http://www.collembola.org/taxa/isotinae.htm - Last updated on 2017.07.17 by Frans Janssens
Checklist of the Collembola: Isotominae

Subfamilia Isotominae Schäffer, 1896
Genus Number
of species
Aackia1
Acanthomurus6
Agrenia13
Antarcticinella1
Araucanocyrtus1
Axelsonia8
Azoritoma1
Cheirotoma1
Chionobora1
Desoria101
Folsomotoma9
Gnathisotoma5
Halisotoma8
Heteroisotoma6
Hydroisotoma1
Isotoma77
Isotomedia1
Isotomurus68
Kaylathalia1
Marisotoma3
Metisotoma2
Misturasotoma1
Mucronia2
Myopia1
Najtia1
Paracerura9
Parisotoma27
Pentacanthella1
Procerura13
Protodesoria1
Protoisotoma2
Psammisotoma6
Pseudisotoma8
Pseudosorensia4
Pteronychella4
Rhyniella1
Semicerura3
Sericeotoma1
Setocerura6
Spinocerura2
Tibiolatra1
Tomocerura6
Vertagopus27
Isotominae442
Genera with number of species
  Syn.: Entomobryidae Isotominini(sic) Börner, C, 1901:43 ISS
Isotominae from Belgium
2006 © Vuijlsteke, M.
Isotoma viridis from the USA
2009.03.10 © Cowen, R.


Genus Protodesoria Christiansen, KA et Nascimbene, P, 2006:346, -
Species granda Christiansen, KA et Nascimbene, P, 2006:346, -

Genus Tibiolatra Salmon, JT, 1941:320

Tibiolatra latronigra from New Zealand
After Salmon, J.T. 1941 Pl.49 Fig.165.
Species latronigra Salmon, JT, 1941:320, t.t.

Genus Acanthomurus Womersley, 1934
Acanthomurus sp. from Tasmania
2014.04.29 © Murray, A.

Species alpinus Salmon, JT, 1941:317
Acanthomurus alpinus from New Zealand
After Salmon J.T., 1941 Pl.50 Fig.169.

  Subspecies obscuratus Salmon, JT, 1943:381
Acanthomurus alpinus
subsp. obscuratus from New Zealand
After Salmon J.T., 1943 Pl.41 Fig.34.

Species plumbeus Womersley, 1934, t.t.
  Subspecies lineatus Womersley, 1934
Species rivalis Wise, 1964
Species setosus Salmon, JT, 1941:319
Acanthomurus setosus from New Zealand
After Salmon J.T., 1941 Pl.49 Fig.162.

  Subspecies violaceus Salmon, JT, 1941:320
Species sp.nov. Greenslade, P in Dallai, R, 1989:116
Species womersleyi Salmon, JT, 1941:318
Acanthomurus womersleyi from New Zealand
After Salmon J.T., 1941 Pl.50 Fig.176.


Genus Axelsonia Börner, 1906
Species crassicornis (Schött, 1893) Yayuk, RS (sic), 1989:8
Species johnstoni Jordana, 1997
Species littoralis (Moniez, 1890) Denis, 1923
            Syn.: Isotoma nitida Handschin, E in Dahl, F, 1929:60 =?
            Syn.: Axelsonia thalassophila Handschin, E in Dahl, F, 1929:60
            Syn.: Axelsonia australis Bagnall, 1939
             Syn.: Isotoma litoralis(sic) Moniez, 1894 ISS
             Syn.: Axelsonia litoralis(sic) Handschin, E in Dahl, F, 1929:60 ISS
             Syn.: Axelsonia litoralis(sic) Kloet, GS et Hincks, WD, 1945:6 ISS
Species nitida (Folsom, JW, 1899:264) Börner, 1906
Isotoma nitida from Japan
After Folsom JW, 1899 Pl.1 Fig.14

            Syn.: Axelsonia littoralis Yosii, 1955
            Syn.: Axelsonia littoralis Yosii, R, 1977:155
            Syn.: Isotoma (Isotoma) pteromucronata Uchida, H, 1965:88
            Syn.: Isotoma (Isotoma) pteromucronata Yosii, R, 1977:155

Species sarahae Christiansen, K et Bellinger, P, 1992:151
Species thalassophila Börner, 1906, t.t.
Species tubifera Strenzke, 1958
Species yinii Huang & Liang, 1992

Genus Isotomurus Börner, 1903 key  key
  Syn.: Collembola Istoma(sic) Childs, GH, 1915:21 ISS
  Syn.: ... Isotama(sic) Stach, J, 1924:114 ISS
  Syn.: ... Proisotomurus Womersley, 1934
  Syn.: ... Isotomorus(sic) Stach, J, 1924:115 ISS
  Syn.: Isotomidae Isotomourus(sic) Baijal, HN, 1958:355 ISS
  Syn.: Isotomidae Isotomorus(sic) Poinsot, N, 1972:687 ISS
  Syn.: Isotomidae Isomurys(sic) Velázquez, DR et Novoa, NM, 2002:7 ISS
Remark by ICZN. 2007:420: Placed on the Official List of Names in Zoology. Opinion 291.

Isotomurus sp. from the USA
2007 © McClarin, J.
Isotomurus sp. from Norway
Triangular head shape
After Fjellberg A, 2010 p.2
Isotomurus sp. from Norway
3+3 macrosetae on abd.4 in triangle
After Fjellberg A, 2010 p.2
Isotomurus palustris from the UK
Triangular head shape
2010.02.08 © Robertson, A.
Isotomurus graminis from Sweden
Difference between bothriotrichum (bot) and macroseta (mac)
2011.09.24 © Hall, K.
Isotomurus sp. from the UK
Pigmented clypeus
2012.12.18 © Phillips, E.

  Syn.: Isotomidae isotomurus(sic) Glime, JM, 2015:26 ISS
Isotomurus sp. juv. from Holland
2015.01.20 © van Duinen, J.
Isotomurus graminis from Germany
Ovoid post-antennal organ (PAO)
2015.02.14 © Kluck, A.
Isotomurus pseudopalustris from Belgium
Macrosetae only present on posterior body segments
2016.01.04 © Huskens, M.-L.
Isotomurus pseudopalustris from Belgium
Macrosetae on abd.4 in triangular constellation
2016.01.04 © Huskens, M.-L.
Isotomurus graminis from the UK
Mucro with 4 teeth
2016.01.26 © Shepherd, M.J.
Isotomurus maculatus? from Belgium
Ovoid post-antennal organ (PAO)
2016.02.25 © Huskens, M.-L.

Species aetnensis Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:116
Isotomurus aetnensis
2001 © Carapelli, A. et al.
Isotomurus nr aetnensis from the Azores
2010.09.22 © E.D.E.N.

Species afghanicus Yosii, 1966
Isotomurus afghanicus from Afghanistan
After Potapov M, 2001 Fig.374

Species alticolus (Carl, J, 1899:313) Handschin, 1919
             Syn.: Isotomurus alticola(sic) Kloet, GS et Hincks, WD, 1945:7 ISS
Isotomurus alticolus from Poland
After Stach, J., 1947 Plate L Fig.1.
Isotomurus alticolus from Poland
After Stach, J., 1947 Plate LI Fig.1.
Isotomurus alticolus from Poland
After Stach, J., 1947 Plate LI Fig.3.
             Syn.: Isotomurus alticola(sic) Rusek, 1977 ISS
             Syn.: Isotomurus alticola(sic) Dallai & Malatesta in Calderini, 1995 ISS
Isotomurus alticolus from Italy
2001 © Carapelli, A. et al.

Species annectens Yosii, 1963
Species atreus Christiansen & Bellinger, 1980
Species balteatus (Reuter, OM, 1876:86) Handschin, E in Dahl, F, 1929:73
Isotoma palustris var. balteata from Sweden
After Schött H, 1893 Taf.V Fig.10
Isotomurus palustris f. balteata from Poland
After Stach, J., 1947 Plate XLVIII Fig.6
Isotomurus balteatus from Korea
After Yosii R & Lee C-E, 1963 Fig.2A
Isotomurus palustris f. balteata from Poland
After Potapov M, 2001 Fig.371
Isotomurus balteatus
2007 © Fjellberg, A.
Isotomurus balteatus
2007 © Fjellberg, A.
Isotomurus balteatus from India
After Mandal GP & Hazra AK, 2009 Fig.17a

Species beskidensis Rusek, 1963
Species bimaculatus Ågren, 1903
Species bimus Christiansen, K et Bellinger, P, 1980:679
            Syn.: Isotomurus palustris var. balteatus Folsom
Isotomurus palustris ssp. balteatus from the USA
After Snider RJ, 1967 Backcover

Isotomurus bimus from the USA
2007.03.31 © Babin, P.
Isotomurus bimus from the USA
2009.07.05 © Zungri, D.
Isotomurus bimus juv. from the USA
2009.07.06 © Zungri, D.
Isotomurus bimus from the USA
2010.08.03 © Zungri, D.
Isotomurus bimus from the USA
2014.07.20 © Dankowicz, E.
Isotomurus bimus from the USA
2014.07.20 © Dankowicz, E.
Isotomurus bimus from the USA
2014.11.01 © Emile.
Isotomurus bimus from the USA
2015.07.22 © Dankowicz, E.
Isotomurus bimus from the USA
2016.12.24 © Montgomery, G.

Species cassagnaui Deharveng & Lek, 1993
Isotomurus cassagnaui
2001 © Carapelli, A. et al.

Species chaos Potapov & Babenko, 2011
Species chiltoni (Carpenter, GH, 1925:9) Womersley, 1934
Species cibus Christiansen & Bellinger, 1980
Species denticulatus Kos, 1937
Species fucicolus (Reuter, 1891) Axelson, 1906
            Syn.: Isotoma janthina Vogler, 1896
            Syn.: Isotomurus italicus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 1995:94
Isotoma palustris var. fucicola from Sweden
After Schött H, 1893 Taf.V Fig.6

             Syn.: Isotomurus palustris ssp. fucicola(sic) Axelson, 1906 ISS
             Syn.: Isotomurus palustris var. fucicola(sic) Gisin, H, 1960:214 ISS
             Syn.: Isotomurus fucicola(sic) da Gama, 1964 ISS
             Syn.: Isotomorus(sic) fucicola(sic) Poinsot, N, 1972:688 ISS
             Syn.: Isotomurus fucicola(sic) Poinsot, N, 1972:689 ISS
Isotomorus(sic) fucicola(sic) from France
After Poinsot N, 1972 Fig.6D
Isotomurus fucicolus
2001 © Carapelli, A. et al.


             Syn.: Isotomurus fucicola(sic) Fanciulli, PP, Summa, D, Dallai, R et Frati, F, 2001:251 ISS
Isotomurus fucicola(sic) from Scandinavia
2007 © Fjellberg, A.
Isotomurus fucicolus from the UK
2009.09.30 © Robertson, A.
Isotomurus fucicolus from the UK
Abd.3 with 4 bothriotricha
2009.11.26 © Robertson, A.
Isotomurus fucicolus from the UK
2012.04.21 © Murray, A.
Isotomurus fucicolus from the UK
2013.08.10 © Fenwick, D.

             Syn.: Isotomurus fucicola(sic) Santeshwari, X, Raghuraman, M et Singh, J, 2015:572 ISS
Species fuscus (Salmon, JT, 1944:145) Greenslade, P, 2003:1, s.i.
Species gallicus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:120
Isotomurus gallicus from France
2001 © Carapelli, A. et al.

             Syn.: Isotomurus palustris var. balteata(sic) Handschin, E, 1942:445 ISS
            Syn.: Isotomurus balteatus Poinsot, N, 1972:689
            Syn.: Isotomurus balteatus Ellis, WN, 1974:121
            Syn.: Isotomurus balteatus Poinsot-Balaguer & Ferard, 1983
            Syn.: Isotomurus balteatus Deharveng & Lek, 1993
            Syn.: Isotomurus balteatus Sousa & al., 1997
Isotomurus gallicus from France
2008.03.15 © Lebeaux, P.
Isotomurus gallicus from France
2008.03.15 © Lebeaux, P.
Isotomurus gallicus from Norway
2010 © Fjellberg, A.

Species ghibellinus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 1995:94
Isotomurus ghibellinus
2001 © Carapelli, A. et al.

Species graminis Fjellberg, A, 2007:96
            Syn.: Isotomurus palustris var. pallida Schäffer, 1896 =?
            Syn.: Isotomurus palustris var. prasinus Deharveng & Lek, 1993
            Syn.: Isotomurus prasinus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:127
Isotomurus prasinus
2001 © Carapelli, A. et al.
Isotomurus prasinus from Sweden
2005 © Hall, K.
Isotomurus prasinus from the UK
2006 © Cornwall, N.J.
Isotomurus prasinus from the UK
2006 © Cornwall, N.J.

Isotomurus graminis
2007 © Fjellberg, A.
Isotomurus graminis from the UK
2007.03.18 © Cornwall, N.J.
Isotomurus graminis from the UK
2007.04.01 © Cornwall, N.J.
Isotomurus graminis from the UK
2009.03.29 © Michaelson, J.
Isotomurus graminis from the UK
2009.03.29 © Robertson, A.
Isotomurus graminis from the UK
2009.11.15 © Robertson, A.
Isotomurus graminis from the UK
2009.11.22 © Robertson, A.
Isotomurus graminis from Sweden
2011.09.24 © Hall, K.
Isotomurus graminis from the UK
2012.11.21 © Murray, A.
Isotomurus graminis from Holland
2014.01.14 © van Duinen, J.
Isotomurus graminis from Belgium
2014.01.18 © Wallays, H.
Isotomurus graminis ♂ from Holland
2014.02.24 © van Duinen, J.
Isotomurus graminis ♂ from Holland
Spiny setae of male
2014.02.24 © van Duinen, J.
Isotomurus graminis from Germany
2014.09.20 © Beck, K.
Isotomurus graminis from Germany
2014.11.09 © Landgraf, A.
Isotomurus graminis from the UK
2015.01.15 © Thomas, Br.
Isotomurus graminis from Holland
2015.02.06 © van Duinen, J.
Isotomurus graminis from Germany
Ovoid post-antennal organ (PAO)
2015.02.14 © Kluck, A.
Isotomurus graminis from Germany
Macrosetae on abd.4 in triangular constellation
2015.02.14 © Kluck, A.
Isotomurus graminis from Belgium
2016.01.10 © Huskens, M.-L.
Isotomurus graminis from the UK
2016.01.26 © Shepherd, M.J.
Isotomurus graminis from Belgium
2016.11.23 © Huskens, M.-L.
Isotomurus graminis from Belgium
2017.01.31 © Huskens, M.-L.

Species griseus Lee BH et Kim JT in Lee, B-H, Kim, B-J et Kim, J-T, 1993:282
Species hadriaticus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:121
Isotomurus hadriaticus
2001 © Carapelli, A. et al.

Species infuscatus Yosii, 1963
            Syn.: Isotomurus palustris ssp. fucicolus Yosii, 1955
            Syn.: Isotomurus palustris ssp. fucicolus Yosii, R, 1977:156
Species italicus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 1995:94
Isotomurus italicus
2007 © Fjellberg, A.

Species lapidosus (Salmon, JT, 1949:25) Greenslade, P, 2003:1, s.i.
Species lateclavus Kos, 1937
Species lineatus (Salmon, JT, 1941:322) Christiansen, K & Bellinger, P, 2000:114
  Subspecies violaceus Salmon, JT, 1941:323
Species louisiana (Scott, 1962) Waltz, RD, 1998:195
Species maculatus (Schäffer, 1896) Ågren, 1903, B
            Syn.: Isotoma bifasciata Bourlet, C, 1839:401
            Syn.: Desoria (Première division) fulvomaculata Nicolet, H, 1842:59
             Syn.: Isotoma fulvo-maculata(sic) Nicolet, 1847 ISS
            Syn.: Desoria (Première division) virescens Nicolet, H, 1842:59
Desoria virescens from Switzerland
After Nicolet, H, 1842 Pl.5 Fig.12.

