http://www.collembola.org/taxa/arrhidae.htm - Last updated on 2024.02.23 by Frans Janssens
Checklist of the Collembola: Arrhopalitidae

Familia Arrhopalitidae Stach, 1956, sensu Bretfeld G, 1999:13 key

Genus Number
of species
Arrhopalites55
Pygmarrhopalites104
Troglopalites1
Arrhopalitidae160
Genera with number of species
  Syn.: ... Smynthuridae(sic) Lubbock, 1870, partim ISS
  Syn.: ... Smynthuridae(sic) Börner, 1906, partim ISS
  Syn.: Collembola Smynthuridae(sic) Lubbock, J, 1873:100 partim ISS
  Syn.: Collembola Sminthuri(sic) Folsom, JW, 1896:446 partim ISS
  Syn.: Katiannoidea Arrhopalitida(sic) Bretfeld, G, 1999:62
  Syn.: Collembola Collophoridae(lapsus) Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:29


  Syn.: Katianninae Arrhopalini(sic) Richards, 1956 ISS
  Syn.: Katianninae Arrhopalini(sic) Richards in 0918:31 ISS
  Syn.: Sminthurinae Arrhopalini(sic) Christiansen, K, 1966:48 ISS
Arrhopalites hirtus from the USA
2005 © Cheung, D. & Schmidt, J.
Arrhopalites sp. from the USA
2006 © Bernard, E.

  Syn.: Symphypleona Arrhopalitdae(sic) Shayanmehr, M, Yahyapour E, Kahrarian, M et Lafooraki EY, 2013:71 ISS



Genus Arrhopalites Börner, C, 1906:182 key  key  key  key  key  key  key  key


  Syn.: Smynthurus(sic) niveus Lebedinsky, 1904
  Syn.: Sminthuridae Pseudarrhopalites Stach, 1945
  Syn.: Arrhopalites Coecarrhopalites(sic) Yosii, 1954,nn IOS
  Syn.: Arrhopalites Coecarrhopalites(sic) Yosii, 1967 IOS
  Syn.: Arrhopalites Coecarrhopalites(sic) Massoud, Z et Betsch, J-M, 1972:58 IOS
  Syn.: Arrhopalites binoculatus Yosii, 1955
  Syn.: Sphyrotheca binoculatus Scott, 1964 partim holotype
  Syn.: Sphyrotheca binoculatus Scott, 1964 partim paratypes
  Syn.: Arrhopalitini Disparrhopalites Richards, WR, 1968:21
  Syn.: Sminthurinus reticulatus Falahati HosseinAbad, A, Kheirodin, A et Bretfeld, G, 2013:208
Sminthurinus reticulatus from Iran
After Falahati Hossein Abad A & al, 2013 Fig.3




  Syn.: Arrhopalites coecus(sic) Stach, J, 1945:7,gr ISS
  Syn.: Arrhopalites coecus(sic) Cassagnau, P et Delamare Deboutteville, C, 1953:135,gr ISS
  Syn.: Arrhopalites coecus(sic) Zeppelini, D, 2006:22,gr ISS
  Syn.: Arrhopalites coecus(sic) Zeppelini, D, 2011:91,gr ISS
  Syn.: Arrhopalitidae Arrhropalites(sic) Palacios-Vargas, JG, 1999:102 ISS
  Syn.: Arrhopalitidae Arropalites(sic) Zeppelini, DF et Palacios-Vargas JG, 1999:80 ISS

Arrhopalites sp. from France
2007.10.07 © Lebeaux, P.
Arrhopalites sp. from Hungary
2010.03.15 © Pfliegler, W.
Arrhopalites sp. from Hungary
2010.03.15 © Pfliegler, W.
Arrhopalites sp. from Croatia
2010.11.19 © Lukic, M.


  Syn.: Arrhopalitidae Arrhoplaites(sic) Vargovitsh, RS et Busmachiu, G, 2015:539 ISS
  Syn.: Collembola Arrthopalites(sic) Gagnon Koudji, E, Despland, E, Caron, A-S et Handa, IT, 2023:5 sp ISS




Species sp.1 Stach, J, 1956:239

             Syn.: Smynthurus(sic) gracilis Absolon, K, 1901:209 ISS

Species sp.3 Stach, J, 1956:240

            Syn.: Smynthurus(sic) niveus Joseph, 1882

Species sp.4 Christiansen, K, 1966:49

Species sp.5 Christiansen, K, 1966:49

Species sp.6 Christiansen, K, 1966:49

Species sp.7 Zeppelini Filho, D et Castaño Meneses, G, 1995:10

Species sp.8 Zeppelini Filho, D, 1996:27

Species sp.9 Culik, de Souza & Ventura, 2002

Species sp.10 Zeppelini, D, 2011:95

Species sp.11 Zeppelini, D, 2011:95

Species sp.12 Janssens F & Dobbs B, 2016

Arrhopalites? sp.nov. from the USA
Ant.3 with basal swelling; ant.4 with 10 subdivisions
2016.12.20 © Dobbs, Br.
Arrhopalites? sp.nov. from the USA
Dorsal colouration
2016.12.20 © Dobbs, Br.

Species pilulifer (Schött, H, 1893:9) Zeppelini, D, 2006:35

            Syn.: Sminthurus piluliferus(sic) Schött, H, 1893:9 IOS


             Syn.: Sminthurus piluliferus(sic) Salmon, JT, 1964:597 ISS
             Syn.: Arrhopalites piluliferus(sic) Zeppelini, D, 2006:35 ISS



Species violaceus (Handschin, 1930) Stach, J, 1956:72, s.i.


Species-group caecus Vargovitsh RS, 2013:27
Species abchasicus Vargovitsh, RS, 2013:17

Arrhopalites abchasicus from Abkhazia
After Vargovitsh RS, 2013 Fig.23
Arrhopalites abchasicus from Abkhazia
After Vargovitsh RS, 2013 Fig.24
            Syn.: Arrhopalites caecus Barjadze & Djanashvili, 2008


Species antrobius Yosii, R, 1954:68

Species brevicornis Godeiro, NN, Zhang, F et Bellini, CB, 2022:165

Arrhopalites brevicornis from China
After Godeiro NN, Zhang F & Bellini BC, 2022 Fig.1

Species caecus (Tullberg, 1871:146 tt) Börner, C, 1906:182, t.t., B

            Syn.: Sminthurus cæcus(sic) Tullberg, 1871 tt IOS
            Syn.: Smynthurus(sic) infernalis Joseph, 1892 =?
            Syn.: Smynthurus(sic) infernalis Hamann, 1896 =?
            Syn.: Sminthurus carniolicus von Dalla Torre, KW, 1895:8
             Syn.: Sminthurus coecus(sic) Stach, 1956, nec Tullberg, 1871 ISS
            Syn.: Smynthurus(sic) benitus Folsom, 1896
             Syn.: Smynthurus(sic) banitus(sic) Zeppelini D, 2006:26 ISS
            Syn.: Sminthurinus aureus Tuxen, 1944


