http://www.collembola.org/taxa/katiidae.htm - Last updated on 2024.04.25 by Frans Janssens
Checklist of the Collembola: Katiannidae

Familia Katiannidae Börner, 1913:321 sensu Bretfeld G, 1999:13 key

Genus Number
of species
Arborianna1
Betschurinus1
Borgesminthurinus1
Cretokatianna1
Dalianus1
Gen.nov.12
Gen.nov.21
Gisinianus1
Katianna51
Katiannellina1
Katiannina3
Keratopygos1
Neokatianna1
Papirinus4
Parakatianna10
Polykatianna11
Pseudokatianna12
Rusekianna5
Sminthurinus98
Stenognathellus4
Vesicephalus8
Zebulonia1
Katiannidae219
 Genera with number of species
  Syn.: ... Smynthuridae(sic) Lubbock, 1862, partim Lubbock, 1870
  Syn.: ... Sminthuridinae Börner, 1906, partim Börner, 1913
  Syn.: Katianniformia Katiannida(sic) Bretfeld, 1986
  Syn.: Katianniformia Katiannida(sic) Bretfeld, G, 1999:98


Two new Katiannidae from the UK
2009.06.06 © Ardron, P.A.
Katiannina macgillivrayi from the USA
2009.11.26 © Justis, S.


Genus Cretokatianna Sánchez-García, A et Engel, MS, 2016:7, - F

Species bucculenta Sánchez-García, A et Engel, MS, 2016:7, -

Cretokatianna bucculenta in Cretaceous amber from Spain
After Sánchez-García A & Engel MS, 2016 Fig.4

            Syn.: Fasciosminthurus? sp. Simón-Benito, Ortuño & Espantaleón, 2002

Tribus Katiannini Börner, 1913




Genus Gen.nov.1 Greenslade P in Ardron, PA, 2009:16

Genus nov.1 sp. nov. from the UK
Third antennal segment with papilla and long fine setae
2009.04.01 © Ardron, P.
Det. Greenslade, P.
Genus nov.1 sp. nov. from the UK
2009.11.15 © Barton, T.
Genus nov.1 sp. nov. from the UK
Third tibiotarsus distally with spatulate setae (*)
2015.11.23 © Phillips, E.

Species sp.nov.1 Greenslade P in Ardron, PA, 2009:16

Genus nov.1 sp. nov. from the UK
After Ardron PA, 2009 p.16

  Syn.: Gen.nov.1 sp.nov.1 f. nigra Janssens F & Phillips E, 2016
Remark by Janssens, F. 2017.02.02: Dark form. Anterior middorsal pale narrow stripe. Dark 6th abdominal segment.

Genus nov.1 sp. nov. f. nigra from the UK
Form with dark abd.6
2016.12.14 © Phillips, E.
Genus nov.1 sp. nov. f. nigra from the UK
Form with dark abd.6
2016.12.14 © Phillips, E.
Genus nov.1 sp. nov. f. nigra from the UK
Form with dark abd.6
2016.12.14 © Phillips, E.
Genus nov.1 sp. nov. f. nigra from the UK
Form with dark abd.6
2017.02.01 © Barton, T.




"Sphyrotheca multifasciata" from the UK
2007.03.04 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.18 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.18 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.18 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.18 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.18 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.20 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.20 © Cornwall, N.J.
"Parakatianna" sp. from the UK
2009.02.23 © Barton, T.
"Parakatianna" sp. from the UK
2009.02.23 © Barton, T.
"Parakatianna" sp. from the UK
2009.02.23 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Polykatianna" sp. ♀ from the UK
Sub-anal appendage
2009.03.27 © Ardron, P.
"Polykatianna" sp. from the UK
2009.04.01 © Ardron, P.
"Polykatianna" sp. from the UK
2009.04.01 © Ardron, P.
"Polykatianna" sp. ♂ from the UK
2009.04.01 © Ardron, P.
"Polykatianna" sp. from the UK
2009.04.01 © Ardron, P.
"Polykatianna" sp. from the UK
2009.04.01 © Ardron, P.
"Polykatianna" sp. from the UK
2009.04.01 © Ardron, P.
"Parakatianna" sp. from the UK
2009.04.21 © Ardron, P.
"Parakatianna" sp. from the UK
2009.04.21 © Ardron, P.
Genus nov.1 sp. nov. from the UK
Mucrones
2009.05.20 © Janssens, F.
Genus nov.1 sp. nov. ♀ from the UK
Female sub-anal appendage
2009.05.20 © Janssens, F.
Genus nov.1 sp. nov. from the UK
2009.11.15 © Barton, T.
Genus nov.1 sp. nov. from the UK
2009.11.21 © Barton, T.
Genus nov.1 sp. nov. ♀ from the UK
Sub-anal appendage
2009.12.11 © Barton, T.
Genus nov.1 sp. nov. from the UK
2009.11.28 © Barton, T.
Genus nov.1 sp. nov. from the UK
2009.11.28 © Barton, T.
Genus nov.1 sp. nov. from the UK
Preparing for jump step 1 : lifting prolegs
2009.12.11 © Barton, T.
Genus nov.1 sp. nov. from the UK
Spermatophore
2009.12.12 © Barton, T.
Genus nov.1 sp. nov. from the UK
Preparing for a jump
2009.12.12 © Barton, T.
Genus nov.1 sp. nov. from the UK
Consumed spermatophore
2009.12.12 © Barton, T.
Genus nov.1 sp. nov. from the UK
2009.12.17 © Jones, A.
Genus nov.1 sp. nov. from the UK
2009.12.20 © Jones, A.
Genus nov.1 sp. nov. from the UK
Egg with polar caps
2009.12.20 © Barton, T.
Genus nov.1 sp. nov. from the UK
2009.12.24 © Barton, T.
Remark by Janssens, F. 2010.02.14: Adult form. Thoracic middorsal pale narrow stripe. Pale 6th abdominal tergum.