            Syn.: Desoria tigrina Bourlet, C, 1845:67
            Syn.: Desoria tigrina Bourlet, C in Erichson, WP, 1846:303
            Syn.: Podura viridis Bourlet, 1843 partim Schött, 1891
            Syn.: Isotoma lineata Lubbock, 1862
            Syn.: Isotoma palustris f. principalis Tullberg, 1872, partim Schött, 1891
            Syn.: Desoria riparia Lubbock, J, 1873:170
            Syn.: Isotomurus palustris f. hortorum Cassagnau, 1987
            Syn.: Isotomurus palustris Greenslade, 2002
             Syn.: Isotomurus palustris var. maculata(sic) Stach, J, 1924:113 ISS
Isotomurus palustris var. maculata(sic) from Tunis
After Stach J, 1924 Taf.II Fig.9
Isotomurus palustris var. maculata(sic) from Malta
After Stach J, 1924 Taf.II Fig.10
Isotomurus palustris var. maculata(sic) from Tunis
After Stach J, 1924 Taf.II Fig.11
Isotomurus palustris var. maculata(sic) from Tunis
After Stach J, 1924 Taf.II Fig.12
Isotomurus palustris f. maculata from Poland
After Stach, J., 1947 Plate XLVIII Fig.1.
Isotomurus palustris f. maculata from Poland
After Stach, J., 1947 Plate XLVIII Fig.4.
Isotomurus palustris f. maculata from Poland
After Stach, J., 1947 Plate XLVIII Fig.5.
Isotomurus palustris ssp. maculatus from France
After Poinsot N, 1972 Fig.6C

             Syn.: Isotomurus maculata(sic) Deharveng, 1981 gr ISS
             Syn.: Isotomurus maculata(sic) Gabriel & al, 1990 gr ISS
Isotomurus maculatus
2001 © Carapelli, A. et al.
Isotomurus maculatus from Italy
After Potapov M, 2001 Fig.368-369
Isotomurus maculatus from the UK
2005 © Hopkin, S.P.
Isotomurus maculatus from the UK
2005 © Brocklehurst, K.
Isotomurus maculatus from Scandinavia
After Fjellberg, A, 2006 Fig.4
Isotomurus maculatus from the UK
2006.04.08 © Read, D.
Isotomurus maculatus from the UK
2007 © Cornwall, N.J.
Isotomurus maculatus from the UK
2007 © Cornwall, N.J.
Isotomurus palustris f. maculatus from the USA
2007 © Moorehead, C.
Isotomurus palustris f. maculatus from the USA
2007 © Moorehead, C.
Isotomurus palustris f. maculatus from the USA
2007 © Moorehead, C.
Isotomurus maculatus
2007 © Fjellberg, A.
Isotomurus maculatus from Italy
2007 © Bartolini, L.
Isotomurus maculatus from the UK
2007.12.01 © Warner, P.
Isotomurus palustris f. maculatus from the USA
2008.05.09 © Zungri, D.
Isotomurus maculatus from Hungary
2008.11.08 © Keresztes, G.
Isotomurus maculatus from the Netherlands
2008.11.14 © Baas, A.H.
Isotomurus maculatus from the Netherlands
2008.11.19 © Baas, A.H.
Isotomurus maculatus from the Netherlands
2008.11.19 © Baas, A.H.
Isotomurus maculatus from the Netherlands
2008.11.19 © Baas, A.H.
Isotomurus maculatus from the Netherlands
2008.11.19 © Baas, A.H.
Isotomurus maculatus from France
2009.02.25 © Lefebvre, V.
Isotomurus maculatus from Belgium
2009.03.08 © De Groot, S.
Isotomurus maculatus from the UK
2009.04.06 © Robertson, A.
Isotomurus maculatus from Tasmania
2009.08.16 © Daley, A.
Isotomurus maculatus from Belgium
2010.01.31 © Wallays, H.

             Syn.: Isotomurus maculata(sic) Greenslade, 2010 gr ISS
             Syn.: Isotomurus maculata(sic) Greenslade, P et Convey, P, 2012:411,gr ISS
Isotomurus maculatus from the UK
2012.11.07 © Jones, A.
Isotomurus maculatus from the UK
2012.11.07 © Jones, A.
Isotomurus maculatus from the UK
2013.12.01 © van Duinen, J.
Isotomurus maculatus from the UK
Pigmented clypeus and labrum
2015.02.05 © Horton, M.
Isotomurus maculatus from the UK
Pre-jump posture
2015.02.05 © Horton, M.
Isotomurus maculatus from Belgium
2016.01.11 © Huskens, M.-L.
Isotomurus maculatus from Belgium
Unguis with basal lateral teeth (*)
2016.03.08 © Huskens, M.-L.
Isotomurus maculatus from Belgium
2016.04.24 © Huskens, M.-L.
Isotomurus maculatus from Belgium
2016.09.04 © Huskens, M.-L.
Isotomurus palustris f. maculatus from the USA
2017.01.22 © Dobbs, Br.
Isotomurus palustris f. maculatus from the USA
2017.02.06 © Dobbs, Br.
Isotomurus maculatus from Belgium
2017.02.07 © Huskens, M.-L.
Isotomurus maculatus from Belgium
Dark clypeus
2017.02.08 © Huskens, M.-L.

Species mankyungensis Kim, JT et Lee, B-H, 2000:181
Species martiniquensis Stach, J, 1947:449
Isotomurus tricuspis ssp. martiniquensis from Martinique
After Stach, J., 1947 Plate XLIX Fig.8.

Species mauretanicus (Handschin, 1926, s.d.) Handschin, 1928
Species myohyangsanus Lee BH et Kim JT in Lee, B-H, Kim, B-J et Kim, J-T, 1993:285
Species nebulosus Lek, S et Carapelli, A, 1998:125
Isotomurus nebulosus from France
After Lek & Carapelli, 1998 Fig.1
Isotomurus nebulosus
2001 © Carapelli, A. et al.

Species nicolae Stomp, 1983
Species novaezealandiae (Salmon, JT, 1941:321) Christiansen, K & Bellinger, P, 2000:114
            Syn.: Proisotomurus novae-zealandiae(sic) Salmon, JT, 1941:321 IOS
Proisotomurus novae-zealandiae(sic) from New Zealand
After Salmon J.T., 1941 Pl.50 Fig.179

             Syn.: Proisotomurus novae-zealandiae(sic) Salmon, JT, 1964:359 ISS
Species obscurus Carpenter, GH, 1916:34
Species opala Christiansen, K et Bellinger, P, 1992:155
            Syn.: Isotomurus palustris var. balteatus Folsom, 1932
Species palliceps (Uzel, 1891) Stach, 1929
Isotomurus palliceps from Poland
After Stach, J., 1947 Plate LII Fig.1.
Isotomurus palliceps from Poland
After Stach, J., 1947 Plate LII Fig.2.

Species palustris (Müller OF, 1776) Szidat, 1926, t.t., B
            Syn.: Podura aquatica cinerea de Geer, C, 1740:63
"Graue Insekten die unter den vorigen schwarzen [Podura aquatica] zusammen auf den Wasser sitzen und etwas grösser sind" from Sweden
After de Geer, C., 1749 Tab.IV Fig.1.

            Syn.: Podura aquatilis Müller OF, 1776
            Syn.: Podura aquatica grisea de Geer, C, 1778:28
Podura aquatica grisea from Sweden
After de Geer, C., 1778 Tom.VII Pl.2 Fig.19.

             Syn.: Podura Grisea(sic) Herbst, JFW, 1787:350 ISS
Podura Grisea(sic) from Germany
After Herbst JFW, 1787 Taf.LXXV Fig.4ab

            Syn.: Podura fuliginosa Templeton, R in Templeton, R et Westwood, JO, 1836:95
            Syn.: Podura Stagnorum(sic) Templeton, R in Templeton, R et Westwood, JO, 1836:93
Podura Stagnorum(sic) from Ireland
After Templeton, R, 1836 Pl.XII Fig.4

            Syn.: Podura armata Nicolet, 1842
            Syn.: Aetheocerus aquaticus Bourlet, 1842
            Syn.: Desoria (Deuxième division) fusca Nicolet, H, 1842:63
            Syn.: Isotoma spilosoma Gervais in Walckenaër, 1844
Isotome spilosome from France
Isotoma spilosoma
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.9


            Syn.: Isotoma palustris var. spilosoma Parfitt, 1891
            Syn.: Desoria riparia Bourlet, C, 1845:67
            Syn.: Desoria riparia Bourlet, C in Erichson, WP, 1846:303
            Syn.: Podura psi Herklots, 1857
            Syn.: Degeeria pi Herklots, 1858
            Syn.: Isotoma tricolor MacGillivray, 1891
            Syn.: Isotoma tricolor MacGillivray, 1896
            Syn.: Isotoma tricolor Handschin, E in Dahl, F, 1929:67
            Syn.: Isotoma tricolor Womersley, 1934
            Syn.: Isotomurus tricolor Womersley, 1934
            Syn.: Isotoma tricolor Stach, J, 1947:422
            Syn.: Isotoma tricolor Cavalloro, 1956
            Syn.: Isotoma tricolor Cavalloro, 1956
            Syn.: Isotomurus tricolor Hart, 1974 =?
            Syn.: Isotomurus tricolor Fjellberg, 1986
            Syn.: Isotoma balteata Reuter, 1876
            Syn.: Isotoma Stuxbergii(sic) Tullberg, 1876 IOS
            Syn.: Isotoma Tullbergi(sic) Moniez, 1889
            Syn.: Isotoma aequalis MacGillivray, 1896
            Syn.: Isotoma palustris f. pallida Schäffer, 1896
             Syn.: Isotomurus palustris ssp. pallida(sic) Salmon, JT, 1964:368 ISS
            Syn.: Isotoma maculata Börner, 1901
            Syn.: Isotoma palustris f. cincta Krausbauer, 1902
            Syn.: Isotoma palustris ab. bimaculata Agren, 1903
            Syn.: Isotomurus palustris ssp. tricuspis Börner, C, 1906:173
            Syn.: Proisotomurus papillatus Womersley, 1934
            Syn.: Isotomurus antennalis Poinsot, N, 1972:687
            Syn.: Isotomurus palustris f. catalana Cassagnau, 1987
Isotomurus palustris f. catalana from France
After Potapov M, 2001 Fig.370

            Syn.: Isotomurus independente Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 1995:94
Isotomurus independente
2001 © Carapelli, A. et al.

            Syn.: Isotomurus sp. Dethier, M et Hubart, J-M, 2000:29,B
Isotoma palustris forma principalis from Sweden
After Schött H, 1893 Taf.V Fig.9
             Syn.: Isotoma pulustris(sic) Stach, J, 1924:114 ISS
Isotomurus palustris from China
After Stach J, 1964 PlateVIII Fig.1
Isotomurus palustris from China
After Stach J, 1964 PlateVIII Fig.2
Isotomurus palustris from China
After Stach J, 1964 PlateVIII Fig.3
Isotomurus palustris ssp. palustris from the USA
After Snider RJ, 1967 Cover
Isotomurus palustris ssp. palustris from France
After Poinsot N, 1972 Fig.6A


             Syn.: Isotomurus palustis(sic) Heungens, A et Van Daele, E, 1973:141,B ISS
             Syn.: Isotomurus pallustris(sic) Greenslade, P in Dallai, R, 1989:111 ISS
Isotomurus palustris
2001 © Carapelli, A. et al.
Isotomurus palustris from Italy
After Potapov M, 2001 Fig.372

Isotomurus palustris from the UK
2005 © Hopkin, S.P.
Isotomurus palustris from the UK
2005 © Hopkin, S.P.
Isotomurus palustris from the UK
2005 © Hopkin, S.P.
Isotomurus palustris from the UK
2006 © Barton, T.
Isotomurus palustris from the UK
2006 © Barton, T.
Isotomurus palustris from Belgium
2006 © Vuijlsteke, M.
Isotomurus palustris from the UK
2007 © Valentine., B
Isotomurus palustris from the USA
2007.02.20 © Moorehead, C.
Isotomurus palustris
2007 © Fjellberg, A.
Isotomurus palustris from the UK
2007.10.16 © Campbell, A.
Isotomurus palustris from the UK
2007.10.19 © Campbell, A.
Isotomurus palustris from the Netherlands
2007.11.08 © Baas, A.H.
Isotomurus palustris from the UK
2007.11.24 © Campbell, A.
Isotomurus palustris from the USA
2008.01.12 © Moorehead, C.
Isotomurus palustris from the UK
2008.01.12 © Farrell, S.
Isotomurus palustris from the Netherlands
2008.01.26 © Baas, A.H.
Isotomurus palustris from Germany
2008.04.17 © Peters, J.
Isotomurus palustris from the Netherlands
2008.11.14 © Baas, A.H.
Isotomurus palustris
2008.12.25 © onefivenine.


             Syn.: Isotomurus palustres(sic) Palacios-Vargas, JG, Mejía-Recamier, BE et Cutz-Pool, LQ, 2009:205 ISS
Isotomurus palustris from the USA
2009.01.07 © Fleming, K.
Isotomurus palustris from the USA
2009.01.31 © Justis, S.
Isotomurus palustris from the USA
2009.02.01 © Justis, S.
Isotomurus palustris from France
2009.02.25 © Lefebvre, V.
Isotomurus palustris from the USA
2009.03.03 © Justis, S.
Isotomurus palustris from the UK
2009.03.07 © Barton, T.
Isotomurus palustris from the Netherlands
2009.03.27 © Baas, A.H.
Isotomurus palustris from the Netherlands
2009.03.27 © Baas, A.H.
Isotomurus palustris from the UK
2009.12.26 © Barton, T.
Isotomurus palustris from the UK
2009.12.26 © Barton, T.
Isotomurus palustris from the UK
Post-antennal organ
2012.01.03 © Kent, J.J.
Isotomurus cf palustris from the UK
2012.08.30 © Murray, A.
Isotomurus cf palustris from the UK
2012.08.30 © Murray, A.
Isotomurus cf palustris from the UK
2012.08.30 © Murray, A.
Isotomurus palustris from the UK
2012.11.07 © Jones, A.
Isotomurus palustris from the UK
2012.11.07 © Jones, A.
Isotomurus palustris from La Palma
2012.12.01 © Barton, T.
Isotomurus palustris from Holland
2014.11.07 © van Duinen, J.
Isotomurus palustris from the UK
Phoretic curled up nematode
2015.02.24 © Thomas., B
Isotomurus palustris from the UK
2015.03.23 © Barton, T.
Isotomurus palustris from the UK
2015.03.23 © Barton, T.
Isotomurus palustris from the UK
Aberrant abdominal pigmentation
2015.12.04 © Barton, T.
Isotomurus palustris from the UK
2016.07.22 © David W.
Isotomurus palustris from the USA
2016.11.28 © Dobbs, Br.
Isotomurus palustris from the USA
2017.01.20 © Fleming, K.

Species palustroides Folsom, JW, 1937:77
Species paraciliatus Cardoso, MA, 1970:109

Isotomurus philfrancisi
2006 © Bernard, E.
Species philfrancisi Bernard, 2006
Species plumosus Bagnall, 1940, B
            Syn.: Isotomurus palustroides ssp. subciliatus Stach, J, 1947:461
Isotomurus palustroides subciliatus from Poland
After Stach, J., 1947 Plate L Fig.1
Isotomurus palustroides subciliatus from Poland
After Stach, J., 1947 Plate L Fig.2

             Syn.: Isotomurus palustroides ssp. subcialiatus(sic) Rusek, J in Dlabola, J, 1977:15 ISS
Isotomurus plumosus from Belgium
2001 © Hopkin, S.P.
Isotomurus plumosus from Poland
After Potapov M, 2001 Fig.373
Isotomurus plumosus from Sweden
2006 © Hall, K.
Isotomurus plumosus
2007 © Fjellberg, A.
Isotomurus plumosus from the UK
2007.04.09 © Cornwall, N.J.
Isotomurus plumosus from Belgium
2007.03.28 © Lindsley, J.
Isotomurus plumosus juv. from France
2008.03.02 © Lebeaux, P.
Isotomurus plumosus from the UK
2010.02.06 © Talbot, M.E.
Isotomurus plumosus from Denmark
2012.03.19 © Bildstrup, O.
Isotomurus plumosus juv. from the UK
2012.10.02 © Murray, A.
Isotomurus plumosus from the UK
2013.06.24 © Fenwick, D.
Isotomurus plumosus from the UK
2013.08.10 © Fenwick, D.
Isotomurus plumosus from Czechia
With Sminthurinus elegans f. ornata
2015.03.05 © Krásenský, P.
Isotomurus plumosus from Holland
2015.12.04 © van Duinen, J.
Isotomurus plumosus from Belgium
2016.03.13 © Janssens, F.
Isotomurus plumosus juv. from Belgium
2016.03.13 © Janssens, F.
Isotomurus plumosus juv. from Belgium
2016.04.17 © Huskens, M.-L.
Isotomurus plumosus from Belgium
2016.07.10 © Janssens, F.
Isotomurus plumosus from Belgium
Postantennal organ (PAO) elliptical with central constriction
2016.08.10 © Huskens, M.-L.

Species pseudopalustris Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:127
            Syn.: Podura palustris ssp. aquatilis Müller OF, 1776
Isotoma aquatilis from the UK
After Lubbock, J., 1873 Pl.37.