             Syn.: Sminthurus (Pilosi) cæcus(sic) Tullberg, T, 1872:33 ISS
             Syn.: Sminthurus coecus(sic) Reuter, OM, 1876:80 ISS
             Syn.: Sminthurus coecus(sic) Reuter, 1890 ISS
             Syn.: Sminthurus coecus(sic) Reuter, OM, 1892:17 ISS
             Syn.: Sminthurus coecus(sic) Reuter, OM, 1892:18 ISS
             Syn.: Sminthurus coecus(sic) Schäffer, 1896 ISS
             Syn.: Sminthurus coecus(sic) Scherbakov, 1898 ISS
             Syn.: Sminthurinus coecus(sic) Börner, 1901 ISS
             Syn.: Arrhopalites coecus(sic) Linnaniemi, WM, 1912:271 ISS
             Syn.: Arrhopalites coecus(sic) Houlbert, 1924 ISS
             Syn.: Arrhopalites coecus(sic) Brown, 1926 ISS
             Syn.: Arrhopalites coecus(sic) Handschin, 1926 ISS
             Syn.: Arrhopalites coecus(sic) Edwards, 1929 ISS
             Syn.: Arrhopalites coecus(sic) Handschin, 1929 ISS
             Syn.: Arrhopalites coecus(sic) Denis, 1932 ISS
             Syn.: Arrhopalites coecus(sic) Womersley, 1932 ISS
             Syn.: Arrhopalites coecus(sic) Denis, J-R, 1933:285 ISS
             Syn.: Arrhopalites coecus(sic) Denis, J-R, 1933:287 ISS
             Syn.: Arrhopalites coecus(sic) Denis, J-R, 1933:295 ISS
             Syn.: Arrhopalites coecus(sic) Denis, J-R, 1933:297 ISS
             Syn.: Arrhopalites coecus(sic) Turk, 1933 ISS
             Syn.: Arrhopalites coecus(sic) Davies, 1934 ISS
             Syn.: Arrhopalites coecus(sic) Dürkop, 1934 ISS
             Syn.: Arrhopalites coecus(sic) Stammer, 1936 ISS
             Syn.: Arrhopalites coecus(sic) Frenzel, 1937 ISS
             Syn.: Arrhopalites coecus(sic) Weidner, 1937 ISS
             Syn.: Arrhopalites coecus(sic) Denis, 1938 ISS
             Syn.: Arrhopalites coecus(sic) Muhlmann, 1942 ISS
             Syn.: Arrhopalites coecus(sic) Kloet, GS et Hincks, WD, 1945:9 ISS
             Syn.: Arrhopalites coecus(sic) Stach, J, 1945:9 ISS
            Syn.: Arrhopalites cæcus(sic) Denis, J-R, 1947:6 IOS
            Syn.: Arrhopalites cæcus(sic) Denis, J-R, 1947:7 IOS
            Syn.: Arrhopalites Cæcus(sic) Denis, J-R, 1947:7 IOS
            Syn.: Arrhopalites cæcus(sic) Denis, J-R, 1947:8 IOS
             Syn.: Arrhopalites coecus(sic) Cassagnau, P et Delamare Deboutteville, C, 1953:133 ISS
             Syn.: Arrhopalites coecus(sic) Weidner, 1953 ISS
             Syn.: Arrhopalites coecus(sic) Bellinger, 1954 ISS
             Syn.: Arrhopalites coecus(sic) Obenberger, 1955 ISS
             Syn.: Arrhopalites coecus(sic) Stach, 1955 ISS
             Syn.: Arrhopalites coecus(sic) Strenzke, 1955 ISS
             Syn.: Arrhopalites coecus(sic) Paclt, 1956 ISS
             Syn.: Arrhopalites coecus(sic) Bödvarsson, H, 1957:38 ISS
             Syn.: Arrhopalites coecus(sic) Strebel, 1957 ISS
             Syn.: Arrhopalites coecus(sic) Grinbergs, 1958 ISS
             Syn.: Arrhopalites coecus(sic) von Törne, 1958 ISS
             Syn.: Arrhopalites coecus(sic) da Gama, 1959 ISS
             Syn.: Arrhopalites coecus(sic) Paclt, 1959 ISS
             Syn.: Arrhopalites coecus(sic) Grinbergs, 1960 ISS
             Syn.: Arrhopalites coecus(sic) Paclt, 1961 ISS
             Syn.: Arrhopalites coecus(sic) Cassagnau, P, 1962:229 ISS
             Syn.: Arrhopalites coecus(sic) Paclt, 1962 ISS
             Syn.: Coecarrhopalites(sic) coecus(sic) Delamare Deboutteville & Massoud, 1963 ISS
             Syn.: Arrhopalites coecus(sic) Pomorski & Skarzynski, 1991 ISS
             Syn.: Arrhopalites coecus(sic) Palacios-Vargas & Zeppelini, [1996] ISS
             Syn.: Arrhopalites coecus(sic) Zeppelini, 1996 ISS
             Syn.: Arrhopalites (Arrhopalites) coecus(sic) Zeppelini Filho, D, 1996:57 ISS
             Syn.: Arrhopalites coecus(sic) Zeppelini, [1997] ISS
             Syn.: Arrhopalites coecus(sic) Zeppelini Filho, D, 1997:122 ISS
             Syn.: Arrhopalites coecus(sic) Klag, J et Swiatek, P, 1999:161 ISS
             Syn.: Arrhopalites coecus(sic) Beruete, E, Baquero, E et Jordana, R, 2002:397 ISS
             Syn.: Arrhopalites coecus(sic) Zeppelini, D et Christiansen, K, 2003:37 ISS
Arrhopalites caecus from the UK
Mucro
2005 © Hopkin, S.P.

             Syn.: Arrhopalites ceacus(sic) Moore, JC, Saunders, P, Selby, G, Horton, H, Chelius, MK, Chapman, A et Horrocks, RD, 2005:116 ISS
Arrhopalites caecus from Hungary
2011.02.11 © Pfliegler, W.P.
Arrhopalites caecus from Hungary
2011.02.11 © Pfliegler, W.P.
Arrhopalites caecus ♀ from Hungary
Sub-anal appendage
2011.02.11 © Pfliegler, W.P.
Arrhopalites caecus from Hungary
Pale colour form
2011.02.11 © Pfliegler, W.P.
Arrhopalites caecus from Hungary
Mandibula with apical teeth and molar plate
2013.01.11 © Pfliegler, W.P.
Arrhopalites caecus ♀ from Hungary
Sub-anal appendage
2013.01.11 © Pfliegler, W.P.
Arrhopalites caecus from Hungary
Mucro with round tip (*), serrated edges and basal ventral spine (**)
2013.01.11 © Pfliegler, W.P.
Arrhopalites caecus from Hungary
Ant.4 subdivisions indistinct
2013.01.11 © Pfliegler, W.P.
Arrhopalites caecus from Hungary
Bothriotricha ABC sub-orthogonal
2013.01.11 © Pfliegler, W.P.
Arrhopalites caecus from the USA
2022.11.29 © Wang, G.
Arrhopalites caecus from the USA
Mucro
2022.11.29 © Wang, G.
Arrhopalites caecus from the USA
Hind foot
Unguiculus broad
2022.11.29 © Wang, G.
Arrhopalites caecus from the USA
2022.12.29 © Wang, G.
Arrhopalites caecus from the USA
Mucro 'bipartite'
2022.12.29 © Wang, G.
Arrhopalites caecus from the USA
Ant.4 with less then 5 subsegments
2022.12.29 © Wang, G.
Arrhopalites caecus from the USA
Unguis with inner tooth
Unguiculus shorter then unguis
2022.12.29 © Wang, G.
Arrhopalites caecus from the USA
Infected by ?Sporodiniella umbellata?
2023.01.21 © Wang, G.
Arrhopalites caecus ♀ from the USA
Curved spine at base of mucro
2023.02.19 © Pavy, R.
Arrhopalites caecus ♀ from the USA
Anal valves with short thick spines inbetween long setae
2023.02.19 © Pavy, R.
Arrhopalites caecus ♀ from the USA
Sub-anal appendage
2023.02.19 © Pavy, R.