Genus nov.1 sp. nov. ♂ from the UK
Furca
2015.03.29 © Phillips, E.
Genus nov.1 sp. nov. from the UK
2010.02.13 © Barton, T.
Genus nov.1 sp. nov. from the UK
2012.01.06 © Nurcombe, E.
Genus nov.1 sp. nov. from the UK
Abd.5 with 2 distinct bothriothricha
Abd.6 without pigment
2012.01.06 © Nurcombe, E.
Genus nov.1 sp. nov. from the UK
2012.01.06 © Nurcombe, E.
Genus nov.1 sp. nov. from the UK
2012.12.15 © Barton, T.
Genus nov.1 sp. nov. from the UK
Eyes
2012.02.16 © Nurcombe, E.
Genus nov.1 sp. nov. from the UK
2014.12.14 © Garlick, T.
Genus nov.1 sp. nov. from the UK
2014.12.18 © Jeffery, J.
Genus nov.1 sp. nov. from the UK
2014.12.21 © Robertson, A.
Genus nov.1 sp. nov. from the UK
2015.12.09 © Boardman, P.
Genus nov.1 sp. nov. from the UK
Footcomplex with knobbed tenent setae (*)
2016.12.01 © Phillips, E.
Genus nov.1 sp. nov. from the UK
Bothriotricha D (*)
2016.12.01 © Phillips, E.
Genus nov.1 sp. nov. ♀ from the UK
Sub-anal appendage (SAA)
2017.01.04 © Phillips, E.
Genus nov.1 sp. nov. ♀ from the UK
Curved sub-anal appendage (SAA) (*)
2017.01.29 © David W.
Genus nov.1 sp. nov. ♀ from the UK
Footcomplex with knobbed tenent setae
2017.02.04 © David W.
Genus nov.1 sp. nov. ♀ from the UK
Curved sub-anal appendage (SAA) (*)
2017.02.04 © David W.
Genus nov.1 sp. nov. ♀ from the UK
With everted collophore vesicles
Ready to jump
2019.11.15 © Barton, T.
Genus nov.1 sp. nov. from the UK
2021.12.11 © Phillips, E.
Genus nov.1 sp. nov. from the UK
Lateral bothriotricha ABC
2021.12.11 © Phillips, E.
Genus nov.1 sp. nov. from the UK
Abd.5 with bothriotrichum D (reddish)
Abd.6 pale
2021.12.11 © Phillips, E.
Genus nov.1 sp. nov. ♂ from the UK
Abd.5 with genital papilla (*)
2022.01.25 © Phillips, E.
Genus nov.1 sp. nov. from the UK
With everted collophore vesicles
Ready to jump
2023.11.16 © Barton, T.
Genus nov.1 sp. nov. from the UK
Ready to jump
Collophore vesicles everted
2023.12.30 © Phillips, E.



Species sp.nov.2 Janssens F & Phillips E, 2017

"Parakatianna" sp. from the UK
2009.04.21 © Ardron, P.
"Parakatianna" sp. from the UK
2009.04.21 © Ardron, P.
"Parakatianna" sp. from the UK
2009.04.21 © Ardron, P.
Genus nov.1 sp. nov. from the UK
Preparing for jump step 3 : everting collophore vesicles
2009.12.11 © Barton, T.
Genus nov.1 sp. nov. from the UK
Everting collophore vesicles before jumping
2016.01.14 © Horton, M.
Genus nov.1 sp. nov. from the UK
Backwards extended furca
2016.01.14 © Horton, M.
Genus nov.1 sp. nov. from the UK
Preparing for jump step 3 : everting collophore vesicles
2020.10.28 © Barton, T.
Genus nov.1 sp. nov. from the UK
Everting collophore vesicles before jump
2021.02.05 © Barton, T.

  Forma juvenile1 Janssens F & Barton T, 2009

"Parakatianna" sp.from the UK
2009.02.23 © Barton, T.
Genus nov.1 sp. nov. from the UK
2009.11.15 © Barton, T.
Genus nov.1 sp. nov. from the UK
2009.12.11 © Barton, T.
Genus nov.1 sp. nov. from the UK
2010.03.09 © Barton, T.
Genus nov.1 sp. nov. from the UK
2013.10.13 © Nurcombe, E.

  Forma juvenile2 Janssens F & Cornwall N, 2007

"Sphyrotheca multifasciata" juv. from the UK
2007.03.18 © Cornwall, N.J.
"Parakatianna" sp. from the UK
2009.02.23 © Barton, T.
"Katianna" sp. from the UK
2009.03.06 © Barton, T.
"Polykatianna" sp. from the UK
2009.04.07 © Shaw, P.
"Parakatianna" sp. from the UK
2009.04.21 © Ardron, P.
Genus nov.1 sp. nov.2 from the UK
2009.10.25 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.11 © Barton, T.