             Syn.: Isotoma aquatelis(sic) Parfitt, 1891 ISS
Isotoma palustris var. aquatilis from Sweden
After Schött H, 1893 Taf.V Fig.7


            Syn.: Isotoma trifasciata Bourlet, C, 1839:402
Isotoma trifasciata from the UK
After McIntire SJ, 1867 Fig.37
After McIntire SJ, 1868
Isotoma trifasciata from the UK
After Lubbock J, 1870 Pl.XXII Fig.20
Isotoma trifasciata from the UK
Foot of anterior leg
After Lubbock J, 1870 Pl.XXII Fig.21
            Syn.: Isotoma Gervaisii(sic) Nicolet, H, 1847:374 IOS
Isotoma Gervaisii(sic) from Switzerland
After Nicolet,, 1847 Pl.6 Fig.9


             Syn.: Isotoma Gervaisii(sic) Brauer, 1855 ISS
             Syn.: Isotoma Gervaisii(sic) Lubbock, J, 1873:176 ISS
             Syn.: Isotoma Gervaisii(sic) Stach, J, 1947:367 ISS
Isotomurus pseudopalustris f. gervaisi from Belgium
2004 © Vuijlsteke, M.
Isotomurus pseudopalustris f. gervaisi from Belgium
2004 © Vuijlsteke, M.
Isotomurus pseudopalustris f. gervaisi from Belgium
2004 © Vuijlsteke, M.
Isotomurus pseudopalustris f. gervaisi from Belgium
2005.06.02 © Vuijlsteke, M.
Isotomurus cf. gervaisi from the USA
2007 © Moorehead, C.
Isotomurus pseudopalustris f. gervaisi from Belgium
2008.12.11 © Vuijlsteke, M.
Isotomurus pseudopalustris f. gervaisi from the UK
2011.08.26 © Barton, T.
Isotomurus pseudopalustris f. gervaisi from Holland
2014.02.11 © van Duinen, J.
Isotomurus pseudopalustris f. gervaisi from the UK
2014.12.20 © Evans, K.
Isotomurus pseudopalustris f. gervaisi from Germany
2015.01.20 © Landgraf, A.
Isotomurus pseudopalustris f. gervaisi from the UK
2015.11.25 © Evans, K.
Isotomurus pseudopalustris f. gervaisi from the UK
2015.12.08 © Horton, M.
Isotomurus nr pseudopalustris f. gervaisi from the USA
2016.04.17 © Dobbs, Br.
Isotomurus nr pseudopalustris f. gervaisi from the USA
2016.04.21 © Dobbs, Br.
Isotomurus pseudopalustris f. gervaisi from Belgium
2016.05.08 © Huskens, M.-L.
Isotomurus pseudopalustris f. gervaisi from Belgium
2016.05.27 © Huskens, M.-L.


            Syn.: Isotoma tricolor Schött, H, 1891:22
            Syn.: Isotoma tricolor MacGillivray, AD, 1894:105
            Syn.: Isotoma palustris forma principalis Schött, H, 1893:63 partim
Isotoma palustris forma principalis from Sweden
After Schött H, 1893 Taf.V Fig.8
Isotomurus palustris forma principalis from Germany
After Heymons R & Heymons H, 1909 Fig.11


            Syn.: Isotomurus palustris f. maculata Stach, J, 1947:459 partim
Isotomurus palustris f. maculata from Poland
After Stach, J., 1947 Plate XLVIII Fig.2.
Isotomurus palustris f. maculata from Poland
After Stach, J., 1947 Plate XLVIII Fig.3.


            Syn.: Isotomurus palustris Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 1995:87
Isotomurus pseudopalustris from Italy
After Potapov M, 2001 Fig.366-367
Isotomurus pseudopalustris from the UK
2006 © Brocklehurst, K.
Isotomurus pseudopalustris from the UK
2006.06.24 © Botting, J.
Isotomurus pseudopalustris from Germany
2008.04.17 © Peters, J.
Isotomurus pseudopalustris from the UK
2008.09.24 © Robertson, A.
Isotomurus cf. pseudopalustris from the USA
20100207 © Zungri, D.
Isotomurus pseudopalustris from the UK
2012.10.21 © Nurcombe, E.
Isotomurus pseudopalustris from the UK
2012.10.21 © Nurcombe, E.
Isotomurus pseudopalustris from the UK
2012.11.03 © Nurcombe, E.
Isotomurus pseudopalustris from France
2012.11.05 © Quintin, C.
Isotomurus nr pseudopalustris from France
2013.01.13 © Quintin, C.
Isotomurus pseudopalustris from the UK
2013.01.25 © Cobby, P.
Isotomurus pseudopalustris from Holland
2015.02.24 © van Duinen, J.
Isotomurus pseudopalustris from Holland
Two pale dots on abd.4
2015.10.19 © van Duinen, J.
Isotomurus pseudopalustris from the UK
2015.12.28 © Evans, K.
Isotomurus pseudopalustris from Belgium
2016.01.04 © Huskens, M.-L.
Isotomurus pseudopalustris from the UK
2016.02.07 © David W.
Isotomurus pseudopalustris from the UK
2016.02.07 © David W.
Isotomurus pseudopalustris from the UK
Unguiculus with distinct inner tooth
2016.02.13 © Phillips, E.
Isotomurus pseudopalustris from Belgium
2016.04.09 © Huskens, M.-L.

Species pseudosensillatus de Mendonça, 1990
Species punctiferus Yosii, 1963
             Syn.: Isotomurus punctatus(sic) Yosii, R, 1967:2 lapsus ISS
             Syn.: Isotomurus punctiferous(sic) Gupta & Mukharji, 1977 ISS
Isotomurus punctiferus from Japan
After Potapov M, 2001 Fig.375
Isotomurus nr punctiferus from Norway
2007.03.29 © Fauskanger, I.
Isotomurus nr punctiferus from France
2008.11.08 © Jacquemin, G.
Isotomurus nr punctiferus from the UK
2012.09.25 © Murray, A.
Isotomurus nr punctiferus from the UK
2012.09.25 © Murray, A.
Isotomurus nr punctiferus from the UK
2012.09.25 © Murray, A.
Isotomurus nr punctiferus from the UK
2012.09.25 © Murray, A.
Isotomurus nr punctiferus juv. from the UK
2012.09.25 © Murray, A.
Isotomurus nr punctiferus juv. from the UK
2012.10.02 © Murray, A.
Isotomurus nr punctiferus juv. from the UK
2012.10.02 © Murray, A.

Species qinpuensis Chen, 1985
Species quadrisetosus Rusek, 1971
Species rabili Deharveng & Lek, 1993
Isotomurus rabili
2001 © Carapelli, A. et al.

Species retardatus Folsom, JW, 1937:71
Species riparius de Mendonça, 1990
Species sensillatus Winter, 1967
Species sp. Palacios-Vargas, 1981:89
Species sp. Maes & Palacios-Vargas, 1988:4
Species sp. Palacios-Vargas, 1993:157
Species sp. Palacios-Vargas & Gómez-Anaya, 1995:26
Species sp. Zeppelini & Castaño, 1995:10
Species stepposus Potapov, MB et Starostenko, EV, 2002:331
Isotomurus stepposus from Ukraine
After Potapov MB & Starostenko EV, 2002 Fig.1

Species stuxbergi (Tullberg, T, 1877:35) Babenko & Bulavintsev, 1993
            Syn.: Isotoma Stuxbergii(sic) Tullberg, T, 1877:35 IOS
            Syn.: Isotoma Stuxbergi(sic) var. prasina Reuter, 1891
             Syn.: Isotomurus palustris var. prasina(sic) Axelson, 1905 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Axelson, 1906 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Linnaniemi, 1907 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Wahlgren, 1907 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Collinge & Shoebotham, 1910 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Linnaniemi, 1911 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Linnaniemi, 1912 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Shoebotham, 1914 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Brown, 1918 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Axelson, 1906 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Stach, 1920 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Denis, 1921 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Stach, 1923 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Thompson, 1924 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Stirnimann, 1926 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Jackson, 1928 ISS
             Syn.: Isotomurus palustris f. prasina(sic) Handschin, E in Dahl, F, 1929:73 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Stach, 1929 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Buitendijk, 1930 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Womersley, 1930 ISS
             Syn.: Isotomurus palustris f. prasina(sic) Kseneman, M, 1932:20 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Schubert, 1933 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Schubert, 1933 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Turk, 1933 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Mills, 1934 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Schubert, 1935 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Drenowski, 1937 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Folsom, 1937 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Frenzel, 1937 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Womersley, 1939 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Szent-Ivany, 1940 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Buitendijk, 1941 ISS
             Syn.: Isotomurus palustris f. prasina(sic) Stach, J, 1947:459 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Frenzel, 1949 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Mills & Richards, 1953 ISS
             Syn.: Isotomurus palustris f. prasina(sic) Yosii, 1954 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Goto, 1955 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Grinbergs, 1958 ISS
             Syn.: Isotomurus palustris var. prasina(sic) von Torne, 1958 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Gisin, H, 1960:214 ISS
             Syn.: Isotomurus (Isotomurus) palustris var. prasina(sic) Grinbergs, 1960 ISS
             Syn.: Isotomurus palustris ssp. prasina(sic) Salmon, JT, 1964:368 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Poinsot-Balaguer, 1976 ISS
             Syn.: Isotomurus palustris f. prasina(sic) Yosii, R, 1977:156 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Fjellberg, A, 2007:98=? ISS
            Syn.: Isotomurus palustris ssp. palustris Brimley, 1942
            Syn.: Isotomurus ciliatus Stach, 1947
            Syn.: Isotomurus tricolor Babenko & Bulavintsev, 1993 =?
Isotomurus stuxbergi
2007 © Fjellberg, A.

Species subterraneus Stach, 1946
Species takahashii (Yosii, R, 1940:115) Yosii, 1963
Isotoma takhaashii(sic) from Formosa
After Yosii R, 1940 Fig.7

            Syn.: Isotomurus alticulus ssp. japonicus Yosii, R, 1956:64
            Syn.: Isotomurus alticolus ssp. japonicus Yosii, R, 1977:156
             Syn.: Isotoma takhaashii(sic) Yosii, R, 1940:116 ISS
Species texensis Folsom, JW, 1937:76
Species tricolor (Packard, AS, 1873:34) Lippert & Butler, 1976
            Syn.: Isotoma palustris Folsom, JW, 1901:159
            Syn.: Isotoma palustris Guthrie, JE, 1903:70
            Syn.: Isotoma palustris f. principalis Guthrie, JE, 1903:70
            Syn.: Isotoma palustris Folsom, JW, 1919:11A
            Syn.: Isotomurus palustris Mills, HB, 1934:54
            Syn.: Isotomurus palustris Maynard, EA, 1951:122
            Syn.: Isotomurus palustris Salmon, JT, 1964:362 partim
            Syn.: Isotomurus palustris Salmon, JT, 1964:363 partim
            Syn.: Isotomurus palustris Christiansen, K et Bellinger, P, 1980:687
            Syn.: Isotomurus palustris Christiansen, K et Bellinger, P, 1998:756
Isotomurus (Isotomurus) tricolor from the USA
After Folsom JW in Christiansen K & Bellinger P, 1980 Fig.546A
Isotomurus tricolor from the USA
2006 © Murray, T.
Isotomurus tricolor from the USA
2007.05.18 © Cowen, R.
Isotomurus tricolor from the USA
2008.01.06 © Moorehead, C.
Isotomurus tricolor from the USA
2009.03.04 © Murray, T.
Isotomurus tricolor from the USA
2009.08.19 © Zungri, D.
Isotomurus tricolor from the USA
2010.02.08 © Zungri, D.
Isotomurus tricolor from the USA
2010.01.21 © Murray, T.
Isotomurus tricolor from the USA
2012.01.14 © Justis, S.
Isotomurus tricolor from the USA
2012.09.22 © Murray, T.

Species tricuspis Börner, C, 1906:173
Species unifasciatus (Börner, C, 1901:50) Stach, J, 1947:459, B
            Syn.: Isotoma palustris f. principalis Schött partim
Isotomurus unifasciatus
2001 © Carapelli, A. et al.
Isotomurus unifasciatus from the UK
2005 © Hopkin, S.P.
Isotomurus unifasciatus from the UK
2007 © Barton, T.
Isotomurus unifasciatus from the UK
2007 © Cornwall, N.J.
Isotomurus unifasciatus from the UK
2007 © Cornwall, N.J.
Isotomurus unifasciatus
2007 © Fjellberg, A.
Isotomurus unifasciatus from the UK
2007 © Spears, G.
Isotomurus unifasciatus from Norway
2009.11.29 © Haraldseide, H.
Isotomurus unifasciatus from the UK
2012.01.26 © Kent, J.J.
Isotomurus unifasciatus from the UK
2012.11.02 © Murray, A.
Isotomurus unifasciatus juv. from the UK
2013.01.18 © Nurcombe, E.
Isotomurus unifasciatus from Holland
2013.02.07 © van Duinen, J.
Isotomurus nr unifasciatus from New Zealand
2014.06.01 © Murray, A.
Isotomurus unifasciatus from Germany
2015.01.05 © Beck, K.
Isotomurus unifasciatus juv. from Germany
2015.03.28 © Kluck, A.

Species venustus Barra, J-A, 2006:67
Species viridipalustris Kos, 1937
Species yamaquizuensis Winter, 1967
            Syn.: Isotomurus yamaquizuensis Winter, 1963 nn
Isotomurus cf. yamaquizuensis from Colombia
After Mari Mutt JA, 1987 Fig.11


Genus Chionobora Greenslade, P et Potapov, M, 2015:334
Species amila Greenslade, P et Potapov, M, 2015:334, t.t.
Chionobora amila from Tasmania
After Greenslade P & Potapov M, 2015 Fig.1
Chionobora amila ♂ from Tasmania
After Stevens MI & D'Haese CA, 2016 Fig.1A
Chionobora amila ♀ from Tasmania
After Stevens MI & D'Haese CA, 2016 Fig.1B


Genus Kaylathalia Stevens, MI et D'Haese, CA, 2016:3
Species klovstadi (Carpenter, 1902) Stevens, MI & D'Haese, CA, 2016:3, t.t.
            Syn.: Isotoma Klovstadi(sic) Carpenter, 1902 IOS
             Syn.: Isotoma Klovstadi(sic) Enderlein, 1909 ISS
             Syn.: Isotoma Klovstadi(sic) McNamara, 1919 ISS
             Syn.: Isotoma Klovstadi(sic) Gressitt & Weber, 1959 ISS
Desoria klovstadi from Antarctica
2008.05.13 © O'Brien., B

             Syn.: Chionobora klovsatdi(sic) Stevens, MI et D'Haese, CA, 2016:3 ISS

Genus Hydroisotoma Stach, J, 1947:249
Species schaefferi (Krausbauer, 1898) Stach, J, 1947:250, t.t.
            Syn.: Isotoma schäfferi(sic) Krausbauer, 1898,tt IOS
            Syn.: Isotoma rivularis Latzel, 1921
            Syn.: Archisotoma dimorpha Denis, 1922
            Syn.: Proisotoma Schäfferi(sic) ssp. aculeata Stach, J, 1930:289
             Syn.: Proisotoma schaefferi ssp. aculata(sic) Wray, DL, 1978:205 ISS
            Syn.: Hydroisotoma stachi Bagnall, 1949
             Syn.: Proisotoma (Ballistura) Schäfferi(sic) Heymons, R et Heymons, H in Brauer, A, 1909:5 ISS
Proisotoma Schäfferi(sic) from Germany
After Heymons R & Heymons H, 1909 Fig.6

             Syn.: Isotoma schäfferi(sic) Folsom, 1928 ISS
             Syn.: Proisotoma Schäfferi(sic) Denis, JR, 1931:116 ISS
             Syn.: Isotoma schäfferi(sic) Tarsia, 1943 ISS
             Syn.: Isotoma schäfferi(sic) Gisin, 1944 ISS
             Syn.: Hydroisotoma schäfferi(sic) Stach, J, 1947:250 ISS
Hydroisotoma schäfferi(sic) from Poland
After Stach, J., 1947 Plate XXIXa Fig.1

             Syn.: Isotoma (Isotoma) schäfferi(sic) Maynard, EA, 1951:108 ISS
             Syn.: Isotoma schäfferi(sic) Maynard, EA, 1951:108 ISS
Isotoma schäfferi(sic) from the USA
After Maynard EA PlateXII Fig.198
Isotoma schäfferi(sic) from the USA
After Maynard EA PlateXII Fig.199

             Syn.: Hydroisotoma schafferi(sic) Gisin, H, 1960:213 ISS
             Syn.: Hydroisotoma schafferi(sic) Snider, RJ, 1967:212 ISS
Hydroisotoma schaefferi ♂ from the USA
Habitus lateral
After Wray DL, 1978 Fig.5
Hydroisotoma schaefferi ♂ from the USA
Habitus dorsal
After Wray DL, 1978 Fig.6
Hydroisotoma schaefferi ♂ from the USA
Head with curled antennae
After Wray DL, 1978 Fig.2
Hydroisotoma schaefferi ♂ from the USA
Antenna with sense organs
After Wray DL, 1978 Fig.1
Hydroisotoma schaefferi from Germany
After Schulz, H.-J., 1989
Hydroisotoma schaefferi from Germany
2006 © Schulz, H.-J. & Schubert H.D.
Hydroisotoma schaefferi from the USA
2007 © Murray, T.
Hydroisotoma schaefferi from the USA
Furca with subcylindrical dentes
2007 © Murray, T.
Hydroisotoma schaefferi from the USA
2007.03.01 © Murray, T.
Hydroisotoma schaefferi from the USA
2008.01.13 © Murray, T.
Hydroisotoma schaefferi from France
2008.03.16 © Lebeaux, P.
Hydroisotoma schaefferi juv. from France
2009.09.dd © Lebeaux, P.
Hydroisotoma schaefferi juv. from France
2009.09.dd © Lebeaux, P.
Hydroisotoma schaefferi from the USA
2011.02.20 © Husic, C.
Hydroisotoma schaefferi from the USA
Furca with subcylindrical dentes
2012.12.02 © Fleming, K.
Hydroisotoma schaefferi ♂ from the USA
Male with bent apical antennal segments and warts on 2nd antennal segment
2013.03.28 © Murray, T.

