yoshii, R et Suhardjno, YR, 1992:17bis

Lepidocyrtoides bicolor from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.10A

Species bicolorangelus Cipola NG et Bellini BC in Cipola, NG, de Morais, JW et Bellini, BC, 2017:220

Lepidocyrtoides bicolorangelus from Brazil
After Cipola NG, de Morais JW & Bellini BC, 2017 Fig.1E

Species caeruleomaculatus Cipola NG et Bellini BC in Cipola, NG, de Morais, JW et Bellini, BC, 2017:202

Lepidocyrtoides caeruleomaculatus from Brazil
After Cipola NG, de Morais JW & Bellini BC, 2017 Fig.1A

Species cheesmani Womersley, 1937

Species colormutatus Cipola NG et Bellini BC in Cipola, NG, de Morais, JW et Bellini, BC, 2017:214

Lepidocyrtoides colormutatus from Brazil
After Cipola NG, de Morais JW & Bellini BC, 2017 Fig.1B
Lepidocyrtoides colormutatus subadult from Brazil
After Cipola NG, de Morais JW & Bellini BC, 2017 Fig.1C
Lepidocyrtoides colormutatus juv. from Brazil
After Cipola NG, de Morais JW & Bellini BC, 2017 Fig.1D

Species cucullaris (Schött, H, 1917:39) Womersley, H, 1939:194, t.t.

Lepidocyrtoides cucullaris from Australia
After Schött H, 1917 Taf.3 Fig.3
             Syn.: Lepidocyrtoides cucularis(sic) Womersley, H, 1939:194 ISS


Species fasciatus (Womersley, 1934) Greenslade, P in Houston, WWK, 1994:94

Mesira fasciata from Australia
After Womersley H, 1939 Fig.62J

Species flavocinctus (Schött, H, 1917:42) Schött, 1925

Lepidocyrtoides flavocinctus from Australia
After Schött H, 1917 Fig.44

  Subspecies unicolor Womersley, 1934

Species hopkini Cipola NG et Bellini BC in Cipola, NG, de Morais, JW et Bellini, BC, 2017:238

Lepidocyrtoides hopkini from Australia
After Cipola NG, de Morais JW & Bellini BC, 2017 Fig.1I
Lepidocyrtoides hopkini juv. from Australia
After Cipola NG, de Morais JW & Bellini BC, 2017 Fig.1J

Species malabaricus Mandal, GP, Suman, KK et Bhattacharya, KK, 2019:18

Lepidocyrtoides malabaricus from India
After Mandal G.P. & al, 2019 Plate I Fig.1

Species matahari Yoshii, R et Suhardjno, YR, 1992:21

Lepidocyrtoides matahari from Indonesia
After Yoshii R & Suhardjono YR, 1992 Fig.13A

Species meraukeae Yoshii, R et Suhardjno, YR, 1992:19

Lepidocyrtoides meraukeae from Indonesia
After Yoshii R & Suhardjono YR, 1992 Fig.12A

Species najtae (Yoshii, 1989) Yoshii, R & Greenslade, P, 1994:11

Species novacaledoniae Yosii, 1960

             Syn.: Lepidocyrtoidea(sic) novaecaledoniae(sic) Deharveng, L et Najt, J in Tillier, S, 1988:18 ISS

Species oliveri Liu, L, Chen, J-X et Greenslade, P, 2008:56

Lepidocyrtoides oliveri from Australia
After Liu L, Chen J-X & Greenslade P, 2008 Fig.1

Species pictus (Schäffer, C, 1898:415) Yoshii, R & Suhardjno, YR, 1992:18

             Syn.: Sira(sic) (Lepidocyrtus) pictus(sic) Börner, C, 1901:11 ISS
Lepidocyrtoides pictus from Indonesia
After Yoshii R & Suhardjono YR, 1992 Fig.11A



Species quatuordecimocellata Prabhoo, 1971

Species striatus (Schött, 1901) Yoshii, R & Greenslade, P, 1994:11

Species tapuia (Arlé & Guimaraes, 1980) Cipola, NG, de Morais, JW & Bellini, BC, 2017:226

Lepidocyrtoides tapuia from Brazil
After Cipola NG, de Morais JW & Bellini BC, 2017 Fig.1F
Lepidocyrtoides tapuia juv. from Brazil
After Cipola NG, de Morais JW & Bellini BC, 2017 Fig.1G

Species timorensis Yoshii, R et Suhardjono, YR, 1992:80

Lepidocyrtoides timorensis from Indonesia
After Yoshii R & Suhardjono YR, 1992 Fig.3A

Species villasboasi (Arlé & Guimaraes, 1981) Cipola, NG, de Morais, JW & Bellini, BC, 2017:232

Lepidocyrtoides villasboasi from Brazil
After Cipola NG, de Morais JW & Bellini BC, 2017 Fig.1H

Species yasumatsui Uchida, 1944

             Syn.: Lepidocyrtoides yatsumatsui(sic) Liu, L, Chen, J-X et Greenslade, P, 2008:56 ISS
             Syn.: Lepidocyrtoides yatsumatsui(sic) Liu, L, Chen, J-X et Greenslade, P, 2008:60 ISS

References