  Forma subadult Janssens F & Cornwall N, 2007

"Sphyrotheca multifasciata" from the UK
2007.03.04 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.18 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.18 © Cornwall, N.J.
"Parakatianna" sp. from the UK
2009.02.23 © Barton, T.
"Parakatianna" sp. from the UK
2009.02.23 © Barton, T.
"Parakatianna" sp. from the UK
2009.02.23 © Barton, T.
"Parakatianna" sp. from the UK
2009.02.23 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.15 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.11.02 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.11.15 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2014.01.21 © Horton, M.
Genus nov.1 sp. nov.2 from the UK
Papilla on 3rd antennal segment (*)
2014.01.21 © Horton, M.
Genus nov.1 sp. nov.2 from the UK
Feeding instar with dark gut contents
2014.01.21 © Horton, M.
Genus nov.1 sp. nov.2 from the UK
With everted collophore vesicles
Ready to jump
2023.11.16 © Barton, T.

  Forma adult

Remark by Janssens, F. 2010.02.14: Adult form. Thoracic and abdominal middorsal pale or orange broad area. Pigmented (dark or reddish) 6th abdominal tergum with 2 pale spots.

"Sphyrotheca multifasciata" from the UK
2007.03.18 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2008.01.20 © Spears, G.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
Form with middorsal pale winged pattern
2009.04.21 © Ardron, P.
"Parakatianna" sp. from the UK
Form with middorsal pale winged pattern
2009.04.21 © Ardron, P.
Genus nov.1 sp. nov.2 from the UK
2009.11.15 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.11.15 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.11.15 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.11.21 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.11.28 © Barton, T.
Genus nov.1 sp. nov.2 pale & dark form from the UK
2009.11.28 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
Grooming bubble blowing
2009.12.11 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.12 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.15 © Jones, A.
Genus nov.1 sp. nov.2 from the UK
2009.12.20 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.24 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.24 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.28 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.28 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.28 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.28 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2010.01.17 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2010.01.17 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2010.01.24 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2010.02.06 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2010.02.13 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2010.02.13 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
With everted collophore vesicles, ready to take-off
2010.02.21 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2012.01.06 © Nurcombe, E.
Genus nov.1 sp. nov.2 from the UK
2012.01.27 © Nurcombe, E.
Genus nov.1 sp. nov.2 ♂ from the UK
Raised genital papilla
2012.01.30 © Nurcombe, E.
Genus nov.1 sp. nov.2 from Ireland
2012.02.19 © Cluskey, D.
Genus nov.1 sp. nov.2 from Ireland
2013.02.02 © Cluskey, D.
Genus nov.1 sp. nov.2 from the UK
Composed eye with 8 ocelli: B,C,H large, A,E,F,G smaller, D smallest
2013.02.07 © Nurcombe, E.
Genus nov.1 sp. nov.2 from the UK
2013.12.17 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2014.11.26 © Nurcombe, E.
Genus nov.1 sp. nov.2 from the UK
2014.12.14 © Garlick, T.
Genus nov.1 sp. nov.2 ♂ from the UK
2015.12.09 © Boardman, P.
Genus nov.1 sp. nov.2 ♂ from the UK
Raised genital papilla (*)
2016.03.11 © Phillips, E.
Genus nov.1 sp. nov.2 ♀ from the UK
Curved sub-anal appendage on basal papilla
2017.02.09 © Phillips, E.
Genus nov.1 sp. nov.2 ♀ from the UK
Broad sub-anal appendage on basal papilla
2017.02.09 © Phillips, E.
Genus nov.1 sp. nov.2 ♂ from the UK
Raised genital papilla (*)
2017.11.16 © Phillips, E.
Genus nov.1 sp. nov.2 ♀ from the UK
2021.01.25 © Uncle, F.
Genus nov.1 sp. nov.2 ♀ from the UK
2021.01.25 © Uncle, F.
Genus nov.1 sp. nov.2 ♀ from the UK
Female gonopore
Sub-anal appendage (saa) on papilla
2021.01.25 © Uncle, F.
Genus nov.1 sp. nov.2 ♀ from the UK
Sub-anal appendage exteriad-anteriad
2021.01.25 © Uncle, F.
?Genus nov.1 sp. nov.2? from Australia
2021.04.15 © Hobern, D.
Genus nov.1 sp. nov.2 from the UK
Prejump posture
With everted collophore vesicles
2023.12.08 © Barton, T.


Genus Gen.nov.2 Greenslade P, 2012:il

Species sp.nov. Greenslade P, 2012:il

            Syn.: Katianna sp.3 Janssens F et Ardron PA in Ardron, PA, 2009:15
Katianna sp.3 from the UK
After Ardron PA, 2009 p.15

            Syn.: Sphyrotheca multifasciata Ardron, PA, 2009:15
            Syn.: Katianna sp.3 f. flammea Ardron, PA, 2009:16
Genus nov.2 sp. nov. f. flammea from the UK
2009.06.06 © Ardron, P.A.
Genus nov.2 sp. nov. f. flammea from the UK
2009.06.06 © Ardron, P.A.
Genus nov.2 sp. nov. f. flammea from the UK
2009.06.06 © Ardron, P.A.
Genus nov.2 sp. nov. f. flammea from the UK
2009.07.24 © Barton, T.
Genus nov.2 sp. nov. f . flammea from the UK
2014.11.12 © Nurcombe, E.
Genus nov.2 sp. nov. f . flammea from the UK
2014.11.12 © Nurcombe, E.
Genus nov.2 sp. nov. f . flammea from the UK
2015.01.07 © Jeffery, J.
Genus nov.2 sp. nov. f . flammea from the UK
2017.10.21 © Cunningham, T.