             Syn.: Hydroisotoma schafferi(sic) Glime, JM, 2015:3 ISS
Hydroisotoma schaefferi from Germany
2015.03.31 © Schulz, H.-J.
Hydroisotoma schaefferi from the USA
2015.04.16 © Mallory, C.


Genus Agrenia Kloet, GS et Hincks, WD, 1945:6 key

Species-group bidenticulata
Remark by Fjellberg, A. & Bernard, E.C. 2009:19: Lateral mucronal seta present.

Species sp.1 Fjellberg, A, 1988:112
Species bidenticulata (Tullberg, T, 1877:35) Handschin, E in Dahl, F, 1929:71, t.t., B
            Syn.: Isotoma lanuginosa Carl, J, 1899:307
            Syn.: Agrenia bidenticulata var. elongata Carpenter, 1908
            Syn.: Agrenia bidenticulata var. nigra Linnaniemi, 1935
Agrenia bidenticulata from Belgium
After Jacquemart S, 1975 p.83
Agrenia bidenticulata from Russia
2006 © Boldsystems.org
Agrenia bidenticulata from the USA
2011.05.30 © Eiseman, C.
Agrenia bidenticulata from Norway
2015.mm.dd © Fjellberg, A.
Agrenia bidenticulata from Buryatia
With long claws
After Potapov M, Gulgenova A & Babykina M, 2016 Fig.37

Species lamellosa Fjellberg, A, 1988:110
Species pilosa Fjellberg, A, 1986:101
Species polymorpha Fjellberg, A, 1986:96

Species-group agilis
Remark by Fjellberg, A. & Bernard, E.C. 2009:20: Lateral mucronal seta absent.

Species agilis Fjellberg, A, 1986:101
Agrenia agilis from the USA
After Fjellberg A & Bernard EC, 2009 Fig.3A

Species atroviridis Fjellberg, A, 1986:104
Agrenia atroviridis from the USA
2008.02.09 © Murray, T.
Agrenia atroviridis from the USA
After Fjellberg A & Bernard EC, 2009 Fig.3B

Species cyanura Fjellberg, A, 1986:102
Agrenia cyanura from the USA
After Fjellberg A & Bernard EC, 2009 Fig.3C
Agrenia cyanura from the USA
After Fjellberg A & Bernard EC, 2009 Fig.3D

Species extrema Fjellberg, A et Bernard, EC, 2009:24
Species falcula Fjellberg, A et Bernard, EC, 2009:25
Agrenia falcula from the USA
After Fjellberg A & Bernard EC, 2009 Fig.5A
Agrenia falcula from the USA
After Fjellberg A & Bernard EC, 2009 Fig.5B

Species parkeri Fjellberg, A et Bernard, EC, 2009:20
Agrenia parkeri from the USA
2006 © Bernard, E.C.
Det. Bernard, E.C., 2010.01.25
Agrenia parkeri ♀ from the USA
After Fjellberg A & Bernard EC, 2009 Fig.4A
Agrenia parkeri ♂ from the USA
After Fjellberg A & Bernard EC, 2009 Fig.4A
Agrenia parkeri from the USA
After Fjellberg A & Bernard EC, 2009 Fig.4B

Species riparia Fjellberg, A, 1986:103
Species tarashchukae Fjellberg, A et Bernard, EC, 2009:27

Genus Pteronychella Börner, C, 1909:113
Species andina Yoshii, 1990
Species babai Yosii, 1939
Species ezoensis Yosii, 1965
Species perpulchra Börner, C, 1909:113, t.t.

Genus Isotomedia Salmon, JT, 1944:148
Species triseta Salmon, JT, 1944:148, t.t.

Genus Procerura Salmon, JT, 1941:326
Species aequaoculata Salmon, JT, 1941:328
Species dissimilis (Salmon, JT, 1944:146) Christiansen, K & Bellinger, P, 2000:107

Procerura fasciata from New Zealand
After Salmon, J.T. 1941 Pl.51 Fig.198.
Species fasciata Salmon, JT, 1941:329

Species fusca (Salmon, JT, 1941:330) Christiansen, K & Bellinger, P, 2000:107
  Subspecies pallida Salmon, JT, 1941:331
Species indica (Baijal, HN, 1958:355) Greenslade, P, 2003:1, s.i.
Species magnifica (Salmon, JT, 1949:29) Greenslade, P, 2003:1, s.i.
Species montana Salmon, JT, 1941:328

Isotoma parva from New Zealand
After Salmon, J.T. 1937 Pl.49 Fig.a.
Species parva (Salmon, JT, 1937:352) Greenslade, P, 2003:1, s.i.
             Syn.: Papillomurus parva(sic) Salmon, JT, 1964:372 ISS
Species purpurea Salmon, JT, 1941:328
Species ochracea (Salmon, JT, 1949:27) Christiansen, K & Bellinger, P, 2000:107
Species serrata Salmon, JT, 1941:329
Species transequatoria (Salmon, 1969) Greenslade, P, 2003:1, s.i.
            Syn.: Papillomurus transequatoria(sic) Salmon, 1969 IOS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:24 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:74 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:78 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:115 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:116 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:126 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:128 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:178 ISS
Species violacea Salmon, JT, 1941:327, t.t.

Genus Spinocerura Salmon, 1941

Spinocerura capillata from New Zealand
After Salmon, J.T. 1941 Pl.52 Fig.212.
Species capillata Salmon, JT, 1941:332, t.t.

Species dreuxi (Delamare Deboutteville & Massoud, 1966)

Genus Setocerura Salmon, JT, 1949:33
Species georgiana (Schäffer, 1891) Wise, KAJ, 1970:198
Species maruiensis (Salmon, JT, 1941:326) Deharveng, L & de Oliveira, EP, 1994:442

Tomocerura rubenota from New Zealand
After Salmon, J.T. 1941 Pl.51 Fig.193.
Species rubenota (Salmon, JT, 1941:325) Salmon, JT, 1949:33, t.t.

Species sp. Greenslade, P in Dallai, R, 1989:112
Species striata (Schäffer, 1897)
Species wahlgreni (Womersley, H, 1939:321) Salmon, 1949

Genus Tomocerura Wahlgren, 1901
Species colonavia Salmon, JT, 1949:31
Species conjungens (Schäffer, 1897) Wahlgren, E, 1906:17
Species faguagae Izarra, 1970
Species georgiana (Schäffer, 1897) Massoud & Rapoport, 1968
Species picta Wahlgren, 1901, t.t.
Species swani (Womersley, 1934)
Isotoma swani from Australia
After Womersley H, 1939 Pl.1 Fig.2.


Genus Isotoma Bourlet, C, 1839:399 key
Isotoma from Germany
Schematic sagittal section
After Anonymous Fig.E
  Syn.: Arthropleona Isootoma(sic) Richards, WR, 1968:15 ISS
Remark by ICZN. 2007:419: Placed on the Official List of Names in Zoology. Opinion 291.

Isotoma sp. from the USA
2009.01.29 © Justis, S.
Isotoma sp. from Tunesia
Predation by pseudoscorpion Garypus
2009.04.14 © Henderickx, H.
Isotoma sp. from Norway
3+3 macrosetae on abd.4 in one line
After Fjellberg A, 2010 p.2
Isotoma sp. from Norway
More elongate head
After Fjellberg A, 2010 p.2
Isotoma nr viridis from the USA
2010.01.15 © Parker, H.S.
Isotoma nr viridis from the USA
2010.01.15 © Parker, H.S.
Isotoma nr viridis from the USA
2010.01.15 © Parker, H.S.

Species acrea Wray, 1953
Species anglicana Lubbock, 1862
Isotoma anglicana from the UK
After Lubbock J, 1873 Pl.38
Isotoma anglicana from the UK
2005 © Hopkin, S.P.
Isotoma anglicana from the UK
Manubrial thickening with 4 teeth
2005 © Hopkin, S.P.
Isotoma anglicana from Germany
Manubrial thickening with 4 teeth
Det. Schulze, H.J.
2005 © Mebes, H.
Isotoma anglicana from Sweden
2006 © Hall, K.
Isotoma anglicana juv. from Holland
On snow
2006.03.02 © Hamers., B
Isotoma anglicana from the UK
2007.03.25 © Barnes, C.
Isotoma anglicana from Holland
2008.01.27 © Baas, A.H.
Isotoma anglicana from Belgium
2008.02.07 © Hanssens., B
Isotoma anglicana from Holland
2008.02.08 © Baas, A.H.
Isotoma anglicana from the UK
2008.04.13 © Kilford., B
Isotoma anglicana from France
2008.11.13 © Lavoué,B.
 
Isotoma anglicana from the UK
2008.12.10 © Talbot, M.
Isotoma anglicana from the UK
2009.03.01 © Kilford., B
Isotoma anglicana from Holland
2009.03.19 © Baas, A.H.
Isotoma anglicana from Holland
2009.03.19 © Baas, A.H.
Isotoma anglicana juv. from Holland
2009.03.19 © Baas, A.H.
Isotoma anglicana from the UK
2009.03.21 © Talbot, M.
Isotoma anglicana from Holland
2009.03.20 © Baas, A.H.
Isotoma anglicana from the UK
2009.08.20 © Ransom, T.
Isotoma anglicana from Norway
After Fjellberg A, 2010 p.3
Isotoma anglicana from Norway
Manubrial dorsal macrosetae: A>B
After Fjellberg A, 2010 p.8
Isotoma anglicana from Norway
3 dorsal basal dental macrosetae
After Fjellberg A, 2010 p.9
Isotoma anglicana juv. from the UK
2010.02.01 © Robertson, A.
 
Isotoma anglicana juv. from the UK
2010.02.01 © Robertson, A.
Isotoma anglicana juv. from the UK
2010.02.01 © Robertson, A.
Isotoma anglicana from Holland
2010.02.06 © van Duinen, J.
Isotoma anglicana from the UK
2010.05.02 © Robertson, A.
Isotoma anglicana from the UK
2011.12.01 © Jones, A.
Isotoma anglicana from the UK
2011.12.01 © Jones, A.
Isotoma anglicana from the UK
2012.02.16 © Talbot, M.
Isotoma anglicana from Holland
Ovoid post-antennal organ (PAO)
2012.05.13 © van Duinen, J.
Isotoma anglicana from Holland
Abdominal pigmentation
2012.12.14 © Hamers., B
Isotoma anglicana from Belgium
On snow
2013.02.15 © Hamers., B
Isotoma anglicana from Holland
2014.02.21 © van Duinen, J.
Isotoma anglicana from Belgium
2014.03.02 © Huskens, M.-L.
Isotoma anglicana from Germany
2015.01.17 © Landgraf, A.
Isotoma anglicana from Holland
2015.02.04 © van Duinen, J.
Isotoma anglicana from Holland
2015.03.10 © van Duinen, J.
Isotoma anglicana from the UK
2015.08.16 © Boardman, P.
Isotoma anglicana from the UK
Manubrial thickening with 2+2 teeth (*)
2015.08.16 © Boardman, P.
Isotoma anglicana from Holland
2016.03.17 © van Duinen, J.
Isotoma anglicana from Holland
Distinctly spotted skin
2016.03.17 © van Duinen, J.

Species arctica Schött, H, 1893:61
Isotoma viridis var. arctica from the American Arctics
After Schött H, 1893 Taf.V Fig.4
Isotoma viridis var. arctica from Alaska
After Folsom JW, 1902 Pl.IV Fig.5

             Syn.: Isotoma viridis var. artica(sic) Stach, J, 1924:114 ISS
Isotoma arctica from Alaska
After Fjellberg A, 2003 Fig.9
Isotoma arctica from the USA
Det. Christiansen, K.
2006 © Schimming, L.

Species aspera Bacon, 1914
Species burkhardti Winter, 1967
Species caeca Reuter, 1880
Species caerulea Bourlet, C, 1839:401, B
             Syn.: Isotoma coerulea(sic) Nicolet, H, 1847:374 ISS
             Syn.: Isotoma viridis f. coerulea(sic) Handschin, E in Dahl, F, 1929:67 ISS
             Syn.: Isotoma (Isotoma) viridis var. coerulea(sic) Palissa, A, 1960:378 ISS
             Syn.: Isotoma (Isotoma) viridis ssp. coerulea(sic) Palissa, A in Schwoerbel, J et Zwick, P, 2000:100 ISS
             Syn.: Isotoma coerulea(sic) Fjellberg, A, 2003:94 ISS
             Syn.: Isotoma cerulea(sic) Gudleiffsson, BE et Bjarnadottir, B, 2008:56 ISS
Isotoma caerulea from Belgium
2008.02.07 © Hanssens., B
Isotoma caerulea from the UK
2008.02.27 © Spears, G.
Isotoma caerulea from the UK
2009.03.25 © Ransom, T.
Isotoma caerulea from Norway
After Fjellberg A, 2010 p.4
Isotoma caerulea from Norway
Manubrial dorsal macrosetae: A<B
After Fjellberg A, 2010 p.8
Isotoma caerulea from Norway
2 dorsal basal dental macrosetae
After Fjellberg A, 2010 p.9
Isotoma caerulea from Denmark
2010.11.10 © Bildstrup, O.
Isotoma caerulea from the UK
2011.12.05 © Jones, A.
Isotoma caerulea from the UK
2012.11.08 © Jones, A.
Isotoma caerulea from the UK
2013.09.22 © Murray, A.
Isotoma caerulea from Germany
2015.12.18 © Kluck, A.
Isotoma caerulea from the UK
2015.12.17 © David W.
Isotoma caerulea from Holland
2016.11.15 © van Duinen, J.
Isotoma caerulea from France
2017.02.19 © Ramel, A.

Species crassicornis (Handschin, 1926) Weitschat, W & Wichard, W, 2002:86, -
Species decorata Brown, JM, 1926:205
Isotoma viridis var. decorata from Mesopotamia
After Brown JM, 1926 Fig.11

            Syn.: Isotoma viridis var. decorata f. obscura Handschin, E, 1942:405
Isotoma viridis-decorata(sic) f. obscura from Palestine
After Handschin E, 1942 Fig.1a

            Syn.: Isotoma viridis var. decorata f. pallida Handschin, E, 1942:405
Isotoma viridis-decorata(sic) f. pallida from Palestine
After Handschin E, 1942 Fig.1b

            Syn.: Isotoma viridis var. decorata f. principalis Handschin, E, 1942:405
             Syn.: Isotoma viridis-decorata(sic) Handschin, E, 1942:404 ISS
Species delta MacGillivray, 1896
            Syn.: Isotoma catena Guthrie, JE, 1903:69
Isotoma delta from the USA
2007.03.03 © Montgomery, G.
Isotoma delta from the USA
2007.12.29 © Bryant, P.
Isotoma delta from the USA
2008.03.08 © Justis, P.S.
Isotoma delta from the USA
2008.03.12 © Houston, S.
Isotoma delta from the USA
2008.05.09 © Bradford, A.
Isotoma delta from the USA
2008.11.08 © Justis, S.
Isotoma delta from the USA
2008.11.15 © Justis, S.
Isotoma delta from the USA
2008.11.15 © Justis, S.
Isotoma delta from the USA
2008.11.26 © Justis, S.
Isotoma delta from the USA
2008.12.15 © of Baltimore City, T.
Isotoma delta from the USA
2009.01.16 © Montgomery, G.
Isotoma delta from the USA
2009.04.16 © of Baltimore City, T.
Isotoma delta from the USA
2009.04.27 © Kropiewnicki, T.
Isotoma delta from the USA
2011.03.18 © Hoyer, R.
Isotoma delta from the USA
2011.03.18 © Hoyer, R.
Isotoma delta from the USA
2012.01.04 © Kouri, J.
Isotoma delta juv. from the USA
2012.01.14 © Justis, S.
Isotoma delta from the USA
2015.03.10 © Himmler, K.
Isotoma delta from the USA
2015.12.31 © Schmidt, M.L.
Isotoma delta from the USA
2016.12.11 © McDonald, G.