  Subspecies attenuatus Carpenter & Evans, 1899





Species coreanus Park & Kang, 2007

Species gul Yosii, R, 1966:559

Remark by Janssens, F. 2023.10.27: "Gul" is a vernacular Korean word that means "cave". The ICZN Rec.11A discourages the use of vernacular words as scientific names. Given appropriate latinisation is the prefered means of formation of names from vernacular words, "cavus" (= pertaining to the cave) would be an example of an appropriate name in line with Rec.11A.
Species incertus Zeppelini, D et Christiansen, K, 2003:36

Species karabiensis Vargovitsh, RS, 2009:10

Arrhopalites karabiensis from Ukraine
After Vargovitsh, R.S., 2009 Fig.2

Species loczyi Loksa, 1960

Species macronyx Vargovitsh, RS, 2012:2

Arrhopalites macronyx from Western Caucasus
After Vargotitsh RS, 2012 Fig.25
Arrhopalites macronyx from Europe
After Vargovitsh R in Fiera C & al, 2021 Fig.1c

Species millsi (Delamare Deboutteville, C & Massoud, Z, 1963:215) Betsch, 1980

            Syn.: Caecarrhopalites Millsi(sic) Delamare Deboutteville, C et Massoud, Z, 1963:215 IOS

Species minor Park & Kang, 2007

Species minutus Yosii, R, 1970:17

Species peculiaris Vargovitsh, RS, 2009:15

Species potapovi Vargovitsh, RS, 2015:102

Arrhopalites potapovi ♀ from Russia
After Vargovitsh RS, 2015 Fig.24

Species profundus Vargovitsh, RS, 2022:6

Arrhopalites profundus ♂ from the Caucasus
After Vargovitsh RS, 2022 Fig.5A

Species pukouensis Wu, M et Christiansen, K, 1997:266

Species remyi Denis, J-R, 1947:5

            Syn.: Arrhopalites Remyi(sic) Denis, J-R, 1947:5 IOS
Arrhopalites Remyi(sic) from Corsica
After Denis J-R, 1947 Fig.3a

            Syn.: Arrhopalites Remyi(sic) Denis, J-R, 1947:6 IOS
            Syn.: Arrhopalites Remyi(sic) Denis, J-R, 1947:7 IOS
            Syn.: Arrhopalites Remyi(sic) Denis, J-R, 1947:8 IOS
            Syn.: Arrhopalites? remyi Stach, J, 1956:72,si
            Syn.: Arrhopalites remyi Bretfeld, G, 1999:65,si
            Syn.: Arrhopalites? remyi Vargovitsh, RS, 2013:27,si




Species tenuis Stach, J, 1945:14

            Syn.: Arrhopalites Remyi(sic) Cassagnau, P et Delamare Deboutteville, C, 1953:141 =? IOS




Species ulehlovae Rusek, 1970


Species-group diversus Vargovitsh RS, 2013:27
Species sp.1 Vargovitsh, RS et Busmachiu, G, 2015:548

Species acanthophthalmus Gisin, 1958

             Syn.: Arrhopalites acantophthalmus(sic) Dallai, R, 1973:560 ISS
             Syn.: Arrhopalites acantophthalmus(sic) Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:895 ISS
             Syn.: Arrhopalites acanthophthalma(sic) Russell, DJ et Griegel, A, 2006:174 ISS

Species antonioi Giuga & Jordana, 2013

Species baccettii Dallai, 1969

             Syn.: Arrhopalites (diversus-group) baccetii(sic) Vargovitsh, RS et Busmachiu, G, 2015:539 ISS

Species diversus Mills, HB, 1934:93

Arrhopalites diversus from the USA
Eyepatch with 1 ocellus
2018.01.17 © Dobbs, Br.
Arrhopalites diversus from the USA
Ant.3 with distinct basal swelling (*)
2018.01.17 © Dobbs, Br.
Arrhopalites diversus from the USA
Ant.1:2:3:4 = 1:2.2:3.75:6.25
Ant.4 not subdivided
2018.01.17 © Dobbs, Br.
Arrhopalites diversus from the USA
2024.01.18 © Coogler, J.
Arrhopalites diversus from Canada
Ant.3 with distinct basal bulge
Ant.4 not subdivided
2024.01.26 © Chan, J.
Arrhopalites diversus from Canada
Ant.3 with distinct basal bulge
2024.01.30 © Chan, J.
Arrhopalites diversus from Canada
Interocular region darker
2024.01.30 © Chan, J.
Arrhopalites diversus from Canada
Ant.4 not subdivided
2024.01.30 © Chan, J.
Arrhopalites diversus from Canada
2024.01.30 © Chan, J.
Arrhopalites diversus from Canada
Subcutaneous white fat tissue
2024.01.30 © Chan, J.

Species ezoensis Zeppelini, D, 2004:4

Species microphthalmus Cassagnau, P et Delamare Deboutteville, C, 1953:133, B

             Syn.: Arrhopalites microphtalmus(sic) Zeppelini Filho, D, 1996:27 ISS
             Syn.: Arrhopalites (Arrhopalites) microphtalmus(sic) Zeppelini Filho, D, 1996:58 ISS
             Syn.: Arrhopalites microphtalmus(sic) Zeppelini Filho, D, [1997]:12 ISS
             Syn.: Arrhopalites microphtalmus(sic) Jacobs, W, 1997:71,B ISS
             Syn.: Arrhopalites microphtalmus(sic) Jacobs, W, De Bruyn, L et Lust, N, 1999:58,B ISS
Arrhopalites ?microphthalmus? from France
2020.02.01 © Reinach, Br.
Arrhopalites ?microphthalmus? from France
2020.02.01 © Reinach, Br.



Species nivalis Yosii, 1966

Species persicus Vargovitsh, R et Kahrarian, M, 2020:339

Arrhopalites persicus from Iran
After Vargovitsh RS & Kahrarian M, 2020 Fig.21

Species prutensis Vargovitsh, RS et Busmachiu, G, 2015:540

Arrhopalites prutensis ♀ from Moldova
After Vargovitsh RS & Busmachiu G, 2015 Fig.2

Species-group harveyi Vargovitsh RS, 2013:27
Species alambariensis Zeppelini, D, 2006:19

Species amorimi Palacios-Vargas, JG et Zeppelini, D, 1996:10

Species anulifer Nayrolles, 1990

Species botuveraensis Zeppelini, D, 2006:22

Species glabrofasciatus Zeppelini, D, Brito, RA et Lima, ECA, 2018:60

Arrhopalites glabrofasciatus ♀ from Brazil
After Zeppelini D & al, 2018 Fig.1

Species gnaspinii Palacios-Vargas, JG et Zeppelini, D, 1996:8

            Syn.: Arrhopalites (Coecarrhopalites(sic)) gnaspinius(sic) Palacios-Vargas, JG et Zeppelini, D, 1996:8 IOS
             Syn.: Arrhopalites gnaspinius(sic) Mari Mutt, JA et Bellinger, PF, 1996:172 ISS
             Syn.: Arrhopalites gnaspinius(sic) Zeppelini Filho, D, 1996:29 ISS
             Syn.: Arrhopalites (Arrhopalites) gnaspinius(sic) Zeppelini Filho, D, 1996:58 ISS
             Syn.: Arrhopalites gnaspinius(sic) Zeppelini Filho, D, [1997]:12 ISS
             Syn.: Arrhopalites gnaspinius(sic) Zeppelini Filho, D, 1997:122 ISS
             Syn.: Arrhopalites gnaspinius(sic) Zeppelini, DF et Palacios-Vargas JG, 1999:81 ISS
             Syn.: Arrhopalites gnaspinius(sic) Culik, MP et Zeppelini, DF, 2003:1131 ISS
             Syn.: Arrhopalites (harveyi-group) gnaspinius(sic) Brito, RA, de Lima, ECA et Zeppelini, D, 2019:410 ISS