  Syn.: Gen.nov.2 sp.nov. f. A Janssens F & Greenslade P, 2012
Remark by Janssens, F. 2017.02.08: Form with pale flanks and middorsal stripe.
Genus nov.2 sp. nov. from the UK
2007 © Barton, T.
Genus nov.2 sp. nov. from the UK
2007.03.04 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.03.18 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.03.18 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.12.05 © Barton, T.
Genus nov.2 sp. nov. from the UK
2007.12.10 © Barton, T.
Genus nov.2 sp. nov. from the UK
2008.08.16 © Ardron, P.A.
Genus nov.2 sp. nov. from the UK
2008.08.16 © Ardron, P.A.
Genus nov.2 sp. nov. from the UK
2008.08.16 © Ardron, P.A.
Spines on head
Genus nov.2 sp. nov. from the UK
2009.04.21 © Ardron, P.A.
Genus nov.2 sp. nov. from Tasmania
2009.08.16 © Daley, A.
Genus nov.2 sp. nov. from the UK
2013.03.21 © Nurcombe, E.
Genus nov.2 sp. nov. from the UK
2013.12.10 © Nurcombe, E.
Genus nov.2 sp. nov. from the UK
2013.12.10 © Nurcombe, E.
Genus nov.2 sp. nov. ♂ from the UK
2014.11.29 © Nurcombe, E.
Genus nov.2 sp. nov. from France
2017.02.05 © Picard, J.
Genus nov.2 sp. nov. from the UK
2017.02.02 © David W.
Genus nov.2 sp. nov. from the UK
2017.02.02 © David W.
Genus nov.2 sp. nov. from the UK
2017.02.17 © David W.
Genus nov.2 sp. nov. from the UK
2017.02.17 © David W.
Genus nov.2 sp. nov. from the UK
2018.10.31 © Marquis, A.
Genus nov.2 sp. nov. from Tasmania
2020.03.05 © Ellingsen, K.
Genus nov.2 sp. nov. from Tasmania
Ventral aspect
2020.03.05 © Ellingsen, K.
Genus nov.2 sp. nov. from Tasmania
2020.03.05 © Ellingsen, K.

  Syn.: Gen.nov.2 sp.nov. f. B Janssens F & Greenslade P, 2012
Remark by Janssens, F. 2017.02.08: Form with dark flanks and middorsal stripe.
Genus nov.2 sp. nov. from the UK
2006 © Barton, T.
Genus nov.2 sp. nov. from the UK
2006 © Barton, T.
Genus nov.2 sp. nov. from the UK
2007.03.04 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.03.04 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.03.18 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.03.18 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.03.18 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.03.18 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.12.05 © Barton, T.
Genus nov.2 sp. nov. from the UK
2007.12.16 © Barton, T.
Genus nov.2 sp. nov. from the UK
2007.12.16 © Barton, T.
Genus nov.2 sp. nov. from Tasmania
2009.10.24 © Daley, A.
Genus nov.2 sp. nov. from Tasmania
2009.10.24 © Daley, A.
Genus nov.2 sp. nov. from Tasmania
2009.10.24 © Daley, A.
Genus nov.2 sp. nov. from the UK
2011.12.07 © Kent, J.J.
Genus nov.2 sp. nov. from the UK
2013.12.10 © Nurcombe, E.
Genus nov.2 sp. nov. from the UK
Eyebrow short and anteriorly pointed
2016.12.17 © David W.
Genus nov.2 sp. nov. from France
2016.12.23 © Picard, J.
Genus nov.2 sp. nov. from France
2016.12.23 © Picard, J.
Genus nov.2 sp. nov. from France
2016.12.24 © Picard, J.
Genus nov.2 sp. nov. ♀ from the UK
2017.02.17 © David W.
Genus nov.2 sp. nov. ♀ from the UK
Sub-anal appendage (saa) on papilla
2017.02.17 © David W.
Genus nov.2 sp. nov. ♀ from the UK
Bothriotrichum D
2017.02.17 © David W.
Genus nov.2 sp. nov. from France
2017.03.08 © Picard, J.
Genus nov.2 sp. nov. ♀ from France
Sub-anal appendages (*) on papilla
2017.03.11 © Picard, J.
Genus nov.2 sp. nov. ♀ & ♂from Tasmania
2020.03.05 © Ellingsen, K.
Genus nov.2 sp. nov. ♀ from the UK
2020.10.11 © Marquis, A.
Genus nov.2 sp. nov. ♀ from the UK
2020.10.11 © Marquis, A.