Species diverticula Yosii, 1966

Isotoma exiguadentata from New Zealand
After Salmon, J.T. 1941 Pl.53 Fig.232.
Species exiguadentata Salmon, JT, 1941:336

Species fasciata Börner, C, 1909:112
            Syn.: Isotoma virgata Yosii, 1963 =?
Species finitima Scherbakov, 1899, s.d.
Species fulva Schäffer, 1897
Species galapagoensis Jacquemart, 1975 in Peck SB & Jacquemart J, 2011:3 u
Species gandhi Rapoport & Rubio, 1968
Species glauca Packard, AS, 1873:33
Species gracilliseta Börner, C, 1909:113
Species grana Lee BH et Kim JT in Lee, B-H, Kim, B-J et Kim, J-T, 1993:285
Species hibernica Carpenter, 1906

Podura himalayana from the Himalayas
After Baijal, H.N., 1955 Fig.1
Species himalayana (Baijal, HN, 1955:175)

            Syn.: Podura himmalayana(sic) Baijal, HN, 1955:175 IOS
Species incerta (Nicolet, H, 1847:370) Brook, G, 1884:280
Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Species innominata Yosii, 1963
Species iranica Arbea, JI et Kahrarian, M, 2015:77
Isotoma iranica from Iran
After Arbea JI & Kahrarian M, 2015 Fig.23

Species kosiana Bagnall, 1949
Species littoralis (Tanaka, 1981)
Species longispina (Mac Gillivray, 1893) Folsom, 1916:480
             Syn.: Isotoma longipenna(sic) Poole, RW in Poole, RW et Gentili, P, 1997:51 ISS
Species mackenziana (Hammer, 1953) Fjellberg, A, 2003:96
Species malvinensis Wahlgren, E, 1906:13
Species manubriata Mac Gillivray, 1896
Species masatierrae Schött, 1921
Species melanocephala Börner, C, 1909:112
Species modesta Nicolet, 1845
Species nepalica (Yosii, 1966)
Species nicoleti Gervais in Walckenaër, 1844
            Syn.: Isotoma Nicoletii(sic) Gervais in Walckenaër, 1844 IOS
             Syn.: Isotama(sic) Nicoletii(sic) Nicolet, H, 1847:374,si ISS
Species nishihirai Yosii, 1965
Species ornata Wahlgren, E, 1906:15
Species pallidafasciata Salmon, JT, 1941:336
Species perkinsi Carpenter, 1904
Species pinnata Börner, C, 1909:110
            Syn.: Isotoma (Isotoma) pinnata var. coracina Börner, C, 1909:112
            Syn.: Isotoma pinnata var. coracina Uchida, 1938
            Syn.: Isotoma pinnata var. coracina Uchida, 1957
            Syn.: Isotoma pinnata var. coracina Yosii, 1963
            Syn.: Isotoma pinnata ssp. coracina Salmon, JT, 1964:392
             Syn.: Isotoma pinnata ssp. voracina(sic) Salmon, JT, 1964:392 ISS
            Syn.: Isotoma pinnata var. coracina Yosii, R, 1977:157
            Syn.: Isotoma (Isotoma) pinnata var. v-album(sic) Börner, C, 1909:112 IOS
             Syn.: Isotoma pinnata var. v-album(sic) Uchida, 1938 ISS
             Syn.: Isotoma pinnata var. v-album(sic) Uchida, 1957 ISS
             Syn.: Isotoma pinnata ssp. album(sic) Salmon, JT, 1964:392 ISS
             Syn.: Isotoma pinnata var. v-album(sic) Yosii, R, 1977:157 ISS
             Syn.: Isotoma pinnata var. v-album(sic) Potapov, M, 2001:282 ISS
            Syn.: Isotoma pinnata var. alba Uchida, 1943
            Syn.: Isotoma pinnata var. alba Uchida, 1957
            Syn.: Isotoma pinnata ssp. alba Salmon, JT, 1964:392
            Syn.: Isotoma pinnata var. alba Yosii, R, 1977:157
Species plumosa (Salmon, 1969) Lawrence, PN, 1978:269
             Syn.: Rhodanella plumose(sic) Mandal, GP et Hazra, AK, 2009:116 ISS
Species protocinerea Handschin, 1926 -
Species rhopalota Scherbakov, AM, 1898:9
Isotoma rhopalota from Russia
After Scherbakov AM, 1898 Table I Fig.9

Species riparia (Nicolet, H, 1842:62) Nicolet, H, 1847:373, B
Desoria riparia from Switzerland
After Nicolet, H., 1842 Pl.6 Fig.6

            Syn.: Podura aquatica grisea de Geer, 1740
            Syn.: Podura palustris Bourlet, 1843
            Syn.: Isotoma viridis var. aquatilis Schött, 1891
Isotoma viridis var. riparia from Sweden
After Schött H, 1893 Taf.V Fig.2
Isotoma viridis var. riparia from Sweden
After Schött H, 1893 Taf.V Fig.3
Isotoma viridis var. riparia from Germany
After Heymons R & Heymons H, 1909 Fig.10
Isotoma riparia from Norway
After Fjellberg A, 2003 Fig.8
Isotoma riparia from Estonia
2006.03.18 © Tartes, U.
Isotoma riparia from the UK
After Shaw P, 2009 ©
Isotoma riparia from France
Predation by pseudoscorpion
2009.11.16 © Lebeaux, P.
Isotoma riparia from Norway
After Fjellberg A, 2010 p.6
Isotoma riparia from Sweden
2013.04.07 © Lind, J.
Isotoma riparia from Sweden
2013.04.07 © Lind, J.
Isotoma riparia from Sweden
2013.04.20 © Lind, J.
Isotoma riparia from Holland
2014.01.22 © Wieringa, P.
Isotoma riparia from Holland
2014.01.22 © Wieringa, P.
Isotoma riparia from the UK
2014.02.04 © Valentine., B
Isotoma riparia juv. from the UK
2014.09.16 © Garlick, T.
Isotoma riparia juv. from the UK
2014.09.16 © Garlick, T.
Isotoma riparia juv. from the UK
2014.10.24 © Garlick, T.
Isotoma nr riparia from Estonia
2015.02.07 © Tartes, U.
Isotoma riparia juv. from Holland
2015.07.02 © van Leeuwen, J.
Isotoma riparia from Holland
2016.03.05 © de Wilde, A.
Isotoma nr riparia from Canada
2016.03.13 © Schmidt, H.
Isotoma nr riparia from Canada
2016.03.13 © Schmidt, H.
Isotoma nr riparia from Canada
2016.03.13 © Schmidt, H.
Isotoma nr riparia from Canada
2016.03.13 © Schmidt, H.

Species sarkundensis Baijal, HN, 1958:354
Species schalleri Winter, 1967
Species semenkevitshi Scherbakov, AM, 1898:8
Species silvatica Schäffer, 1897
Species sinensis Yue & Yin, 1999
Species sp. auct. anon., 1879
Species sp. Wise, 1970, s.i.
Species sp. Ellis, 1974:121, i.s.
Species sp. Maes & Palacios-Vargas, 1988:4
Species sp. Palacios-Vargas, 1990:21
Species sp. Palacios-Vargas & Gómez, 1991
Species sp. Palacios-Vargas & Gómez-Anaya, 1995:26
Species sp. Zeppelini & Castaño, 1995:10
Species sp. Palacios-Vargas, JG et Gamboa-Vargas, JA, 1997:90
Species sp. Brand, RH et Dunn, CP, 1998:238
Species sp. Brand, RH et Dunn, CP, 1998:238
Species sp. Rebek, EJ, Young, DK et Hogg, DB, 1999:60
Species spelaea Joseph, 1882
Species spinicauda (Bonet, 1930) Denis, JR, 1936:263
            Syn.: Isotoma turkestanica Stach, 1947
            Syn.: Isotoma decorata Martynova & Chikatunov, 1968 =?
            Syn.: Isotoma decorata Martynova, 1968 =?
            Syn.: Isotoma decorata Yosii, 1971 =?
            Syn.: Isotoma decorata Martynova & Vtorov, 1973 =?
            Syn.: Isotoma decorata Vtorov & Martynova, 1974 =?
            Syn.: Isotoma decorata Vtorov & Martynova, 1975 =?
            Syn.: Isotoma (Isotoma) decorata Tanaka & al, 1978=?
            Syn.: Isotoma (Isotoma) decorata Janetschek, 1990=?
            Syn.: Acanthisotoma quadridentata Huang & Liang, 1994 =?
Species subcaerulea Kos, 1952
Species subviridis Folsom, JW, 1937:115
Species tahitiensis Salmon, 1964
Species termitophila Womersley, 1934
Species tesselata Rapoport, 1962
Species tigrinella Wahlgren, E, 1906:14
            Syn.: Isotoma tigrina Schött, 1899 nec Nicolet, 1842
Species tridenticulata Schäffer, 1896
             Syn.: Isotoma 3-denticulata(sic) Absolon, K, 1901:32 ISS
Species tridentifera Schött, H, 1917:9
Isotoma tridentifera from Australia
After Schött H, 1917 Fig.12

  Subspecies edenticulata Womersley, 1934
Species tridentina Tarsia in Curia, 1941
Species turbotti (Salmon, JT, 1949:30) Greenslade P, 1990
Species vaillanti Murphy, 1958
Species variodentata Dunger, WG, 1982:60
  Forma marginata Dunger, 1982:62
  Forma principalis Dunger, 1982:63
  Forma unidentata Dunger, 1982:62
Species viridis Bourlet, C, 1839:401, B
            Syn.: Podura viatica Linnæus, 1746
            Syn.: Podura aquatilis Müller OF, 1776, sensu Gisin, 1960
            Syn.: Podura plumbea Müller, 1776
Isotoma plumbea from the USA
After Packard AS, 1889 Pl.10 Fig.6
Isotoma plumbea from the USA
After Packard AS, 1889 Pl.10 Fig.7


            Syn.: Podura fuliginosa Templeton, R in Templeton, R et Westwood, JO, 1836:95
            Syn.: Isotoma arborea Bourlet, C, 1839:94
            Syn.: Desoria (Deuxième division) cylindrica Nicolet, H in Desor, E, 1841:385
            Syn.: Desoria (Deuxième division) annulata Nicolet, H, 1842:62
Isotoma viridis var. annulata from the USA
2007.03.27 © McClarin, J.
Isotoma viridis var. annulata from the USA
2007.04.30 © Cowen, R.
Isotoma viridis var. annulata from Holland
2008.10.10 © Baas, A.H.
Isotoma viridis var. annulata from the UK
2013.10.05 © Evans, J.
Isotoma viridis var. annulata from Germany
2015.01.01 © Beck, K.
Isotoma viridis var. annulata juv. from Germany
2015.01.16 © Landgraf, A.
Isotoma viridis var. annulata from Holland
2015.02.05 © van Duinen, J.
Isotoma viridis var. annulata from the USA
2015.03.22 © Abbott, J.C.
Isotoma viridis var. annulata juv. from Belgium
2016.03.28 © Huskens, M.-L.


            Syn.: Desoria (Deuxième division) ebriosa Nicolet, H, 1842:61
            Syn.: Desoria (Deuxième division) fusca Nicolet, H, 1842:63
            Syn.: Desoria (Deuxième division) fusca var. var. Nicolet, H, 1842:63
            Syn.: Desoria (Deuxième division) pallida Nicolet, H, 1842:61
Isotoma viridis var. pallida from Germany
2005 © Menzel, E.
Isotoma viridis var. pallida from the UK
2006 © Barton, T.
Isotoma viridis var. pallida from Sweden
2006 © Hall, K.
Isotoma viridis var. pallida
2014.01.19 © Evans, J.


            Syn.: Desoria (Première division) virescens Nicolet, H, 1842:59
            Syn.: Heterotoma chlorata Gervais in Walckenaër, 1844
Hétérotome verte from France
Heterotoma chlorata
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.6

            Syn.: Isotoma Desmarestii(sic) Gervais in Walckenaër, 1844 IOS
Isotome de Desmarest from France
Isotoma Desmarestii(sic)
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.11

             Syn.: Isotoma Desmarestii(sic) Nicolet, H, 1847:374,si ISS

            Syn.: Isotoma lineata Lubbock, 1862
            Syn.: Isotoma palustris Tullberg, 1871
            Syn.: Isotoma palustris var. unicolor Tullberg, 1871
            Syn.: Isotoma Belfragei(sic) Packard, AS, 1873:33 IOS
             Syn.: Isotoma Belfragei(sic) MacGillivray, 1891 ISS
            Syn.: Isotoma purpurascens Packard, AS, 1873:34
            Syn.: Isotoma tricolor Packard, 1873 partim
            Syn.: Isotoma tricolor MacGillivray, 1891 partim
            Syn.: Isotoma tricolor Schött, 1893
            Syn.: Isotoma tricolor von Dalla Torre, 1895
            Syn.: Isotoma tricolor MacGillivray, 1896 partim
            Syn.: Isotoma tricolor Folsom, 1903 partim
            Syn.: Isotoma tricolor Linnaniemi, 1912
            Syn.: Isotoma tricolor Bacon, 1914 partim
            Syn.: Isotoma tricolor Bartholin, 1916
            Syn.: Isotoma tricolor Folsom, JW, 1919:12A
            Syn.: Isotoma tricolor Handschin, E in Dahl, F, 1929:67
            Syn.: Isotoma tricolor Mills, HB, 1934:63
            Syn.: Isotoma tricolor Stach, J, 1947:422 partim
            Syn.: Isotoma tricolor Maynard, EA, 1951:110
            Syn.: Isotoma tricolor Uchida, 1957
            Syn.: Isotoma tricolor Salmon, JT, 1964:403
            Syn.: Isotoma tricolor Christiansen, K et Bellinger, P, 1998:853
            Syn.: Isotoma cincta Tullberg, 1876
Isotoma viridis var. cincta from Novaja Selmja
After Schött H, 1893 Taf.V Fig.5

            Syn.: Isotoma viridis var. arctica Schött, H, 1893:61
            Syn.: Isotoma riparia Lameere, A, 1895:594
            Syn.: Isotoma capitola MacGillivray, 1896
            Syn.: Isotoma delta Mac Gillivray, 1896
            Syn.: Isotoma glauca MacGillivray, 1896
            Syn.: Isotoma montana MacGillivray, 1896
            Syn.: Isotoma tridentata MacGillivray, 1896
            Syn.: Isotoma viridis var. violacea Lie-Pettersen, 1898
Isotoma viridis var. violacea from Holland
2008.01.23 © Baas, A.H.
Isotoma viridis var. violacea from Holland
2008.01.27 © Baas, A.H.
Isotoma viridis var. violacea from Holland
2009.04.05 © Baas, A.H.
Isotoma viridis var. violacea from Holland
2014.01.05 © van Duinen, J.
Isotoma viridis var. violacea from France
2014.01.17 © Sandra.
Isotoma viridis var. violacea from France
2014.01.17 © Sandra.


            Syn.: Isotoma paradoxa Carl, J, 1899:317
            Syn.: Isotoma viridis f. genuina Börner, C, 1901:50
            Syn.: Isotoma catena Guthrie, JE, 1903:69
            Syn.: Isotoma viridis f. micans Ågren, 1903
            Syn.: Isotoma hibernica Carpenter, 1906 =?
            Syn.: Isotoma aspera Bacon, 1914
            Syn.: Isotoma spinicauda Bonet, 1930
            Syn.: Isotoma (Isotoma) viridis var. pomona Folsom, JW, 1937:115
            Syn.: Isotoma turkestanica Stach, 1947
            Syn.: Isotomurus bemakus Wray, 1952
            Syn.: Orchesella rubra Scott, 1963 partim holotype
            Syn.: Rhodanella plumosa Salmon, 1969 =?
            Syn.: Tetracanthura mirabilis Martynova, 1971
            Syn.: Desoria riparia Poole, RW in Poole, RW et Gentili, P, 1997:51
            Syn.: Desoria riparia Christiansen, K et Bellinger, P, 1998:853
Isotoma viridis forma principalis from Sweden
After Schött H, 1893 Taf.V Fig.1
Isotoma viridis from Alaska
After Folsom JW, 1902 Pl.IV Fig.4
Isotoma viridis f. typica from Germany
After Heymons R & Heymons H, 1909 Fig.9
             Syn.: Isotoma ciridis(sic) Salmon, JT, 1964:407 ISS
Isotoma viridis from the UK
2005 © Hopkin, S.P.
Isotoma viridis from the UK
2006 © Valentine., B
Isotoma viridis from Germany
2006 © Steiner, A.
Isotoma viridis from the UK
2006.04.08 © Read, D.
Isotoma viridis from the USA
2007 © Murray, T.
Isotoma viridis from the USA
2007 © Murray, T.
Isotoma viridis from Germany
2007 © Peters, J.
Isotoma viridis from the UK
2007.02.23 © Valentine., B
Isotoma viridis from the UK
2007.02.24 © Barton, T.
Isotoma viridis from the UK
2007.02.28 © Valentine., B
Isotoma viridis from the UK
2007.03.02 © Barton, T.
Isotoma viridis from the UK
2007.03.05 © Barton, T.
Isotoma viridis from the Netherlands
2007.03.21 © Baas, A.H.
Isotoma viridis from the USA
2007.10.21 © Moorehead, C.
Isotoma viridis from the UK
2007.12.29 © Barton, T.
Isotoma viridis from the UK
2007.12.29 © Barton, T.
Isotoma viridis from the Netherlands
2008.01.22 © Baas, A.H.
Isotoma viridis from the UK
2008.02.03 © Tonsbeek, M.
Isotoma viridis from Belgium
2008.03.23 © Oger, P.
Isotoma viridis from the UK
2008.04.07 © Robertson, A.
Isotoma viridis from the Netherlands
2008.10.08 © Baas, A.H.
Isotoma viridis from the Netherlands
2008.11.16 © Baas, A.H.
Isotoma viridis from Germany
2009.01.12 © Peters, J.
Isotoma viridis from Germany
2009.01.12 © Peters, J.
Isotoma viridis from the USA
2009.02.10 © Cowen, R.
Isotoma viridis from the USA
2009.03.05 © Cowen, R.
Isotoma viridis from the USA
2009.03.04 © Murray, T.
Isotoma viridis from Norway
After Fjellberg A, 2010 p.5
Isotoma viridis from Norway
Single ventroapical manubrial tooth
After Fjellberg A, 2010 p.7
Isotoma viridis from the UK
2010.02.06 © Barton, T.
Isotoma viridis from the UK
2010.02.14 © Robertson, A.
Isotoma viridis from the UK
2011.12.03 © Jones, A.
Isotoma viridis (bar=2µm)
Hexagonal-rhombic epicuticular ultrastructure
After Nickerl J & al, 2012 Fig.2B'
Isotoma viridis (bar=500nm)
Rhombic epicuticular ultrastructure
After Nickerl J & al, 2012 Fig.2B''
Isotoma viridis from Holland
2012.01.27 © van Duinen, J.
Isotoma viridis from the UK
2012.11.08 © Jones, A.
Isotoma viridis from Holland
2015.02.06 © van Duinen, J.
Isotoma viridis from Germany
2015.02.15 © Kluck, A.
Isotoma viridis from Holland
With parasitic larva of Acari:Erythraeoidea
2015.07.26 © van Duinen, J.
Isotoma viridis from Belgium
Collophore pressed against surfacewaterfilm
2016.01.14 © Huskens, M.-L.
Isotoma viridis from the UK
2016.02.15 © Horton, M.
Isotoma viridis from Belgium
Cephalic pigmentation
2016.11.27 © Huskens, M.-L.
Isotoma viridis from Belgium
Post antennal organ (PAO), half the size of nearest ocellus
2016.11.27 © Huskens, M.-L.