Species harveyi Denis, J-R, 1933:285

            Syn.: Arrhopalites Harveyi(sic) Denis, J-R, 1933:285 IOS
            Syn.: Arrhopalites Harveyi(sic) Denis, J-R, 1933:286 IOS
            Syn.: Arrhopalites Harveyi(sic) Denis, J-R, 1933:297 IOS
            Syn.: Arrhopalites Harveyi(sic) Denis, J-R, 1933:315 IOS
            Syn.: Arrhopalites Harveyi(sic) Denis, J-R, 1933:320 IOS
            Syn.: Coecarrhopalites(sic) Harveyi(sic) Delamare Deboutteville & Massoud, 1963 IOS

Species heteroculatus Zeppelini, D, 2006:20

Species lawrencei Palacios-Vargas, JG et Zeppelini, D, 1996:12

             Syn.: Arrhopalites (Coecarrhopalites(sic)) lawrenci(sic) Palacios-Vargas, JG et Zeppelini, D, 1996:23 ISS
             Syn.: Arrhopalites (harveyi-group) lawrenci(sic) Brito, RA, de Lima, ECA et Zeppelini, D, 2019:410 ISS

Species mendoncae Brito, RA, de Lima, ECA et Zeppelini, D, 2019:402

Species paranaensis Zeppelini, D, 2006:24

Species vazquezae Palacios-Vargas, JG et Zeppelini, D, 1996:14


Genus Pygmarrhopalites Vargovitsh, RS, 2009:20

  Syn.: Sminthurinus Articulati Börner, C, 1901:100
  Syn.: Collembola Smynthrus(sic) Christiansen, K, 1966:53 ISS
  Syn.: Arrhopalites Novarrhopalites Zeppelini Filho, D, 1996:57 =u
  Syn.: Arrhopalites pygmaeus-group Stach, J, 1945:7
  Syn.: Katianninae Arrhopalites Yosii, R, 1977:164 partim
  Syn.: Arrhopalitidae Arrhopalites Marx, MT et Weber, D, 2013:234
  Syn.: Pygmarrhopalites? remyi Vargovitsh, RS, 2021:il,si


Arrhopalites sp. from Chile
After Jordana R & Baquero E, 2008 Fig.5
Remark by Zeppelini, D. 2011:96: The name Pygmarrhopalites has only a didactic meaning, being a convenience for referring to species in the pygmaeus-group, which follows the system proposed by Bretfeld (1999). Thus the generic name Pygmarrhopalites should be treated as a junior synonym of Arrhopalites s.l. Börner.

  Syn.: Collembola Pygmaarhopalites(sic) Saifutdinov, RA et Korobushkin, DI, 2020:307 ISS


Species sp.1 Zeppelini Filho, D, 1996:27

Species sp.2 Vargovitsh, RS et Busmachiu, G, 2015:540

Species sp.3 Vargovitsh, RS et Busmachiu, G, 2015:540

Species sp.4 Janssens F sensu Arbea & al, 2021

            Syn.: Arrhopalites terricola(sic) Arbea & Pérez Fernández, 2021
            Syn.: Arrhopalites terricola(sic) Arbea, 2013
            Syn.: Pygmarrhopalites terricola(sic) Arbea, JI, Baquero, E, Beruete, E, Pérez Fernández, T et Jordana, R, 2021:62 IOS
            Syn.: Pygmarrhopalites sp.inédita Arbea, JI, Baquero, E, Beruete, E, Pérez Fernández, T et Jordana, R, 2021:62




Species aggtelekiensis (Stach, J, 1930:308) Vargovitsh, RS, 2009:26

             Syn.: Arrhopalites aggtelekensis(sic) Gisin, 1960 ISS
             Syn.: Arrhopalites aggtelekensis(sic) Christiansen, 1966 ISS
             Syn.: Arrhopalites aggtelekensis(sic) Loksa, I, 1967:294 ISS
             Syn.: Arrhopalites aggtelkiensis(sic) Nosek, J et Paoletti, MG, 1981:70 ISS
             Syn.: Arrhopalites aggtelekensis(sic) Zeppelini Filho, D, 1996:27 ISS
             Syn.: Arrhopalites aggtelekensis(sic) Hubart, J-M et Dethier, M, 1999:168 ISS
Pygmarrhopalites aggtelekiensis from Slovakia
2012 © Kovác, L.



Species alticolus (Yosii, R, 1970:15) Vargovitsh, RS, 2009:36

             Syn.: Arrhopalites alticola(sic) Zeppelini, D, 2004:3 ISS
             Syn.: Pygmarrhopalites alticola(sic) Vargovitsh, RS, 2009:36 ISS
             Syn.: Pygmarrhopalites alticola(sic) Vargovitsh, RS, 2009:e18 ISS
             Syn.: Pygmarrhopalites alticola(sic) Jordana, R, Fadrique, F et Baquero, E, 2012:63 ISS

Species altus (Christiansen, K, 1966:51) Vargovitsh, RS, 2009:e18

Pygmarrhopalites altus from the USA
2023.01.01 © Wang, G.
Pygmarrhopalites altus from the USA
2023.01.01 © Wang, G.

Species amarus (Christiansen, K, 1966:62) Vargovitsh, RS, 2009:20

Species arcus (Zeppelini, D & Christiansen, K, 2003:38) Vargovitsh, RS, 2009:e18

            Syn.: Arrhopalites arca(sic) Zeppelini, D et Christiansen, K, 2003:38 IOS
             Syn.: Pygmarrhopalites arca(sic) Zeppelini, D, Taylor, SJ et Slay, ME, 2009:1 ISS
             Syn.: Pygmarrhopalites (principalis-group) arca(sic) Vargovitsh, RS, 2017:33 ISS



Species ashcraftensis Zeppelini, D, Taylor, SJ et Slay, ME, 2009:13

Species ater (Christiansen, K & Bellinger, P, 1998:1247) Vargovitsh, RS, 2009:e18

Pygmarrhopalites ater ♀ from the USA
2023.11.19 © Wang, G.
Pygmarrhopalites ater ♀ from the USA
Sub-anal appendage curved and acuminate
2023.11.19 © Wang, G.
Pygmarrhopalites ater ♀ from the USA
Ant.3 w/o basal bulge
2023.11.19 © Wang, G.
Pygmarrhopalites ater ♀ from the USA
Ant.4 with 5 subdivisions
2023.11.19 © Wang, G.
Pygmarrhopalites ?ater? from the USA
2023.11.24 © Wang, G.
Pygmarrhopalites ?ater? from the USA
2023.11.24 © Wang, G.