  Syn.: Gen.nov.2 sp.nov. f. obscura Janssens F & Jeffery J, 2015
Genus nov.2 sp. nov. f. obscura from the UK
2015.11.05 © Jeffery, J.
Genus nov.2 sp. nov. f. obscura? from Australia
2016.07.10 © Steve & Alison.
Genus nov.2 sp. nov. f. obscura? from Australia
2016.07.10 © Steve & Alison.
Genus nov.2 sp. nov. f. obscura? from Australia
2016.07.10 © Steve & Alison.
Genus nov.2 sp. nov. f. obscura from the UK
2016.12.03 © David W.
Genus nov.2 sp. nov. f. obscura from the UK
2016.12.03 © David W.
Genus nov.2 sp. nov. f. obscura from the UK
At least 3 tenent setae (*)
2016.12.03 © David W.







Genus Keratopygos Christiansen, KA et Pike, E, 2002:180, -

Species megalos (Christiansen, KA & Pike, E, 2002:181, -) Christiansen, KA & Pike, E, 2002:181, -, t.t.


Genus Arborianna Bretfeld, 2002

Species cupiubae Bretfeld, 2002, t.t.


Genus Parakatianna Womersley, H, 1932:24

  Syn.: KatiannaGroup Longkingia Salmon, JT, 1946:40


Species albirubrafrons Salmon, JT, 1943:6

  Subspecies niveanota Salmon, JT, 1943:7

             Syn.: Parakatianna albirubrafrons ssp. niveanoa(sic) Salmon, JT, 1943:plate1 ISS

Species anomala Womersley, 1933

Species cortica Salmon, JT, 1943:8

Parakatianna cortica from New Zealand
After Salmon JT, 1943 Pl.1 Fig.9

Species diversitata Salmon, JT, 1943:7

Parakatianna diversitata from New Zealand
After Salmon JT, 1943 Pl.1 Fig.4

  Subspecies viridis Salmon, JT, 1943:8

Parakatianna diversitata
subsp. viridis from New Zealand
After Salmon JT, 1943 Pl.1 Fig.11

Species hexagona Salmon, JT, 1941:414

Parakatianna hexagona from New Zealand
After Salmon JT, 1941 Pl.71 Fig.510

Species homerica (Salmon, JT, 1946:44) Betsch, 1980

Species prospina (Salmon, JT, 1946:40) Betsch, 1980, t.t.

Species salmoni Wise, 1964

Species spinata Womersley, H, 1932:25, t.t.

Species superba (Salmon, JT, 1946:42) Betsch, 1980

Longkingia superba from New Zealand
After Salmon JT, 1946 Fig.28


Genus Katianna Börner, C, 1906:182

  Syn.: ... Metakatiannidae Stach, 1956 partim
  Syn.: Katiannini Metakatianna Denis, J-R, 1933:291
  Syn.: Katiannini Metakatianna Richards, WR, 1968:22 partim
  Syn.: Katiannini P.(sic) Denis, J-R, 1933:293 lapsus IOS


  Syn.: Symphypleona Katiana(sic) de Izarra, DC, 1965:275 ISS
Katianna nr schotti from the UK
Spinelike setae on head
2008.05.06 © Ardron, P.A.
Katianna nr schotti from the UK
Peglike papilla on 3rd antennal segment
2008.05.06 © Ardron, P.A.
Katianna nr schotti ♂? & ♀? from the UK
2008.12.09 © Ardron, P.A.
Katianna sp. from Australia
2009.08.29 © Hort, J.
Katianna sp. from Peru
2013.03.17 © Cuadros Barrenechea.M., B
Katianna sp. from Tasmania
2014.04.18 © Murray, A.
Katianna sp. from Tasmania
2016.01.11 © Murray, A.
?Katianna? sp. from Chile
2019.08.19 © Guzmán, V.V.
?Katianna? sp. from Chile
2019.08.19 © Guzmán, V.V.
?Katianna? sp. from Chile
2019.08.19 © Guzmán, V.V.
?Katianna? sp. from Chile
2019.08.19 © Guzmán, V.V.
?Katianna? sp. from Taiwan
2023.05.31 © Cheng, H.-J.
?Katianna? sp. from Taiwan
2023.05.31 © Cheng, H.-J.
?Katianna? sp. from Taiwan
2021.08.05 © Cheng, H.-J.
?Katianna? sp. from Chile
2021.10.23 © Gargiulo, M.
?Katianna? sp. from Taiwan
2023.11.10 © Cheng, H.-J.
?Katianna? sp. from Taiwan
2023.11.10 © Cheng, H.-J.





Species sp.1 Delamare Deboutteville, C et Massoud, Z, 1963:241

Species sp.2 Greenslade, P et Ireson, JE, 1986:282

Katianna sp. from Australia
After Greenslade P & Ireson JE, 1986 Fig.2

Species sp.21 Ireson, JE, 1990:147

Katianna sp.1 from Tasmania
Yellow-green colour form
After Ireson JE, 1990 Fig.51
Katianna sp.1 from Tasmania
Striped colour form
After Ireson JE, 1990 Fig.52