Genus Pseudisotoma Handschin, 1924
  Syn.: Isotoma Pseudoisotoma(sic) Kloet, GS et Hincks, WD, 1945:6 ISS
  Syn.: Collembola Psedisotoma(sic) Hornung-Leoni, CT et Palacios-Vargas, JG, 2012:67 ISS
Species agrana (Lee BH & Kim JT in Lee, B-H, Kim, B-J & Kim, J-T, 1993:288) Potapov, 2001

Pseudisotoma anamola from Lebanon
After Christiansen, K., 1958 Pl.10 Fig.29.
Species anamola Christiansen, 1958:76

Species himalayana Yosii, 1971
Species microchaeta Cassagnau, P, 1959:85
Species monochaeta (Kos, 1942) Jacquemart, S, 1967:4, B
Species ohtanii (Yosii, R, 1972:90) Potapov, 2001
Species sensibilis (Tullberg, T, 1877:36) Agrell, I, 1939:8, t.t., B
            Syn.: Isotoma turicensis Haller, 1880
            Syn.: Isotoma dubia Reuter, 1895
            Syn.: Isotoma determinata Mac Gillivray, 1896
            Syn.: Isotoma parva Mac Gillivray, 1896
            Syn.: Isotoma longidens Schäffer, 1896
            Syn.: Isotoma monstrosa Schäffer, 1896
            Syn.: Isotoma inopinata Axelson, 1902
            Syn.: Isotoma arborea Evans, 1908
Pseudisotoma sensibilis from Belgium
2006.10.12 © Schelfhout, L.
Pseudisotoma sensibilis from Belgium
2008.06.09 © Vuijlsteke, M.
Pseudisotoma sensibilis from Holland
2012.10.28 © van Duinen, J.
Pseudisotoma sensibilis from the USA
Abd.5+6 fused
2016.12.21 © Dobbs, Br.
Pseudisotoma sensibilis from the USA
2016.12.21 © Dobbs, Br.
Pseudisotoma sensibilis from the USA
2016.12.21 © Dobbs, Br.

Species wisei Deharveng, 1980

Genus Semicerura Maynard, EA, 1951:99
Species bishopi Maynard, EA, 1951:99, t.t.
Semicerura nr bishopi
2012.01.25 © Walter, D.E.
Semicerura nr bishopi
2012.01.25 © Walter, D.E.

Species goryshini Martynova, 1969
Semicerura goryshini
2006 © Boldsystems.org
Semicerura goryshini
2006 © Boldsystems.org
Semicerura goryshini
2006 © Boldsystems.org

Species multispinata (James, 1933) Christiansen, KA in Dindal, DL, 1990:987
             Syn.: Semicerura miltispinata(sic) Christiansen, KA in Dindal, DL, 1990:987 ISS

Genus Heteroisotoma Stach, 1947
  Syn.: Isotomidae Sibirisotoma Rusek, 1991
Species carpenteri (Börner, C, 1909:109), t.t.
            Syn.: Isotoma (Isotoma) andrei Mills, HB, 1934:58
            Syn.: Isotoma (Isotoma) difficilis Folsom, JW, 1937:89
Species dilatosetosa Martynova, 1980
Species heteronomica Jie, D, Potapov, M et Sokolova, E, 2011:55
            Syn.: Heteroisotoma sp.nov. Potapov & Polyakov, 2007
Species jacutica Martynova, 1980
Species sinorossica Jie, D, Potapov, M et Sokolova, E, 2011:51
Heteroisotoma sinorossica from China
After Jie & al, 2011 Fig.25

Species stebaevae (Rusek, 1991) Potapov & Stebaeva, 1992
            Syn.: Sibirisotoma stebajevae(sic) Rusek, 1991 IOS
             Syn.: Heteroisotoma stebajevae(sic) Jie, D, Potapov, M et Sokolova, E, 2011:55 ISS
             Syn.: Heteroisotoma stebaievae(sic) Stevens, MI et D'Haese, CA, 2016:4 ISS

Genus Folsomotoma Bagnall, 1949
Species anomala (Salmon, 1948)
Species bioculata (Womersley, 1934) Salmon, JT, 1964:412, t.t.
            Syn.: Isotoma bi-oculata(sic) Womersley, 1934,tt IOS
Species boerneri (Enderlein, 1903)
            Syn.: Isotoma börneri(sic) Enderlein, 1903 IOS
Species kerguelensis (Enderlein, 1903)
             Syn.: Isotoma octo-oculata(sic) ssp. Kerguelensis(sic) Carpenter, GH, 1907:476 ISS
Species marionensis (Deharveng, 1981) McGaughran, A, Stevens, MI & Holland, BR, 2010:50
Species minuta Salmon, JT, 1949:24
Species octooculata (Willem, V, 1901:262)
            Syn.: Isotoma octo-oculata(sic) Willem, V, 1901:262 IOS
            Syn.: Isotoma octo-oculata(sic) var. gracilis Carpenter, GH, 1907:476
Isotoma octo-oculatus(sic) var. gracilis
from Antarctica
After Carpenter GH, 1907 Plate II Fig.9

            Syn.: Isotoma masatierrae Stach, 1947 =?
             Syn.: Isotoma octo-oculata(sic) Willem, V, 1902:13 ISS
Isotoma octo-oculata from Antarctica
After Willem V, 1902 Pl.IV Fig.7

             Syn.: Isotoma octo-oculata(sic) Womersley, 1937 ISS
             Syn.: Parisotoma octo-oculata(sic) Salmon, 1949 ISS
             Syn.: Parisotoma octo-oculata(sic) Brown, 1964 ISS
             Syn.: Parisotoma octo-oculata(sic) Salmon, 1964 ISS
             Syn.: Isotoma 8-oculata(sic) Deharveng, 1981 ISS
  Subspecies ovata Salmon, JT, 1949:39
Species punctata (Wahlgren, E, 1906:14)
Species subflava (Salmon, JT, 1949:22)

Genus Desoria Agassiz M & Nicolet H in Desor E, 1841:384
  Syn.: Isotomurs(sic) Fjellberg, A, 2007:99 ISS
Desoria sp. from France
2008.02.24 © Jacquemin, G.
Desoria sp. from the USA
2009.12.29 © Parker, H.S.

Remark by Duffin, C. 2009:60: Bearing in mind how much Cécile (wife of Agassiz) detested Desor as a social parasite blighting the Agassiz family it is interesting to speculate how she might have appreciated the irony of his lending his name to a 'flea'.

Desoria sp. from the USA
2010.01.23 © Justis, S.
Desoria sp. from Singapore
2011.01.04 © Koh, J.
Desoria sp. from the UK
2011.12.20 © Kent, J.J.
Desoria sp. from Tasmania
2014.04.16 © Murray, A.

Species sp.1 Fjellberg, A, 2010:40
Species sp.2 Fjellberg, A, 2010:40
Species ahiehie Christiansen, K et Bellinger, P, 1992:166
Species alaskensis Fjellberg, 1978
Species albella (Packard, AS, 1873:32)
            Syn.: Isotoma Walkerii(sic) Packard, 1871
             Syn.: Isotoma walkerii(sic) Poole, RW in Poole, RW et Gentili, P, 1997:51 ISS
            Syn.: Isotoma leonina Packard, AS, 1873:32
Species albicornis Fjellberg, A, 2010:32
Desoria albicornis from the USA
After Fjellberg A, 2010 Fig.115C
Desoria albicornis from the USA
After Fjellberg A, 2010 Fig.111

Species antennalis (Bagnall, 1940), B
Isotomurus(sic) antennalis
2007 © Fjellberg, A.
Desoria antennalis from the UK
2010.04.04 © Botting, J.
Desoria antennalis from the UK
2012.07.15 © Murray, A.
Desoria antennalis from the UK
2012.07.15 © Murray, A.
Desoria antennalis from the UK
2012.07.15 © Murray, A.
Desoria antennalis from the UK
2012.07.15 © Murray, A.
Desoria antennalis from the UK
2012.07.15 © Murray, A.
Desoria antennalis from the UK
2012.07.18 © Murray, A.
Desoria antennalis from the UK
2013.06.12 © Murray, A.
Desoria antennalis from Holland
Diffuse pigmentation
2016.02.16 © Wieringa, P.
Desoria antennalis from Holland
2016.02.19 © van Duinen, J.
Desoria antennalis from Holland
2016.02.19 © van Duinen, J.
Desoria antennalis from Holland
Diffuse pigmentation
2016.02.19 © Wieringa, P.

Species arctica White A in Sutherland, PC, 1852:ccviii
Desoria arctica from the Arctic region
After White A in Sutherland PC, 1852 Fig.7.

Species ater Fjellberg, A, 2010:34
Desoria ater from the USA
After Fjellberg A, 2010 Fig.115D
Desoria ater from the USA
After Fjellberg A, 2010 Fig.113

Species atkasukiensis Fjellberg, 1978
Species banffensis Christiansen, K et Bellinger, P, 1998:794
Species bendixenae Christiansen, K et Bellinger, P, 1992:168
Species blekeni Leinaas, 1980
Species blufusata Fjellberg, 1978
Species caeruleatra (Guthrie, JE, 1903:70)
Species canadensis Brown, 1932
            Syn.: Isotoma olivacea Stach, J, 1947:404
Isotoma (Desoria) canadensis from Canada
After Brown in Christiansen K & Bellinger P, 1980 Fig.583A
Desoria canadensis from Canada
2016.03.12 © Schmidt, H.
Desoria canadensis from Canada
2017.01.02 © Levac, G.

Species cancellarei (Christiansen & Bellinger, 1980) Fjellberg, A, 2010:33
Species capra Fjellberg, A, 2010:40
Species cera Christiansen, K et Bellinger, P, 1998:804
Species choi (Lee, B-H, 1977:166)
Species christianseni Fjellberg, 1978
Species creli Fjellberg, 1978
Species cryophila Fjellberg, A, 2010:38
Desoria cryophila from the USA
After Fjellberg A, 2010 Fig.115G

Species divergens (Axelson, 1900) Kaprus, IJ, Pomorski, RJ, Skarzynski, D & Potapov, MB, 2005:894
Species duodecimoculata Denis, 1927
             Syn.: Desoria duodecemaculata(sic) Fiera, C, 2007:23 ISS
Species elegans (Carl, J, 1899:319)
Species eureka Christiansen, K et Bellinger, P, 1998:806
Species fennica (Reuter, 1895)
             Syn.: Isotoma Formáneki(sic) Absolon, K, 1901:212 ISS
Species fideli (Rapoport & Izarra, 1962)
Species fjellbergi Najt, 1981
Species flora (Christiansen & Bellinger, 1980)
Desoria flora from the USA
2014.02.19 © Klein, T.

Species garibaldii Fjellberg, A, 2010:36
Desoria garibaldii from the USA
After Fjellberg A, 2010 Fig.115F

Species gelida Folsom, JW, 1937:106
Desoria gelida from the USA
2010.04.01 © Murray, T.
Desoria gelida from the USA
2010.04.30 © Hodge, K.

Species gorodkovi Martynova, 1969
Species hiemalis (Schött, H, 1893:70) Bulimar, 1987
            Syn.: Isotoma Theobaldi(sic) Carl, J, 1899:315 IOS
            Syn.: Isotoma violacea var. mucronata Axelson, 1900
            Syn.: Isotoma nivicola Axelson, 1900
Desoria hiemalis from Germany
2006 © Kohl, F.
Desoria hiemalis from the USA
2007.02.18 © Murray, T.
Desoria hiemalis from Western Estonia
2009.02.26 © Tartes, U.
Desoria hiemalis from Russia
2013.01.01 © Serge, L.

Species hissarica Martynova & Chikatunov, 1968
Species hoodensis Christiansen, K et Bellinger, P, 1998:814
Desoria hoodensis? from the USA
2017.01.28 © Dobbs, Br.
Desoria hoodensis? from the USA
2017.01.28 © Dobbs, Br.
Desoria hoodensis? from the USA
2017.01.28 © Dobbs, Br.
Desoria hoodensis? from the USA
2017.01.28 © Dobbs, Br.
Desoria hoodensis? from the USA
2017.03.12 © Dobbs, Br.

Species husaini Yosii & Ashraf, 1965
Species imparidentata (Stach, J, 1964:13) Potapov, 2001
Species infuscata Murphy, 1959
Species intermedia (Schött, 1902) Fjellberg, A, 2007:125, B
            Syn.: Isotoma germanica Hüther & Winter, 1961
Remark by Potapov, M. 2001:120: Obscure species.

Desoria intermedia from Scandinavia
2008.04.04 © Fjellberg, A.

Species iuxta Dunger, WG, 1982:68
Species japonica (Yosii, 1939) Yosii, 1954
Species jayasrae Bhattacharjee, 1984
Species juberthiei Gers & Najt, 1983
Species kaszabi Dunger, WG, 1982:70
Species komarkovae Fjellberg, 1978
Species kuraica (Martynova, 1974:19)
Species lucama Wray, 1952
Species manitobae Fjellberg, 1978
Species marissa (Folsom, JW, 1937:89) Miranda Rangel & Palacios-Vargas, 1992
Species maxillosa Fjellberg, 1978
Species mazda Yosii, 1971
Species mossopi (Womersley, H, 1934:454)
Species multisetis (Carpenter & Phillips, 1922) Miranda Rangel & Palacios-Vargas, 1992
Species nahuelensis (Rapoport & Rubio, 1968)
Species negishina (Börner, C, 1909:108) Yosii, R, 1977:156, s.i.
            Syn.: Isotoma sp. Andreen, A, Malterre, J, Needham, J, Olivo, O, Pessoa, SB et Zhukov-Verezhnikov, NN, 1952:18
Species neglecta (Schäffer, 1900) Dányi, L & Traser, G, 2008:46
            Syn.: Isotoma affinis Axelson, 1900
            Syn.: Isotoma tigrina ab. coerulea Ågren, 1903
Desoria neglecta from Norway
2011 © Fjellberg, A.

Species nigra Mac Gillivray, 1896
Species nigrifrons Folsom JW in Mills, HB, 1934:60
Species nivalis Papon, 1856
Species nivalis (Carl, 1910) Miranda Rangel & Palacios-Vargas, 1992
Species nivea (Schäffer, 1896) Fjellberg, 1978
            Syn.: Isotoma albella? Skarzynski, D, Pomorski, RJ, Smolis, A, Weiner, WM, Szeptycki, A, Slawska, M et Sterzynska, M, 2002:35=?
Species nixoni Fjellberg, 1978
Species nympha Snider, RJ et Calandrino, FJ, 1987:5
Isotoma (Desoria) nympha from the USA
After Snider RJ & Calandrino FJ, 1987 Fig.1

Species occulta (Börner, C, 1909:110) Yosii, R, 1977:156
Species olivacea (Tullberg, 1871) Yosii, R, 1977:156, B
            Syn.: Isotoma grisea Lubbock, 1870
            Syn.: Isotoma voraginum Uzel, 1891
            Syn.: Isotoma olivacea f. genuina Börner, C, 1901:56
            Syn.: Isotoma olivacea f. principalis Linnaniemi, 1912
            Syn.: Spinisotoma regina Stach, J, 1930:290
            Syn.: Isotoma grisescens Denis, 1931
            Syn.: Isotoma tigrina Denis, 1937
            Syn.: Isotoma trispinata Tuxen, 1944 partim
            Syn.: Spinisotoma ephippiata Brian, 1945
            Syn.: Proisotoma subsegmentata Scott, 1959
Desoria olivacea from Holland
2009.04.05 © Baas, A.H.
Desoria olivacea from Holland
2010.04.06 © van Duinen, J.
Desoria olivacea from Holland
2010.04.06 © van Duinen, J.
Desoria olivacea from Norway
2011 © Fjellberg, A.
Desoria olivacea from the UK
2012.10.31 © Murray, A.
Desoria olivacea from the UK
2012.10.31 © Murray, A.
Desoria olivacea from the UK
Ovoid post-antennal organ (PAO)
2012.10.31 © Murray, A.
Desoria olivacea from the UK
Abd.5 mesosetae 2/3rds length abd.5
2012.10.31 © Murray, A.
Desoria olivacea from Holland
2014.05.04 © Wieringa, P.
Desoria olivacea from Belgium
2017.04.20 © Huskens, M.-L.