Species bellingeri (Christiansen, K, 1966:51) Vargovitsh, RS, 2009:e18

Pygmarrhopalites bellingeri from Canada
2024.01.26 © Chan, J.
Pygmarrhopalites bellingeri from Canada
Ant.4 with about 7 subdivisions
2024.01.26 © Chan, J.
Pygmarrhopalites bellingeri from Canada
2024.01.26 © Chan, J.
Pygmarrhopalites bellingeri from Canada
Eye with 2 osculating ocelli
2024.01.26 © Chan, J.
Pygmarrhopalites bellingeri from Canada
Thoracic intersegmental membranes pale
Body with numerous pale spots
2024.01.26 © Chan, J.
Pygmarrhopalites bellingeri from Canada
2024.01.30 © Chan, J.
Pygmarrhopalites bellingeri from Canada
Dorsal pattern
2024.01.30 © Chan, J.
Pygmarrhopalites bellingeri from Canada
Caudal pattern
2024.01.30 © Chan, J.

Species benitus (Folsom, JW, 1896:446) Vargovitsh, RS, 2009:e18

Smynthurus(sic) benitus from the USA
Terminal antennal segment
After Folsom JW, 1896 Plate10 Fig.5

            Syn.: Sminthurus caecus Guthrie, 1903
            Syn.: Arrhopalites caecus Linnaniemi, 1907
            Syn.: Arrhopalites sub-binoculatus(sic) Denis, J-R, 1933:287 IOS
            Syn.: Arrhopalites sub-binoculatus(sic) Denis, J-R, 1933:288 IOS
            Syn.: Arrhopalites sub-binoculatus(sic) Denis, J-R, 1933:297 IOS
             Syn.: Arrhopalites sub-binoculatus(sic) Stach, J, 1956:70 ISS
            Syn.: Arrhopalites binoculatus Denis, 1931
            Syn.: Arrhopalites binoculatus Denis, J-R, 1933:320
            Syn.: Arrhopalites binoculatus Mills, 1934
             Syn.: ... bimaculatus(sic) Brimley, 1938 lapsus ISS
            Syn.: Arrhopalites binoculatus Maynard, 1951
            Syn.: Arrhopalites principalis Stach, 1945 =?




Species bifidus (Stach, J, 1945:34) Vargovitsh, RS, 2009:20, B

             Syn.: Arrhopalites bificus(sic) Salmon, JT, 1964:571 ISS


Pygmarrhopalites bifidus from the UK
2012.08.19 © Murray, A.
Pygmarrhopalites bifidus from the UK
2012.08.19 © Murray, A.
Pygmarrhopalites bifidus from the UK
2012.08.19 © Murray, A.
Pygmarrhopalites bifidus from the UK
Sub-anal appendage
2012.08.19 © Murray, A.
Pygmarrhopalites bifidus ♀ from the UK
2015.03.04 © Murray, A.
Pygmarrhopalites bifidus ♀ from the UK
Bifid sub-anal appendages (*)
2015.03.04 © Murray, A.
Pygmarrhopalites bifidus juv. from the UK
2015.03.04 © Murray, A.
Pygmarrhopalites bifidus from the UK
Reddish eyepatches
2015.03.04 © Murray, A.
Pygmarrhopalites bifidus from the UK
Ant.4 with 5 subdivisions
2015.03.04 © Murray, A.
Pygmarrhopalites bifidus from Holland
Ant.4 with 5 subdivisions
2017.11.22 © Wieringa, P.
Pygmarrhopalites bifidus from Holland
2017.11.22 © Wieringa, P.
Pygmarrhopalites bifidus from Holland
2017.11.22 © Van duinen, J.

Species bimus (Christiansen, K, 1966:55) Vargovitsh, RS, 2009:e18

Species boneti (Stach, J, 1945:16) Vargovitsh, RS, 2009:31

            Syn.: Arrhopalites pygmaeus Bonet, 1931 nec Wankel, 1860


             Syn.: Arrhopalites Boneti(sic) Denis, J-R, 1947:7 ISS
             Syn.: Arrhopalites Boneti(sic) Denis, J-R, 1947:8 ISS
             Syn.: Arrhopalites Boneti(sic) Cassagnau, P et Delamare Deboutteville, C, 1953:135 ISS

Species buekkensis (Loksa, I, 1969:359) Vargovitsh, RS, 2009:26

Species buffaloensis Zeppelini, D, Taylor, SJ et Slay, ME, 2009:10

Species caedus (Christiansen, K & Bellinger, P, 1996:170) Vargovitsh, RS, 2009:e18

Species cantavetulae Jordana, R, Fadrique, F et Baquero, E, 2012:58

Species canzianus (Stach, J, 1945:28) Vargovitsh, RS, 2009:20

Species carolynae (Christiansen, K & Bellinger, P, 1996:171) Vargovitsh, RS, 2009:e18

             Syn.: Pygmarrhopalites carolinae(sic) Soto-Adames, FN et Taylor, SJ, 2013:173 ISS

Species carpathicus (Vargovich, RS, 1999:89) Vargovitsh, RS, 2009:20

Pygmarrhopalites carpathicus from the Carpathians
2011.03.11 © Vargovich, R.

Species changbaishanensis (Wu, M & Zhong, W-Y, 1997:215) Vargovitsh, RS, 2009:20

Species chiangdaoensis (Nayrolles, 1990) Vargovitsh, RS, 2009:20

Species chopardi (Cassagnau, P & Delamare Deboutteville, C, 1955:388) Vargovitsh, RS, 2009:e18

            Syn.: Arrhopalites Chopardi(sic) Cassagnau, P et Delamare Deboutteville, C, 1955:388 IOS

Species clarus (Christiansen, K, 1966:57) Vargovitsh, RS, 2009:e18

Species cochlearifer (Gisin, 1947) Vargovitsh, RS, 2009:e18

             Syn.: Arrhopalites cochleafer(sic) Christiansen, 1966 ISS
             Syn.: Pygmarrhopalites cochlaerifer(sic) Zeppelini, D, 2011:95,gr ISS
Pygmarrhopalites cochlearifer from the UK
Dark colour form
2013.06.dd © Shaw, P.
Pygmarrhopalites cochlearifer from the UK
Ant.4 with 6 subsegments
2013.06.dd © Shaw, P.
Pygmarrhopalites cochlearifer from the UK
Distinctly strong anterior cephalic spines
2013.06.dd © Shaw, P.



Species commorus (Christiansen, K & Bellinger, P, 1996:172) Vargovitsh, RS, 2009:26

Species crepidinis Jordana R et Baquero E in Jordana, R, Barranco, P, Amezcua, A et Baquero, E, 2017:106

Pygmarrhopallites crepidinis from Spain
After Jordana R & Baquero E in Jordana R & al, 2017 Fig.2A

Species custodum Baquero E et Jordana R in Baquero, E, Jordana, R et Ortuño, VM, 2021:8

Species dbari Vargovitsh, RS, 2017:24

Pygmarrhopalites dbari from Abkhazia
After Vargovitsh RS, 2017 Fig.25
Pygmarrhopalites dbari from Europe
After Vargovitsh R in Fiera C & al, 2021 Fig.1d

Species delamarei (Nosek, J & Paoletti, MG, 1984:211) Vargovitsh, RS, 2009:e18

            Syn.: Arrhopalites delamarei Nosek, J et Paoletti, MG, 1981:70 nn

Species dubius (Christiansen, 1966:59) Vargovitsh, RS, 2009:e18

             Syn.: Arrhopalites dubias(sic) Miranda Rangel & Palacios-Vargas, 1992 ISS

Species dudichi (Loksa, I & Rubio, I, 1966:154) Vargovitsh, RS, 2009:20

Species elegans (Cassagnau, P & Delamare Deboutteville, C, 1953:138) Vargovitsh, RS, 2009:e18