Species sp.4 Janssens F et Ardron PA in Ardron, PA, 2009:16

Katianna sp.4 from the UK
After Ardron PA, 2009 p.16
Katianna sp. from the UK
2009.04.27 © Ardron, P.A.
Katianna sp. from the UK
2009.04.27 © Ardron, P.A.
Katianna sp. from the UK
Female subanal appendage
2009.04.27 © Ardron, P.A.
Katianna sp. from the UK
2009.06.06 © Ardron, P.A.
Katianna sp. from the UK
2009.06.06 © Ardron, P.A.
Katianna sp. from the UK
Dorsal pigmentation
2012.11.24 © Bird, S.
Katianna sp. from the UK
Lateral pigmentation
2012.11.24 © Bird, S.
Katianna sp. from the UK
Face pigmentation
2012.11.24 © Bird, S.
Katianna sp. from the UK
Posterior dorsal pigmentation
2012.11.24 © Bird, S.
Katianna sp. from the UK
Footcomplex of 3rd leg
2012.11.24 © Bird, S.
Katianna sp. from the UK
Mucro
2012.11.24 © Bird, S.
Katianna sp. from the UK
2018.05.03 © Ardron, P.A.
Katianna sp. from the UK
2018.05.03 © Ardron, P.A.
Katianna sp. from the UK
2018.05.03 © Ardron, P.A.

Species sp.5 Ardron, PA, 2009:19

Unknown sp.1 from the UK
After Ardron PA, 2009 p.19

Species sp.6 Ardron, PA, 2009:19

Katianna s.l. sp. from the UK
2008.mm.dd © Ardron, P.
Katianna s.l. sp. from the UK
2008.mm.dd © Ardron, P.
Sminthurinus nr elegans var. dorsalis from the UK
After Ardron PA, 2009 p.18
Unknown sp.2 from the UK
After Ardron PA, 2009 p.19

Species sp.8 Janssens F & Bouffler E, 2014

Katianna sp. from Australia
2014.07.03 © Bouffler, E.
Katianna sp. from Australia
2014.07.03 © Bouffler, E.
Katianna sp. from Australia
2014.07.03 © Bouffler, E.
Katianna sp. from Australia
2014.07.03 © Bouffler, E.
Katianna sp. from the UK
2019.12.30 © McCulloch, J.
Katianna sp. from the UK
2019.12.30 © McCulloch, J.
Katianna sp. from the UK
Head
2020.01.01 © McCulloch, J.
Katianna sp. from the UK
Head dorsally with curved spine-like setae
2020.01.01 © McCulloch, J.
Katianna sp. from the UK
Ant.3 with peg-like sensory organ
2020.01.01 © McCulloch, J.
Katianna sp. from the UK
Ant.3 with distinctly stronger setae
2020.01.01 © McCulloch, J.
Katianna sp. from the UK
Ant.4 with annulations
2020.01.01 © McCulloch, J.

Species sp.9 Janssens F & Ardron PA, 2015

Katianna sp. from the UK
2015.04.26 © Ardron, P.A.
Katianna sp. from the UK
2015.04.26 © Ardron, P.A.
Katianna sp. from the UK
2015.04.26 © Ardron, P.A.
Katianna sp. from the UK
Caudal aspect
2015.04.26 © Ardron, P.A.

Species sp.10 Janssens F & Dobbs B, 2016

Katianna sp. from the USA
2016.01.02 © Justis, S.
Katianna sp. from the USA
2016.11.24 © Dobbs, Br.
Katianna sp. from the USA
2016.11.24 © Dobbs, Br.
Katianna sp. from the USA
2016.11.24 © Dobbs, Br.
Katianna sp. from the USA
2016.11.24 © Dobbs, Br.
Katianna sp. from the USA
Antenna
2016.11.24 © Dobbs, Br.
Katianna sp. from the USA
Ant.3 with wart (*)
2016.11.25 © Dobbs, Br.
Katianna sp. from the USA
Dorsal pigmentation
2016.11.25 © Dobbs, Br.
Katianna sp. from the USA
2017.01.01 © Hart, J.
Katianna sp. from the USA
2017.01.01 © Hart, J.

Species sp.12 Bernard EC, Janssens F & Abbott J, 2018

Katianna sp. from the USA
2009.01.02 © Justis, S.
Katianna sp. from the USA
2009.01.02 © Justis, S.
Katianna sp. from the USA
2009.01.02 © Justis, S.
Katianna sp. from the USA
2009.01.24 © Justis, S.
Katianna sp. from the USA
2009.01.24 © Justis, S.
Katianna sp. from the USA
2009.01.24 © Justis, S.
Katianna sp. from the USA
2009.01.24 © Justis, S.
Katianna sp. from the USA
2009.02.07 © Justis, S.
Katianna sp. from the USA
2009.02.24 © Justis, S.
Katianna sp. from the USA
2009.02.24 © Justis, S.
Katianna sp. from the USA
2018.01.10 © Abbott, J.C.
Katianna sp. from the USA
Bothriotricha ABC in angle
2018.01.10 © Abbott, J.C.
Katianna sp. from the USA
2018.01.10 © Abbott, J.C.
Katianna sp. from the USA
Footcomplex with 3 clavate tenent setae (*)
2018.01.10 © Abbott, J.C.
Katianna sp. from the USA
2018.03.13 © Pugh, D.E.
Katianna sp. from the USA
2022.01.14 © Charny, V.
Katianna sp. from the USA
2022.01.14 © Charny, V.
Katianna sp. from the USA
Bothriotricha ABC in obtuse angle
2022.04.19 © Charny, V.