Species olympica Fjellberg, A, 2010:39
Species pallipes (Kos, 1952)
Species pamirensis Martynova, 1975
Species pilifrons Fjellberg, A, 2010:37
             Syn.: Desoria pilifons(sic) Fjellberg, A, 2010:28 ISS
Species pjasini Martynova, 1974
             Syn.: Desoria pjaisini(sic) Greenslade, P et Potapov, M, 2015:338 ISS
Species potapovi Fjellberg, A, 2007:126
Species propinqua (Axelson, 1902) Potapov, 2001
            Syn.: Isotoma poppiusi Axelson, 1903
            Syn.: Spinisotoma pectinata Stach, 1926
Spinisotoma pectinata from Poland
After Stach, J., 1947 Plate XXXVIII Fig.1.

            Syn.: ... propinqua var. hexaspina Agrell, 1936
            Syn.: Desoria sp. Yosii, R, 1977:155
            Syn.: Isotoma (Desoria) graeca Najt, 1981

Species pseudomaritima (Stach, J, 1947:436)
Isotoma pseudomaritima from Poland
After Stach, J., 1947 Plate XLVII Fig.1.

Species quadra Christiansen & Bellinger, 1980
Species randiella Fjellberg, 1978
Species rosea Fjellberg, A, 2010:38
Species saltans Agassiz M & Nicolet H in Desor E, 1841:385 tt
Desoria saltans from Switzerland
After Nicolet H. in Desor E., 1841 Pl.2 fig.1.
            Syn.: ... puce des neiges rougeatre et velue Aristote 350 BC
            Syn.: Desoria (Première division) glacialis Nicolet, H, 1842:58
Desoria glacialis from Switzerland
After Nicolet H., 1842 Pl.5 fig.10.

Isotoma saltans from Austria
After Kopeszki H, 2000 Abb.3
Desoria saltans from Austria
2016.02.27 © van Leeuwen, J.

Species sandersoni Wray, 1952
Species spatiosa (Uchida, H & Tamura, H, 1968:10) Potapov, 2001
Pteronychella spatiosa from Japan
After Suetsugu N, Satoh F & Koike T, 2010 Photo31

Species speciosa Mac Gillivray, 1896
Species subaequalis Folsom, JW, 1937:107
Species subantarctica (Izarra, 1972)
Species tadzhika Martynova & Chikatunov, 1968
Species taigicola Fjellberg, 1978
Species taimyrica (Martynova, 1974) Potapov, 2001
Species tianshanica Hao & Huang, 1995
Species tigrina Nicolet, H, 1842:59, B
Desoria tigrina from Switzerland
After Nicolet H, 1842 Pl.5 Fig.11

            Syn.: Isotoma grisea Lubbock, J, 1870:278
Isotoma grisea from the UK
After Lubbock J, 1873 Pl.39

            Syn.: Isotoma grisescens Schäffer, 1896
             Syn.: Isotoma olivacea var. griscescens(sic) Handschin, E in Dahl, F, 1929:70 ISS
            Syn.: Isotoma affinis Axelson
            Syn.: Isotoma stachi Denis, 1929
            Syn.: Spinisotoma dimorpha Womersley, 1934
            Syn.: Isotoma olivacea Womersley, 1934
            Syn.: Isotoma olivacea Womersley, 1939
            Syn.: Isotoma klovstadi Womersley, 1937
Desoria tigrina from the UK
Setae on fifth abdominal segment
2005 © Hopkin, S.P.
Desoria tigrina from France
2008.02.24 © Jacquemin, G.
Desoria tigrina from Holland
2011.03.22 © van Duinen, J.
Desoria tigrina from the UK
2012.11.05 © Jones, A.
Desoria tigrina from the UK
2012.11.05 © Jones, A.
Desoria tigrina juv. from Iran
Ecomorphic form
After Kahrarian M & Arbea J, 2013 Fig.2
Desoria tigrina juv. from Iran
Ecomorphic abdominal spines
After Kahrarian M & Arbea J, 2013 Fig.3
Desoria tigrina from Iran
After Kahrarian M & Arbea J, 2013 Fig.4
Desoria tigrina from Holland
2015.02.06 © van Duinen, J.
Desoria tigrina from the UK
2016.01.07 © Barton, T.
Desoria tigrina from France
2016.12.06 © Garcelon, P.
Desoria tigrina from the UK
2017.02.01 © David W.
Desoria-tigrina from France
2017.02.05 © Picard, J.
Desoria tigrina juv. ecomorph from Belgium
Abd.5 with transverse row of spines
2017.04.26 © Huskens, M.-L.
Desoria tigrina juv. ecomorph from Belgium
Abd.5 with transverse row of spines
2017.04.26 © Huskens, M.-L.

Species tolya Fjellberg, A, 2007:120
Species torildae Fjellberg, 1978
Species triangularis Fjellberg, A, 2010:33
Desoria triangularis from the USA
After Fjellberg A, 2010 Fig.115E
Desoria triangularis from the USA
After Fjellberg A, 2010 Fig.110

Species trispinata (Mac Gillivray, 1896) Thibaud, 1977
            Syn.: Isotoma maritima ssp. meridionalis Altner, H, 1963:47
             Syn.: Isotoma meridionale(sic) Poinsot, 1971 ISS
             Syn.: Isotoma meridionale(sic) Poinsot, N, 1972:686 ISS
            Syn.: Isotoma setinornata Lee, B-H, 1977:164
            Syn.: Isotoma setinornata Yi, 1976 nn
             Syn.: Isotoma tarspinata(sic) Lesshafft, 1977 ISS
Desoria trispinata from the Azores
2010.09.15 © E.D.E.N.
Desoria trispinata from the UK
2010.04.22 © Shaw, P.
Det.: Fjellberg, A.

             Syn.: Desoria trispina(sic) Cruz-Leal, JI, Escalante-Jiménez, AL, Salgado-Ortiz, J et Palacios-Vargas, JG, 2013:1666 ISS
Desoria trispinata from Japan
2013.03.12 © Hara, Y.
Desoria trispinata from Japan
Abd.4-6
2013.03.12 © Hara, Y.
Desoria trispinata from Germany
2014.09.27 © Beck, K.
Desoria trispinata from Germany
2014.10.05 © Jäschke, J.
Isotoma (Desoria) trispinata from India
After Santeshwari, Raghuraman M & Singh J, 2015 p.574
Desoria trispinata from Holland
2016.02.11 © van Duinen, J.

Species truncata Waltz & Hart, 1995
Desoria truncata from the USA
2006 © Bernard, E.C.
Desoria truncata from the USA
2007.09.05 © Barnd., B

Species tshernovi Martynova, 1974
            Syn.: Isotoma nanseni Fjellberg, 1978
             Syn.: Isotoma tschernovi(sic) Fjellberg, 1995 ISS
Desoria tshernovi from Norway
2011 © Fjellberg, A.

Species tuckeri Christiansen & Bellinger, 1980
Species tunica Christiansen & Bellinger, 1980
Species uniens (Christiansen, K et Bellinger, P, 1980:760)
Desoria uniens from the USA
After Fjellberg A, 2010 Fig.115B

Species violacea (Tullberg, T, 1877:36), B
            Syn.: Isotoma neglecta Schäffer
            Syn.: Isotoma violacea var. mucronata Axelson, 1900
            Syn.: Isotoma olivacea Tuxen, 1944 partim
            Syn.: Isotoma trispinata Tuxen, 1944 partim
            Syn.: Isotoma inupikella Fjellberg, 1978
            Syn.: Desoria tolya Hågvar, S, 2010:283
Desoria violacea from the UK
Mucro with 4 teeth
2005.04.01 © Hopkin, S.P.
Desoria violacea (bar=2µm)
Rhombic epicuticular ultrastructure
After Nickerl J & al, 2012 Fig.4E
Desoria violacea from the UK
2014.01.19 © Valentine., B

Species yukinomi (Yosii, 1939) Yosii, R, 1977:157
Species zlotini Martynova, 1968

Genus Gnathisotoma Cassagnau, P, 1957:393
Species bicolor Cassagnau, P, 1957:393, t.t.
Species deharvengi Najt, 1981
Species pallens Yoshii, 1990
Species patagonica Yoshii, 1990
Species spinolabris Fjellberg, A, 2010:42
Gnathisotoma spinolabris from the USA
After Fjellberg A, 2010 Fig.115G
Gnathisotoma spinolabris from the USA
After Fjellberg A, 2010 Fig.114


Genus Parisotoma Bagnall, RS, 1940:171
  Syn.: Collembola Isotoma Bagnall, RS, 1940:163 partim
  Syn.: Isotomidae Proisotoma(sic) Greenslade, 2002 lapsus
Species agrelli (Delamare Deboutteville, 1950), B
Species amurica (Potapov, MB, 1991:282) Potapov, 2001
Species appressopilosa Potapov, MB, 1991:279
Species atroculata Potapov, MB, 1991:285
Species coeca Yosii, 1966
Species confusoculata Salmon, JT, 1944:150
Species dichaeta (Yosii, 1969)
Species dividua Salmon, JT, 1944:150
Species ekmani (Fjellberg, 1977) Fjellberg, A, 2007:47
            Syn.: Isotoma notabilis f. pallida Agrell, I, 1939:2
             Syn.: Isotoma ekmany(sic) Palacios-Vargas, JG in Sánchez-Ramos, G, Reyes-Castillo, P et Dirzo, R, 2005:365 ISS
             Syn.: Isotoma ekmany(sic) Palacios-Vargas, JG in Sánchez-Ramos, G, Reyes-Castillo, P et Dirzo, R, 2005:366,af ISS
Parisotoma ekmani from the UK
2012.11.18 © Lewington, A.
Det.: Shaw, P.

Species greensladeae Rusek, J, 1984:364
Species hyonosenensis (Yosii, 1939) Potapov, 2001
            Syn.: Desoria pseudosetispinosa Chen, 1985
            Syn.: Isotoma setispinosa Lee, B-H, 1977:162
             Syn.: Isotoma hyonosensensis(sic) Yosii, R, 1977:156 ISS
             Syn.: Parisotoma hyonosonensis(sic) Gao, Y, Xiong, Y et Potapov, MB, 2009:59 ISS
Species insularis Deharveng, 1981
Parisotoma insularis from Antarctica
After Davis, G. in Greenslade, P., 2006 Fig.69.

Species longa Potapov, MB, 1991:283
Species notabilis (Schäffer, 1896) Bagnall, RS, 1940:171, t.t., B
            Syn.: Isotoma menotabilis Börner, 1903
            Syn.: Isotoma delicatula Brown, 1929
            Syn.: Isotoma (Isotoma) eunotabilis Folsom J in Mills, HB, 1934:58
            Syn.: Desoria monticola Hao & Huang, 1995
             Syn.: Parisotoma nitabilis(sic) Boitaud, L, Salmon, S Bourlette, C et Ponge J-F, 2006:452 ISS
Parisotoma notabilis from the UK
2010.03.14 © Barton, T.
Parisotoma notabilis from Norway
2011 © Fjellberg, A.
Parisotoma notabilis from the UK
2012.03.21 © Nurcombe, E.
Parisotoma notabilis from the UK
2012.04.03 © Murray, A.
Parisotoma notabilis from the UK
2012.08.23 © Murray, A.
Parisotoma notabilis from the UK
2012.08.23 © Murray, A.
Parisotoma notabilis from Iran
After Kahrarian M & Arbea J, 2013 Fig.7
Parisotoma notabilis from the UK
With square eyepatch
2013.02.03 © Jones, A.
Parisotoma notabilis juv. from Canada
2013.10.22 © Hochhalter, R.
Parisotoma notabilis from Holland
With square eyepatch of 4 ocelli
2013.11.13 © van Duinen, J.
Parisotoma notabilis from the UK
With reticulate gray body pigmentation
2013.12.23 © Murray, A.
Parisotoma notabilis from Germany
2014.10.20 © Beck, K.
Parisotoma notabilis from the UK
2016.01.13 © Horton, M.
Parisotoma notabilis from Belgium
2017.04.20 © Huskens, M.-L.

Species obscurocellata Potapov, Janion & Deharveng, 2011
Species picea Salmon, JT, 1949:36
Species postantennala Salmon, JT, 1949:38
Species quinquedentata Salmon, JT, 1943:382
             Syn.: Parisotoma quinquedendata(sic) Salmon, JT, 1943:382 ISS
Species raffi (Womersley, 1934)
Species reducta Rusek, J, 1984:360
Species removeopthalma Martynova, 1977
Species sexsetosa Potapov, Janion & Deharveng, 2011
Species tariva (Wray, 1953) Palacios-Vargas, JG, Mejía-Recamier, BE & Cutz-Pool, LQ, 2009:205
Species terricola Rusek, J, 1984:357
Species travei Deharveng, 1981
Species trichaetosa Martynova, 1977
Species vtorovi Martynova, 1977

Genus Misturasotoma Bernard, EC et Christiansen, KA, 2010:58
Species brucealla (Wray, 1953) Bernard, EC & Christiansen, KA, 2010:58, t.t.
            Syn.: Isotoma brucei Wray, 1953, nec Carpenter, 1907
Misturasotoma brucealla from the USA
After Bernard EC & Christiansen KA, 2010 Fig.1


Genus Vertagopus Börner, C, 1906:171 key
Vertagopus? sp. from the Arctic region
After White A in Sutherland PC, 1852 Fig.8
Vertagopus sp. from Germany
2006 © Kohl, F.
Vertagopus sp. from the USA
2006.12.13 © Cresswell, S.
Vertagopus sp. from the UK
2007.11.10 © Valentine., B
Vertagopus arboreus from the UK
Legs with clavate tenent setae
2015.12.29 © Cunningham, T.

Species abeloosi Poinsot, 1965
Species alpinus Haybach, 1972
Vertagopus alpinus from Austria
After Schulz H-J, 2010 Abb.4
Vertagopus alpinus from Austria
Left eyepatch : G and H smaller
Modified after Schulz H-J, 2010 Abb.3

Species alpus Christiansen & Bellinger, 1980
Species arborata Fusheng, Huifen & Yueling, 1993
Species arboreus (Linnæus, C, 1758:609) Ågren, 1903, B
            Syn.: Podura nemotrosa Linnæus, C, 1748:67 i
            Syn.: Podura nigra pedibus albis Hill, 1752 i
            Syn.: Podura annularis de Fourcroy, AF, 1785:524
            Syn.: Podura oblonga nigra de Geer, C, 1778:17
            Syn.: Podura fuliginosa Templeton, R in Templeton, R et Westwood, JO, 1836:95
Podura fuliginosa from Ireland
After Templeton, R, 1836 Pl.XII Fig.3

            Syn.: Isotoma rubricauda Bourlet, C, 1839:402
            Syn.: Desoria (Deuxième division) cylindrica Nicolet, H, 1842:60
            Syn.: Desoria (Deuxième division) ebriosa Nicolet, 1842
            Syn.: Desoria pallida Nicolet, 1842
            Syn.: Isotoma viatica Lubbock, J, 1873:168
Isotoma viatica from the UK
After Lubbock, J., 1873 Pl.36.
 
 

            Syn.: Isotoma synonymica Mac Gillivray, 1896
            Syn.: Isotoma terminata Mac Gillivray, 1896
            Syn.: Isotoma denticulata Schäffer, 1896
"Kleine schwarze Insekten, welche unter den
Rinden alter Bäume sitzen, und mit dem Sterze
hüpfen" from Sweden
After de Geer, C., 1749 Tab.I Fig.1.

             Syn.: Podura Arborea(sic) Scopoli, JA, 1763:380 ISS
Podura arborea nigra from Sweden
After de Geer, C., 1778 Tom.VII Pl.2 Fig.2.
 