Species fransjanssensi Soto-Adames, FN et Taylor, SJ, 2013:164

            Syn.: Pygmarrhopalites fransjanssens(sic) Soto-Adames, FN et Taylor, SJ, 2013:164 IOS
             Syn.: Pygmarrhopalites fransjanssnes(sic) Soto-Adames, FN et Taylor, SJ, 2013:173 ISS



Species furcatus (Stach, J, 1945:22) Vargovitsh, RS, 2009:20

Species giovannensis (Cassagnau, P & Delamare Deboutteville, C, 1953:136) Vargovitsh, RS, 2009:e18

Species gisini (Nosek, 1960) Vargovitsh, RS, 2009:e18

Species habei (Yosii, R, 1956:98) Vargovitsh, RS, 2009:20

            Syn.: Arrhopalites binoculatus Yosii, 1954
            Syn.: Arrhopalites binoculatus Yosii, 1955
            Syn.: Arrhopalites binoculatus Yosii, R, 1977:164
            Syn.: Arrhopalites spadica Salmon, 1963


             Syn.: Arrhopalites habet(sic) Yosii, R, 1956:99 ISS
             Syn.: Arrhopalites havei(sic) Christiansen, K, 1966:61 ISS
             Syn.: Arrhopalites Habei(sic) Zeppelini, D, 2004:13 ISS
Pymarrhopalites habei from Japan
2019.02.27 © Hara, Y.
Pymarrhopalites habei from Japan
2019.02.27 © Hara, Y.



Species hedrosensis (Selga, D, 1963:465) Vargovitsh, RS, 2009:e18

Species hirtus (Christiansen, K, 1966:61) Soto-Adames, FN & Taylor, SJ, 2013:148 lapsus

            Syn.: Pygmarrhopalites hirsuta(sic) Soto-Adames, FN et Taylor, SJ, 2013:148 lapsus


Arrhopalites hirtus from the USA
2005 © Cheung, D. & Schmidt, J.

Species hubbardi (Zeppelini, D & Christiansen, K, 2003:37) Vargovitsh, RS, 2009:e18

Arrhopalites hubbardi from the USA
2007.03.15 © Steinmann, D.

Species hungaricus (Loksa, I, 1967:294) Vargovitsh, RS, 2009:e18

Species incantator Soto-Adames, FN et Taylor, SJ, 2013:164

Species infrasecundarius (Loksa, I & Rubio, I, 1966:154) Vargovitsh, RS, 2009:36

Species intermedius (Loksa, I, 1969:357) Vargovitsh, RS, 2009:e18

Species japonicus (Yosii, R, 1956:99) Vargovitsh, RS, 2009:20

            Syn.: Arrhopalites binoculatus Yosii, 1954, nec Börner
            Syn.: Arrhopalites binoculatus Yosii, R, 1956:99


Species jay (Christiansen, K & Bellinger, P, 1996:173) Vargovitsh, RS, 2009:20

             Syn.: Arrhopalites jax(sic) Zeppelini Filho, D, 1996:27 ISS
             Syn.: Arrhopalites (Novarrhopalites) jax(sic) Zeppelini Filho, D, 1996:58 ISS
             Syn.: Arrhopalites jax(sic) Zeppelini Filho, D, [1997]:12 ISS

Species jeanneli (Cassagnau, P & Delamare Deboutteville, C, 1955:390) Vargovitsh, RS, 2009:e18

            Syn.: Arrhopalites Jeanneli(sic) Cassagnau, P et Delamare Deboutteville, C, 1955:390 IOS

Species kaprusi Vargovitsh, RS, 2009:26

Species kolymensis (Tshelnokov, VG, 1990:351) Vargovitsh, RS, 2009:36

             Syn.: Arrhopalites kolymiensis(sic) Tshelnokov, VG, 1990:350 ISS

Species kovali Vargovitsh, RS, 2017:33

Pygmarrhopalites kvali from Abkhazia
After Vargovitsh RS, 2017 Fig.57

Species kristiani (Vargovich, RS, 2005:67) Vargovitsh, RS, 2009:20

Arrhopalites kristiani from Ukraine
After Vargovich RS, 2005 Fig.1

Species lacuna (Christiansen, K & Bellinger, P, 1996:174) Vargovitsh, RS, 2009:e18

Species leonardwoodensis Zeppelini, D, Taylor, SJ et Slay, ME, 2009:3

Species lewisi (Christiansen, K & Bellinger, P, 1998:1249) Vargovitsh, RS, 2009:e18

Species longicornis (Cassagnau, P & Delamare Deboutteville, C, 1953:141) Vargovitsh, RS, 2009:20

Species madonnensis (Zeppelini, D & Christiansen, K, 2003:41) Vargovitsh, RS, 2009:e18

Species maestrazgoensis Jordana, R, Fadrique, F et Baquero, E, 2012:52

Species marshalli (Christiansen, K & Bellinger, P, 1996:175) Vargovitsh, RS, 2009:e18

Species mauli (Delamare Deboutteville, C & Bassot, J-M, 1957:78) Vargovitsh, RS, 2009:e18

             Syn.: Arrhopalites Mauli(sic) Delamare Deboutteville, C et Bassot, J-M, 1957:79 ISS
             Syn.: Arrhopalites Mauli(sic) Delamare Deboutteville, C et Bassot, J-M, 1957:86 ISS

Species miravetensis (Baquero, E, Herrando-Pérez, S & Jordana, R, 2005:82) Vargovitsh, RS, 2009:20

Species nanjingensis (Lin & Chen, 1997) Vargovitsh, RS, 2009:20

Species nigripes (Park & Kang, 2007) Vargovitsh, RS, 2009:20

Species obtusus (Zeppelini, D & Christiansen, K, 2003:40) Vargovitsh, RS, 2009:e18

Species octacanthus (Yosii, R, 1970:14) Vargovitsh, RS, 2009:20

Species ornatus (Stach, J, 1945:24) Vargovitsh, RS, 2009:20

Species pavo (Christiansen, K & Bellinger, P, 1996:176) Vargovitsh, RS, 2009:e18

Species perezi Arbea, JI, 2013:37

Pygmarrhopalites perezi from Spain
After Arbea JI, 2013 Fig.16

Species plectrifer (Hüther, 1963) Vargovitsh, RS, 2009:20

Species plethorasari Zeppelini, D, Taylor, SJ et Slay, ME, 2009:6

Species postumicoides (Cassagnau, P & Delamare Deboutteville, C, 1953:137) Vargovitsh, RS, 2009:e18

             Syn.: Arrhopalites postumicoïdes(sic) Cassagnau, P et Delamare Deboutteville, C, 1953:142 ISS

Species postumicus (Stach, J, 1945:25) Vargovitsh, RS, 2009:20

Pygmarrhopalites postumicus ♀ from Holland
Sub-anal appendage in lateral aspect
2019.12.29 © Berg, M.
Pygmarrhopalites postumicus ♀ from Holland
Sub-anal appendage in ventral aspect
2019.12.29 © Berg, M.
Pygmarrhopalites postumicus ♀ from Holland
Ant.4 with 6 subdivisions
2019.12.29 © Berg, M.