Species antennapartitia Salmon, JT, 1941:412

             Syn.: Katianna antennapartita(sic) Salmon, JT, 1943:4 ISS

Species aurantiaca (Womersley, H, 1935:218) Greenslade, P in Houston, WWK, 1994:124

            Syn.: Arrhopalites aurantiaca(sic) Womersley, H, 1935:218 IOS
            Syn.: Arrhopalites aurantiaca(sic) Womersley, H, 1939:922 IOS
            Syn.: Arrhopalites aurantiaca(sic) Stach, J, 1945:6,is IOS
            Syn.: Arrhopalites aurantiaca(sic) Stach, J, 1956:71 IOS
            Syn.: Arrhopalites aurantiaca(sic) Salmon, JT, 1964:570 IOS

Species australis Womersley, H, 1931:5

Katianna australis from Tasmania
After Womersley H, 1931 Pl.I Fig.1
Katianna australis from Australia
After Womersley H, 1939 Pl.1 Fig.5
Katianna australis from Australia
After Greenslade P & Ireson JE, 1986 Fig.6
Katianna australis from Tasmania
After Ireson JE, 1990 Fig.44
Katianna australis from Tasmania
After Ireson JE, 1990 Fig.45

             Syn.: Katianna australia(sic) Ireson, JE, 1990:83 ISS
             Syn.: Katianna australia(sic) Ireson, JE, 1990:147 ISS
Katianna australis from Australia
After Thurmer J in Greenslade P, 1994 p.117

  Subspecies tillyardi Womersley, H, 1932:23

  Subspecies viridula Womersley, H, 1932:23



Species banzarei Salmon, JT, 1964:314

             Syn.: Katianna banzare(sic) Greenslade P, 1990 ISS
Katianna banzarei from Antarctica
After Davis, G. in Greenslade, P., 2006 Fig.81.
Katianna banzarei from Australia
2010 © Stevens, M.



Species cardoni (Delamare Deboutteville & Massoud, 1963:237) Rapoport, 1968

            Syn.: Katianna Cardoni Delamare Deboutteville & Massoud, 1963 IOS

Species cobold Börner, 1907

            Syn.: Sminthurus luteus Parona, 1895
            Syn.: Sminthurus luteus Börner, 1907
            Syn.: Sminthurus viridis Parona, 1895 partim
            Syn.: Sminthurus viridis Börner, 1907 partim
            Syn.: Katianna cobold var. decora Börner, 1907
            Syn.: Katianna cobold var. guttulata Börner, 1907


Species coeruleocephala Handschin, 1920

Species drummondi (Womersley, H, 1932:18) Denis, J-R, 1933:321

Species gloriosa Salmon, JT, 1946:38

Species houssayi Delamare Deboutteville & Massoud, 1963:225

            Syn.: Katianna Houssayi(sic) Delamare Deboutteville & Massoud, 1963 IOS

Species jeanneli Delamare Deboutteville & Massoud, 1963:222

Species kerguelenensis Denis, 1947

Remark by Deharveng, L. 1981 (cfr Greenslade, P., 2007.10.09:in litt.): This species probably belongs in the genus Pseudokatianna because it lacks the generic character of spines on the head according to the figures.

             Syn.: Katianna kerguelensis(sic) Greenslade, P, 1997-:www0312 ISS
Remark by Greenslade, P. 2007.10.09:in litt.: Taking into consideration the form of the mucro, the chaetotaxy of the dens, the slight antennal annulation and other features, the species probably is not a Pseudokatianna but may belong in Polykatianna, especially as Denis noted it had conspicuous interocular bosses. However, neither of these two genera are well defined, so the correct generic placement for this species is not possible to determine at present.



Species kuscheli Delamare Deboutteville & Massoud, 1963:228

            Syn.: Katianna Kuscheli(sic) Delamare Deboutteville & Massoud, 1963 IOS

Species liliputana (Nicolet, H, 1847:360) Stach, J, 1956:239, i.s.

Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Smynthurus(sic) liliputanus from Chile
After Nicolet H in Gay C, 1854 Lam.1 Fig.9
             Syn.: Smynthurus Liliputanus(sic) Lubbock, J, 1873:116 ISS


Species maryae Bernard, EC, 2014:149

Katianna sp. from the USA
2006 © Bernard, E.C.
Katianna maryae from the USA
Scale bar = 500 µm
After Bernard EC, 2014 Fig.1E
Katianna maryae ♀ from the USA
Scale bar = 500 µm
After Bernard EC, 2014 Fig.1A
Katianna maryae ♀ from the USA
Scale bar = 500 µm
After Bernard EC, 2014 Fig.1B
Katianna maryae ♀ from the USA
Scale bar = 500 µm
After Bernard EC, 2014 Fig.1C
Katianna maryae ♂ from the USA
Scale bar = 500 µm
After Bernard EC, 2014 Fig.4A
Katianna maryae from the USA
2021.01.11 © Shoemaker, C.
Katianna maryae from the USA
2021.01.12 © Shoemaker, C.
Katianna maryae from the USA
2021.01.21 © Shoemaker, C.
Katianna maryae from the USA
2021.01.21 © Shoemaker, C.
Katianna maryae from the USA
2021.02.12 © Shoemaker, C.
Katianna maryae from the USA
2022.01.26 © Charny, V.
Katianna maryae from the USA
2022.01.26 © Charny, V.
Katianna maryae from the USA
2022.01.26 © Charny, V.
Katianna maryae from the USA
2022.12.14 © Charny, V.
Katianna maryae from the USA
2022.12.14 © Charny, V.
Katianna maryae from the USA
2022.12.15 © Fleming, K.
Katianna maryae from the USA
2022.12.15 © Fleming, K.
Katianna maryae from the USA
2024.01.07 © Coogler, J.
Katianna maryae from the USA
2024.01.07 © Coogler, J.
Katianna maryae from the USA
Third footcomplex
With clavate tenent macrosetae
2024.01.07 © Coogler, J.
Katianna maryae from the USA
2024.01.09 © Coogler, J.
Katianna maryae from the USA
Ant.4 with about 12 subdivisions
2024.01.09 © Coogler, J.