 


             Syn.: Vertagopus arborea(sic) Christiansen, 1958 ISS
             Syn.: Vertagopus arborea(sic) Snider, RJ, 1967:216 ISS
             Syn.: Vertagopus arborea(sic) Massoud & Rapoport, 1968 ISS
             Syn.: Vertagopus arborea(sic) Raynal, G, 1974:189 ISS
             Syn.: Vertagopus arborea(sic) Rubio, 1974 ISS
             Syn.: Vertagopus arborea(sic) Rusek, 1977 ISS
             Syn.: Vertagopus arborea(sic) Bödvarsson, H, 1982:63 ISS
             Syn.: Vertagopus arborea(sic) André, HM, 1983:274,B ISS
             Syn.: Vertagopus arborea(sic) Szeptycki & Weiner in Razowski, 1990 ISS
             Syn.: Vertagopus arborea(sic) Dallai & Malatesta in Calderini, 1995 ISS
Vertagopus arboreus from the USA
2006 © Bernard, E.
Vertagopus arboreus from the UK
2006 © Valentine., B
Vertagopus arboreus from the UK
2006 © Barton, T.
Vertagopus arboreus from the UK
2007 © Valentine., B
Vertagopus arboreus from the UK
2007 © Barton, T.
Vertagopus arboreus from Germany
2007 © Peters, J.
Vertagopus arboreus from the UK
2007 © Barton, T.
Vertagopus arboreus from Germany
2007.02.15 © Peters, J.
Vertagopus arboreus from the USA
2007.03.10 © Murray, T.
Vertagopus arboreus from the USA, juvenile
2007.03.13 © Murray, T.
Vertagopus arboreus from the USA
2007.03.14 © Murray, T.
Vertagopus arboreus from the USA
2007.11.04 © Moorehead, C.
Vertagopus arboreus from the UK
2007.12.30 © Valentine., B
Vertagopus arboreus from the UK
2008.01.04 © Oakshott, A.
Vertagopus arboreus from the UK
2008.01.12 © Farrell, S.
Vertagopus arboreus from the UK
2008.01.12 © Tonsbeek, M.
Vertagopus arboreus from Germany
2008.01.14 © Peters, J.
Vertagopus arboreus from the UK
2008.02.20 © Robertson, A.
Vertagopus arboreus from the UK
2008.02.21 © Valentine., B
Vertagopus arboreus from the UK
2008.03.25 © Robertson, A.
Vertagopus arboreus from the UK
2008.04.01 © Valentine., B
Vertagopus arboreus from the UK
2009.01.12 © Valentine., B
Vertagopus arboreus from the USA
2009.02.15 © Murray, T.
Vertagopus arboreus from the UK
2009.12.10 © Valentine., B
Vertagopus arboreus from Holland
2011.05.01 © van Duinen, J.
Vertagopus arboreus juv. from the UK
2012.11.01 © Murray, A.
Vertagopus arboreus juv. from the UK
2012.11.01 © Murray, A.
Vertagopus arboreus juv. from the UK
2012.11.01 © Murray, A.
Vertagopus arboreus from the UK
2012.12.13 © Valentine., B
Vertagopus arboreus from Holland
Dense cover of short setae
2012.12.16 © van Duinen, J.
Vertagopus arboreus juv. from the UK
2012.12.25 © Murray, A.
Vertagopus arboreus from the UK
Dense cover of short setae
2013.01.04 © Phillips, E.
Vertagopus arboreus juv. & adult from Germany
2014.02.05 © Beck, K.
Vertagopus arboreus subadult from the UK
2015.12.04 © Barton, T.
Vertagopus arboreus from the UK
Eyepatch with 6 ocelli with crystal + 2 ocelli without crystal
2015.12.29 © Cunningham, T.
Vertagopus arboreus from Belgium
2015.12.31 © Janssens, F.
Vertagopus arboreus from Belgium
Ready to jump
2016.11.01 © Huskens, M.-L.
Vertagopus arboreus from Belgium
Knobbed tenent setae (*)
2016.11.01 © Huskens, M.-L.
Vertagopus arboreus from Holland
Adult with early instar juvenile
2016.12.28 © van Duinen, J.
Vertagopus arboreus from Holland
Tibiotarsus with clavate tenent seta (*)
2016.12.28 © van Duinen, J.

Species arcticus Martynova, 1969
Species asiaticus Potapov, M, Gulgenova, A et Babykina, M, 2016:115
Vertagopus asiaticus from Buryatia
After Potapov M, Gulgenova A & Babykina M, 2016 Fig.27

            Syn.: Vertagopus arboreus Uchida, 1969 =?
            Syn.: Vertagopus arboreus Martynova, 1979 =?
Species beta Christiansen & Bellinger, 1980
Species brevicaudus (Carpenter, 1900) Fjellberg, A, 1996:100
Species ceratus Potapov, M, Gulgenova, A et Babykina, M, 2016:113
Vertagopus ceratus from Buryatia Barguzin Range
After Potapov M, Gulgenova A & Babykina M, 2016 Fig.25
Vertagopus ceratus from Buryatia Balan-Tamur Range
After Potapov M, Gulgenova A & Babykina M, 2016 Fig.26

Species ciliatus Christiansen, K, 1958:75
Vertagopus ciliatus from Lebanon
After Christiansen K, 1958 Pl.9 Fig.22

Species cinereus (Nicolet, H, 1842:60) Marlier, GJ, 1942:9, t.t., B
            Syn.: Isotoma quadridenticulata Tullberg, 1871
            Syn.: Isotoma lateraria MacGillivray, 1896
             Syn.: Isotoma latraria(sic) Mills, HB, 1934:57 ISS
            Syn.: Isotoma clavigera Krausbauer, 1901
             Syn.: Vertagopus cinerea(sic) Marlier, GJ, 1942:9,B ISS
             Syn.: Vertagopus cinerea(sic) Snider, RJ, 1967:216 ISS
             Syn.: Vertagopus cinerea(sic) Rusek, 1977 ISS
             Syn.: Vertagopus cinerea(sic) Szeptycki & Weiner in Razowski, 1990 ISS
             Syn.: Vertagopus cinerea(sic) Dallai & Malatesta in Calderini, 1995 ISS
Vertagopus cinereus
After Potapov M, 2001 Fig.1
Vertagopus cinereus ♀
After Potapov M, 2001 Fig.2
Vertagopus cinereus from the UK
2008.09.21 © Robertson, A.
Vertagopus cinereus from Holland
2009.04.12 © Baas, A.H.
Vertagopus cinereus from Holland
2009.04.12 © Baas, A.H.
Vertagopus cinereus from Holland
2009.04.12 © Baas, A.H.
Vertagopus cinereus from Holland
2009.04.12 © Baas, A.H.
Vertagopus cinereus from Holland
2009.04.18 © Baas, A.H.
Vertagopus cinereus from Holland
2009.04.21 © Baas, A.H.
Vertagopus cinereus from Holland
2009.04.21 © Baas, A.H.
Vertagopus cinereus from the UK
2010.11.14 © Robertson, A.
Vertagopus cinereus from France
2013.04.13 © Sandra.
Vertagopus cinereus from Holland
2013.12.11 © Baas, A.H.
Vertagopus cinereus juv. from Germany
2014.02.14 © Beck, K.
Vertagopus cinereus juv. from Holland
2014.11.07 © van Duinen, J.
Vertagopus cinereus juv. from Holland
2014.11.07 © van Duinen, J.
Vertagopus cinereus from the UK
2015.01.01 © Barton, T.

Species haagvari Fjellberg, A, 1996:100
Species helveticus Haybach, 1980
Species laricis Martynova, 1975
Vertagopus laricis from Buryatia
After Potapov M, Gulgenova A & Babykina M, 2016 Fig.23
Vertagopus laricis from Buryatia
After Potapov M, Gulgenova A & Babykina M, 2016 Fig.28

Species monta (Christiansen & Bellinger, 1980) Fjellberg, A, 1996:100,cf
Species montanus Stach, 1947
Species obscurus Wahlgren, 1906
Species pallidus Martynova, 1974
Species persea Wray, 1952
Species pseudocinereus Fjellberg, 1975
            Syn.: Isotoma dilatata Mac Gillivray, 1896 =?
            Syn.: Isotoma inclinata Mac Gillivray, 1896 =?
            Syn.: Isotoma lateraria Mac Gillivray, 1896 =?
            Syn.: Isotoma unica Mac Gillivray, 1896 =?
Vertagopus pseudocinereus
Courtship behaviour
After Goloschapova & al, 2006 Fig.1
Vertagopus pseudocinereus from Holland
2014.01.07 © van Duinen, J.
Vertagopus pseudocinereus from Holland
2014.02.22 © van Duinen, J.
Vertagopus pseudocinereus from Holland
With short dorsal macrosetae on each trunk segment
2014.02.22 © van Duinen, J.
Vertagopus pseudocinereus from Holland
2014.02.26 © van Duinen, J.
Vertagopus pseudocinereus from Holland
2014.02.26 © van Duinen, J.
Vertagopus pseudocinereus ♂ from the USA
With bent 2nd antennal segments
2014.06.27 © Curtis, R.
Vertagopus pseudocinereus from Germany
2014.10.19 © Beck, K.
Vertagopus pseudocinereus from Germany
2014.10.20 © Beck, K.
Vertagopus pseudocinereus from Germany
2014.11.02 © Beck, K.
Vertagopus pseudocinereus from the USA
2014.12.23 © Montgomery, G.
Vertagopus pseudocinereus from the USA
2015.03.09 © Eiseman, C.
Vertagopus pseudocinereus from the USA
Head dorsal pattern
2015.03.09 © Eiseman, C.
Vertagopus pseudocinereus from the USA
Pale colour form
2015.03.09 © Eiseman, C.
Vertagopus pseudocinereus from the USA
Elliptic postantennal organ (PAO)
2015.03.09 © Eiseman, C.
ertagopus pseudocinereus from Canada
2015.11.09 © Hogue, A.

Species reuteri (Schött, H, 1893:71) Fjellberg
Species sarekensis (Wahlgren, 1906) Fjellberg, A, 1996:100
Vertagopus sarekensis from Scandinavia
After Fjellberg, A, 2007 Fig.61O

Species sp.1 Fjellberg, A, 1996:100
Species sp.2 Fjellberg, A, 1996:100
Species tianschanicus Martynova, 1969
Species westerlundi (Reuter, 1898) Schubert, 1933
            Syn.: Isotoma brevicauda Carpenter, 1900
            Syn.: Isotoma sarekensis Wahlgren, 1906
Vertagopus westerlundi from Italy
2006 © Paolucci, P.
Vertagopus westerlundi from Buryatia
After Potapov M, Gulgenova A & Babykina M, 2016 Fig.24


Genus Paracerura Deharveng, L et de Oliveira, EP, 1994:442 key
Species airesi de Mendonça, Abrantes & Fernandes, 2009
Species cristinae Abrantes, EA et Duarte, M, 2013:1394
Paracerura cristinae from Brazil
After Abrantes EA & Duarte M, 2013 Fig.8

Species gandarela de Mendonça & da Silveira, 2013
Species itatiaiensis (Arlé, R, 1960:167) Deharveng, L & de Oliveira, EP, 1994:443
Species pallida Abrantes, EA et Duarte, M, 2013:1392
Paracerura pallida from Brazil
After Abrantes EA & Duarte M, 2013 Fig.1

Species paulista Abrantes, EA et Duarte, M, 2013:1397
Paracerura paulista from Brazil
After Abrantes EA & Duarte M, 2013 Fig.15

Species pindorama Queiroz & de Mendonça, 2010
Species serrana de Mendonça, Abrantes & Fernandes, 2009
Species virgata Deharveng, L et de Oliveira, EP, 1994:443, t.t.

Genus Marisotoma Fjellberg, A, 1997:26
Species canaliculata Fjellberg, A, 1997:26, t.t.
Species iremeli (Potapov, MB, 1991:278) Potapov, 2001
Species tenuicornis (Axelson, WM, 1903:10) Bulimar & Hutu, 1984, B

Genus Araucanocyrtus Massoud & Rapoport, 1968
Species fileri Massoud & Rapoport, 1968, t.t.

Genus Psammisotoma Greenslade, P et Deharveng, L, 1986:89 key
Species sp.1 Thibaud, J-M et Palacios-Vargas, JG, 1999:31
Species dispar (Christiansen, K et Bellinger, P, 1988:233)
Species galapagoensis Thibaud, J-M, Najt, J et Jaquemart (sic), S (+), 1994:202
Species kingae Greenslade, P et Deharveng, L, 1986:90, t.t.
Species mariagalanteae Thibaud, 1993
Species restingae Abrantes, EA et de Mendonça, MC, 2009:28
Psammisotoma restingae from Brazil
After Abrantes EA & de Mendonça MC, 2009 Fig.13


Genus Aackia Yosii, 1966
Species karakoramensis Yosii, 1966, t.t.

Genus Antarcticinella Salmon, 1965
Species monoculata Salmon, 1965

Genus Cheirotoma Bagnall, 1949
Species spatulata (Chamberlain, 1943) Bagnall, 1949

Genus Halisotoma Bagnall, 1949 key
Halisotoma sp. from the UK
2016.09.23 © Trewhella, S.
Det.: Fjellberg, A.

Species arenicola Fjellberg, A, 2009:135
Species boneti (Delamare Deboutteville, 1954)
Species kainui (Christiansen, K et Bellinger, P, 1992:164)
Species marisca (Christiansen & Bellinger, 1988:235)
Species maritima (Tullberg, 1871 tt) Bagnall, 1949
            Syn.: Isotoma pacifica Yosii, 1971
            Syn.: Isotoma pacifica Yosii, R, 1977:156
Species poseidonis (Bagnall, 1939)
             Syn.: Halisotoma poseidons(sic) Fjellberg, A, 2009:138 ISS
Species pritchardi (Womersley, H, 1936:478) Bagnall, 1949
            Syn.: Isotoma maritima Womersley, H, 1936:321
Species sindentata (Salmon, JT, 1943:382)
Parisotoma sindentata from New Zealand
After Salmon JT, 1943 Pl.41 Fig.38


Genus Metisotoma Maynard, 1951
Species grandiceps (Reuter, 1891) Cassagnau, 1972
            Syn.: Isotoma macnamarai Folsom, 1918
            Syn.: Metisotoma capitona Maynard, EA, 1951:95
            Syn.: Metisotoma spiniseta Maynard, EA, 1951:94
Isotoma grandiceps from Siberia
After Schött H, 1893 Taf.VI Fig.24
Isotoma grandiceps from Siberia
After Schött H, 1893 Taf.VI Fig.23
Metisotoma grandiceps from the USA
2007.02.22 © Bernard, E.C.
Metisotoma grandiceps from the USA
2007.02.22 © Bernard, E.C.
Metisotoma grandiceps from the Canada
2016.03.12 © Schmidt, H.
Metisotoma grandiceps from the Canada
Wide head; short antennae
2016.03.12 © Schmidt, H.

Species ursi (Yosii, R, 1972:90)

Genus Myopia Christiansen, K et Bellinger, P, 1980:768
Species alaskana (Christiansen, K & Bellinger, P, 1980:769) Fjellberg, A, 2010:28, t.t.
Myopia alaskana from Alaska
2007.06.12 © Takeuchi, N.
Myopia alaskana gr. from the USA
2008.12.23 © Fjellberg, A.
Myopia alaskana from the USA
After Fjellberg A, 2010 Fig.112
Myopia alaskana from the USA
After Fjellberg A, 2010 Fig.115A


Genus Najtia Arlé & Mendonça, 1986
Species vicaria (Arlé, R, 1960:166) Arlé & Mendonça, 1986

Genus Pentacanthella Deharveng, 1979
Species decemoculata Deharveng, 1979

Genus Pseudosorensia de Izarra, DC, 1972:77
  Syn.: Isotoma Pseudosorencia(sic) Thibaud, J-M et Palacios-Vargas, JG, 1999:25 ISS
Species atlantica (Wise, KAJ, 1970:197)
Species fueguensis de Izarra, DC, 1972:77, t.t.
Species sp. Thibaud, J-M et Palacios-Vargas, JG, 1999:29
Species subcaeca Deharveng, 1981

Genus Azoritoma Greenslade, P et Potapov, M, 2008:89
Species macquariensis Greenslade, P et Potapov, M, 2008:90, t.t.

Genus Sericeotoma Potapov, MB, 1991:287
Species knissi Potapov, MB, 1991:288

Genus Mucronia Fjellberg, A, 2010:43
Species enigmatica Fjellberg, A, 2010:43, t.t.
Mucronia enigmatica from the USA
After Fjellberg A, 2010 Fig.93

Species fjellbergi Potapov, M et Babenko, A, 2014:1
Mucronia fjellbergi from Russia
After Potapov M & Babenko A, 2014 Fig.3B


Genus Rhyniella Hirst & Maulik, 1926, -
Species praecursor Hirst & Maulik, 1926 -
Rhyniella praecursor from the UK Rhynie Chert
After Hirst S & Maulik S, 1926 Fig.A
Holotype as fixed by Tillyard, 1928
Rhyniella praecursor from the UK Rhynie Chert
After Scourfield DJ, 1940


Genus Protoisotoma Christiansen, KA et Pike, E, 2002:171, -
Species burma Christiansen, KA et Nascimbene, P, 2006:343, -
Species micromucra Christiansen, KA et Pike, E, 2002:171, -



References