Species principalis (Stach, J, 1945:37) Vargovitsh, RS, 2009:9, B

            Syn.: Arrhopalites binoculatus Linnaniemi, 1912, nec Börner, 1901
            Syn.: Arrhopalites binoculatus f. principalis Linnaniemi, 1912, nec Börner, 1901
            Syn.: Sminthurinus binoculatus ab. ? Börner, 1903
            Syn.: Arrhopalites binoculatus Womersley, 1939, nec Börner, 1901


Arrhopalites principalis from the UK
Antenna
2005 © Hopkin, S.P.
Pygmarrhopalites principalis from Belgium
2011.09.06 © Henderickx, H.
Pygmarrhopalites principalis from Belgium
2011.09.06 © Henderickx, H.
Pygmarrhopalites principalis from the UK
2012.11.22 © Murray, A.
Pygmarrhopalites principalis from the UK
4th antennal segment with 6 subdivisions
2012.11.22 © Murray, A.
Pygmarrhopalites principalis from the UK
Anal segment
2012.11.22 © Murray, A.
Pygmarrhopalites principalis from the UK
2013.01.31 © Murray, A.
Pygmarrhopalites principalis from the UK
2013.01.31 © Murray, A.
Pygmarrhopalites principalis from the UK
2013.01.31 © Murray, A.
Pygmarrhopalites principalis from France
2013.06.02 © Alonso, C.
Pygmarrhopalites principalis from France
2018.01.07 © Tiky, S.
Pygmarrhopalites principalis from France
2018.01.07 © Tiky, S.
Pygmarrhopalites principalis from France
Ant.3 at basis enlarged (*)
2018.01.07 © Tiky, S.
Pygmarrhopalites principalis from France
2018.02.17 © Picard, J.
Pygmarrhopalites principalis from Denmark
2019.01.22 © Schlütter, P.
Pygmarrhopalites principalis from France
2019.10.31 © Garcelon, P.
Pygmarrhopalites principalis from France
Cephalic pattern
2019.10.31 © Garcelon, P.
Pygmarrhopalites principalis from France
2019.10.31 © Garcelon, P.
Pygmarrhopalites principalis from France
Facial pattern
2019.10.31 © Garcelon, P.
Pygmarrhopalites principalis from France
2020.03.08 © Garcelon, P.
Pygmarrhopalites principalis from Holland
2021.12.28 © Kamsteeg, G.
Pygmarrhopalites principalis from Holland
Middorsal thoracic pattern
2021.12.28 © Kamsteeg, G.
Pygmarrhopalites principalis from Holland
Middorsal abdominal pattern
2021.12.28 © Kamsteeg, G.
Pygmarrhopalites principalis from Holland
Sub-anal appendage (SAA) apically feathered
2022.01.01 © Kamsteeg, G.


  Subspecies pallida Linnaniemi, WM, 1912:274

            Syn.: Arrhopalites binoculatus var. pallida(sic) Linnaniemi, WM, 1912:274 IOS
            Syn.: Arrhopalites binoculatus var. pallida(sic) Bonet, F, 1928:258 IOS
            Syn.: Arrhopalites binoculatus var. pallida(sic) Delamare Deboutteville, C, 1947:239 IOS
            Syn.: Arrhopalites binoculatus ssp. pallida(sic) Salmon, JT, 1964:571 IOS
            Syn.: Arrhopalites pygmaeus ssp. pallida(sic) Salmon, JT, 1964:575 IOS
            Syn.: Pygmarrhopalites (principalis-group) principalis ssp. pallida(sic) Arbea, JI, 2013:41 IOS

  Subspecies skelicus Vargovitsh, RS, 2009:36



Species pseudoappendices (Rusek, 1967:23) Vargovitsh, RS, 2009:e18

Species pseudoprincipalis Vargovitsh, RS, 2009:31

Species pygmaeus (Wankel, 1860:256 tt) Vargovitsh, RS, 2009:8, B

            Syn.: Smynthurus(sic) ferrugineus Packard, 1888
             Syn.: Arrhopalites ferruginaeus(sic) Christiansen, K, 1966:43 ISS
            Syn.: Smynthurus(sic) mammouthia Banks, 1897
             Syn.: Arrhopalites? mamouthia(sic) Bonet, F et Tellez, C, 1947:193 ISS
             Syn.: Sminthurus mammonthia(sic) Stach, 1956,is ISS
            Syn.: Sminthurinus affinis Börner, C, 1901:710
            Syn.: Sminthurinus binocolatus(sic) Börner, 1901 IOS
            Syn.: Sminthurinus (Articulati) binoculatus Börner, C, 1901:100 n
             Syn.: Arrhopalites binocolatus(sic) Kloet, GS et Hincks, WD, 1945:9 ISS
            Syn.: Arrhopalites binoculatus var. cavernarum Bonet, F, 1928:258
            Syn.: Arrhopalites subboneti Cassagnau & Delamare Deboutteville, 1953
            Syn.: Arrhopalites sp. Stach, 1956
            Syn.: Arrhopalites longicornis Gisin, 1963
            Syn.: Sphyrotheca binoculata Scott, 1964, nec Börner, 1901


             Syn.: Dicyrtoma pigmaea(sic) Absolon, K, 1899:317 ISS
             Syn.: Arrhopalites pygmaea(sic) Bonet, 1931 ISS
             Syn.: Arrhopalites pygmaeeus(sic) Selga, D, 1963:454,B ISS
             Syn.: Arrhopalites pygmaeu(sic) Selga, D, 1963:455,B ISS
             Syn.: Arrhopalites pygmeus(sic) Szeptycki & Weiner in Razowski, 1990 ISS
             Syn.: Arrhopalites pygmeus(sic) Skarzynski, D et Smolis, A, 2002:304 ISS


Arrhopalites pygmaeus from the UK
Antenna
2005 © Hopkin, S.P.
Arrhopalites pygmaeus (bar=2µm)
Aberrant epicuticular ultrastructure with secondary granules
After Nickerl J & al, 2012 Fig.2D'
Arrhopalites pygmaeus (bar=500nm)
Epicuticular secondary granule
After Nickerl J & al, 2012 Fig.2D''
Pygmarrhopalites pygmaeus from the UK
2012.09.06 © Murray, A.
 
Pygmarrhopalites pygmaeus from the UK
2012.09.06 © Murray, A.
 
Pygmarrhopalites pygmaeus from the UK
2012.09.06 © Murray, A.
Pygmarrhopalites pygmaeus from the UK
2012.09.07 © Murray, A.
Pygmarrhopalites pygmaeus from the UK
2013.01.29 © Murray, A.
Pygmarrhopalites pygmaeus from the UK
Subdivisions of 4th antennal segment
2013.01.29 © Murray, A.
Pygmarrhopalites pygmaeus from Canada
2015.07.16 © Murray, A.
Pygmarrhopalites pygmaeus from Canada
2015.07.16 © Murray, A.
Pygmarrhopalites pygmaeus gr. from France
Ant.4 with 5 subdivisions
2018.05.02 © Picard, J.
Pygmarrhopalites pygmaeus gr. from France
2018.05.02 © Picard, J.
Pygmarrhopalites pygmaeus gr. from France
2018.05.02 © Picard, J.
Pygmarrhopalites pygmaeus gr. from France
Spiky cephalic setae (*)
2018.05.02 © Picard, J.
Pygmarrhopalites pygmaeus gr. from France
Unguiculus (u) with long filament reaching beyond tip of unguis (U)
2018.05.02 © Picard, J.
Pygmarrhopalites pygmaeus from Germany
2021.11.14 © Fabian, Be.
Pygmarrhopalites pygmaeus from Germany
Spiky setae on vertex
2021.11.14 © Fabian, Be.
Pygmarrhopalites pygmaeus from Germany
2021.11.14 © Fabian, Be.
Pygmarrhopalites pygmaeus from Germany
Ant.4 with 5 subdivisions
Basal and apical longer
2021.11.14 © Fabian, Be.

  Subspecies grisea Axelson, 1907


            Syn.: Sminthurinus binoculatus var. grisea(sic) Axelson, 1907 IOS
            Syn.: Arrhopalites binoculatus var. grisea(sic) Linnaniemi, WM, 1912:275 IOS
            Syn.: Arrhopalites binoculatus var. grisea(sic) Bonet, F, 1928:258 IOS
            Syn.: Arrhopalites pygmaeus f. grisea(sic) Handschin, E in Dahl, F, 1929:112 IOS
            Syn.: Arrhopalites pygmaeus ssp. grisea(sic) Salmon, JT, 1964:575 IOS



Species ruseki (Nosek, J, 1975:45) Vargovitsh, RS, 2009:20

Species rystsovi Vargovitsh, RS, 2022:14

Pygmarrhopalites rystsovi ♀ from the Caucasus
After Vargovitsh RS, 2022 Fig.10A

Species sacer (Christiansen, K & Bellinger, P, 1996:177) Vargovitsh, RS, 2009:26

Species salemensis Soto-Adames, FN et Taylor, SJ, 2013:167

             Syn.: Pygmarrhopalites salemsis(sic) Soto-Adames, FN et Taylor, SJ, 2013:174 ISS

Species sapo (Zeppelini, D & Christiansen, K, 2003:38) Vargovitsh, RS, 2009:e18

Species secundarius (Gisin, 1958) Vargovitsh, RS, 2009:36

Species sericus (Gisin, 1947) Vargovitsh, RS, 2009:e18, B

Species sextus (Zeppelini, D & Christiansen, K, 2003:40) Vargovitsh, RS, 2009:26

Species shoshoneiensis Zeppelini, D, Taylor, SJ et Slay, ME, 2009:11

Species silvus (Christiansen, K & Bellinger, P, 1996:177) Vargovitsh, RS, 2009:e18

Species slovacicus (Nosek, 1975) Vargovitsh, RS, 2009:20

Species spinosus (Rusek, 1967:23) Vargovitsh, RS, 2009:20, B

Species styriacus (Nosek & Neuherz, 1976) Vargovitsh, RS, 2009:20

Species subbifidus (Travé, J, Gadea, E & Delamare Deboutteville, C, 1954:213) Vargovitsh, RS, 2009:e18

Species subboneti (Cassagnau, P & Delamare Deboutteville, C, 1953:135) Vargovitsh, RS, 2009:e18

Species tauricus Vargovitsh, RS, 2009:20

Species terricolus (Gisin, 1958) Vargovitsh, RS, 2009:e18

            Syn.: Arrhopalites terricola(sic) Gisin, 1958 IOS
            Syn.: Arrhopalites terricola(sic) Gisin, H, 1960:277 IOS
            Syn.: Arrhopalites terricola(sic) Selga, D, 1963:464 IOS
            Syn.: Arrhopalites terricola(sic) Salmon, JT, 1964:576 IOS
            Syn.: Arrhopalites terricola(sic) Betsch, J-M et Cassagnau, P, 1966:123 IOS
            Syn.: Arrhopalites terricola(sic) Rusek, J, 1967:23 IOS
            Syn.: Arrhopalites terricola(sic) Nosek, J, 1975:45 IOS
            Syn.: Arrhopalites terricola(sic) Rusek, J in Dlabola, J, 1977:16 IOS
            Syn.: Arrhopalites terricola(sic) Dallai, R et Abukar, MM, 1981:266 IOS
            Syn.: Arrhopalites terricola(sic) Mateos, E, 1988:101 IOS
            Syn.: Arrhopalites terricola(sic) Dallai, R, Malatesta, E et Ramellini, P in Minelli, A, Ruffo, S et La Posta, S, 1995:13 IOS
            Syn.: Arrhopalites terricola(sic) Nayrolles, P, 1996:130 IOS
            Syn.: Arrhopalites terricola(sic) Nayrolles, P, 1996:131 IOS
            Syn.: Arrhopalites terricola(sic) Nayrolles, P, 1996:135 IOS
            Syn.: Arrhopalites terricola(sic) Curcic, BPM et Lucic, LR, 1997:31P IOS
            Syn.: Arrhopalites terricola(sic) Salmon, S et Ponge, J-F, 1999:1163 IOS
            Syn.: Arrhopalites terricola(sic) Baquero, E, Herrando-Pérez, S et Jordana, R, 2005:81 IOS
            Syn.: Arrhopalites terricola(sic) Russell, DJ et Griegel, A, 2006:174 IOS
            Syn.: Arrhopalites terricola(sic) Fiera, C, 2007:31 IOS
            Syn.: Arrhopalites terricola(sic) Dányi, L et Traser, G, 2008:63 IOS
            Syn.: Arrhopalites terricola(sic) Vargovitsh, RS, 2009:7 IOS
            Syn.: Pygmarrhopalites terricola(sic) Vargovitsh, RS, 2009:e18 IOS
            Syn.: Pygmarrhopalites terricola(sic) Dányi, L, 2011:425 IOS
            Syn.: Pygmarrhopalites terricola(sic) Arbea, JI et Pérez Fernández, T, 2012:17 IOS
            Syn.: Pygmarrhopalites terricola(sic) Arbea, JI, 2013:105 IOS
            Syn.: Pygmarrhopalites terricola(sic) Vargovitsh, RS et Busmachiu, G, 2015:550 IOS
            Syn.: Arrhopalites terricola(sic) Burkhardt, U, Schulz, H-J et Stark, A, 2016:13 IOS
            Syn.: Pygmarrhopalites terricola(sic) Grujic, NZ, Winkler, D et Daghighi, E, 2023:749 IOS

Species texensis (Christiansen, K & Bellinger, P, 1996:178) Vargovitsh, RS, 2009:e18

             Syn.: Arrhopalites texesensis(sic) Zeppelini Filho, D, [1997]:12 ISS

Species thermophilus (Loksa, I, 1964:96) Vargovitsh, RS, 2009:il

Species troglophilus (Palissa, 2000) Vargovitsh, RS, 2009:e19

Species uenoi (Yosii, R, 1956:100) Vargovitsh, RS, 2009:26

Species whitesidei (Jacot, 1938) Vargovitsh, RS, 2009:e19

Species yosiii (Zeppelini, D, 2004:8) Vargovitsh, RS, 2009:20

            Syn.: Arrhopalites yosii(sic) Zeppelini, D, 2004:8 IOS

Species youngsteadtorum Zeppelini, D, Taylor, SJ et Slay, ME, 2009:8

            Syn.: Pygmarrhopalites (pygmaeus-group ss) youngsteadtii(sic) Zeppelini, D, Taylor, SJ et Slay, ME, 2009:8 IOS

Species zlotensis Curcic, BPM et Lucic, LR, 1997:31P

            Syn.: Arrhopalites zloti(sic) Curcic, BPM et Lucic, LR, 1997:31P IOS
Arrhopalites zloti(sic) from East Serbia
After Curcic BPM & Lucic LR, 1997 Fig.1
Arrhopalites zloti(sic) from East Serbia
After Curcic BPM & Lucic LR, 1997 Fig.2

             Syn.: Arrhopalites zloti(sic) Jaksic, P, 2005:194 ISS





Genus Troglopalites Vargovitsh, RS, 2012:10

Species stygios Vargovitsh, RS, 2012:11, t.t.

Troglopalites stygios from Western Caucasus
After Vargotitsh RS, 2012 Fig.60
Troglopalites stygios from Western Caucasus
After Vargotitsh RS, 2012 Fig.61

References