Species mnemosyne Börner, C, 1906:182, t.t.

            Syn.: Sminthurus multifasciatus Parona, 1895
            Syn.: Sminthurus multifasciatus Börner, C, 1906:182


Katianna-mnemosyne ♀ from Argentina
After Bonet F, 1933 Fig.XXI

Species mucina Womersley, 1933

Katianna mucina from Australia
After Greenslade P & Ireson JE, 1986 Fig.3
Katianna mucina from Tasmania
After Ireson JE, 1990 Fig.46

Species nunezi (Najt, 1967) Betsch, 1977

Species obscura (Womersley, H, 1936:485) Greenslade, P & Ireson, JE, 1986:275

Parakatianna obscura from Australia
After Womersley H, 1936 Fig.7a
Parakatianna obscura from Australia
After Womersley H, 1936 Fig.7b
Parakatianna obscura from Australia
After Womersley H, 1936 Fig.7c
Katianna sp. cf. obscura from Australia
After Greenslade P & Ireson JE, 1986 Fig.5
Katianna cf. obscura from Tasmania
After Ireson JE, 1990 Fig.47

Species oceanica Schött, H, 1917:57

Katianna oceanica from Australia
After Schött H, 1917 Taf.4 Fig.6
Katianna oceanica from Australia
Abdomen posterior part
After Schött H, 1917 Taf.4 Fig.7

Species ornata Womersley, H, 1932:23

Katianna ornata from Tasmania
After Ireson JE, 1990 Fig.48

Species patagonica (Delamare Deboutteville, C & Massoud, Z, 1963:198) Betsch, 1977

Remark by Mari Mutt J.A. & Bellinger P.F. 1990:138: The genus Metakatianna houses a number of species based on first instars of species of Katianna, Sminthurinus, Pseudokatianna, etc. (Betch, 1980:12). We have listed the species of Metakatianna under Katianna but the position of many is still undetermined.
Species perplexa Salmon, JT, 1944:176

Species pescotti Womersley, H, 1935:218

Katianna cf. pescotti from Tasmania
After Ireson JE, 1990 Fig.50
Katianna pescotti from Tasmania
2014.04.28 © Murray, A.
Katianna pescotti from Tasmania
Body entirely yellow; strong macrosetae on head and body
2015.09.26 © Tony.
Katianna pescotti from Tasmania
Antennae bluish
2015.09.26 © Tony.
Katianna pescotti from Tasmania
Ant.3 with peg (*); ant.4 with 12 subdivisions
2015.09.26 © Tony.

Species poivrei Delamare Deboutteville, C et Massoud, Z, 1963:233

            Syn.: Katianna Poivrei(sic) Delamare Deboutteville, C et Massoud, Z, 1963:233 IOS

Species puella (Denis, J-R, 1933:281) Betsch, 1977

Metakatianna puella from Costa Rica
After Denis J-R, 1933 Fig.86
             Syn.: Metakatianna peulla(sic) Salmon, JT, 1964:589 ISS
             Syn.: Katianna paella(sic) Palacios-Vargas, 1992 ISS


Species purpuravirida Salmon, JT, 1941:413

Katianna purpuravirida from New Zealand
After Salmon JT, 1941 Pl.71 Fig.502
             Syn.: Katianna purpurvirida(sic) Salmon, JT, 1946:38 ISS


Species reducta Salmon, JT, 1944:176

Species richardsi Najt, 1967

            Syn.: Katianna richardsi Rapoport & Najt, 1966,nn


Species ruberoculata Salmon, JT, 1944:175

Species schotti Womersley, 1933

            Syn.: Katianna oceanica var. schötti(sic) Womersley, 1933 IOS
            Syn.: Katianna oceanica King, Hutchinson & Greenslade P, 1976
            Syn.: Katianna oceanica Greenslade P, 1981


             Syn.: Katianna oceania(sic) var. schötti(sic) Womersley, H, 1935:218 ISS
             Syn.: Katianna oceanica var. schötti(sic) Womersley, H, 1939:235 ISS
             Syn.: Katianna schoetti(sic) King, Greenslade P & Hutchinson, 1985 ISS
             Syn.: Katianna oceanica var. schoetti(sic) Greenslade, P et Ireson, JE, 1986:275 ISS
Katianna oceanica var. schoetti(sic) from Australia
After Greenslade P & Ireson JE, 1986 Fig.4

             Syn.: Katianna oceanica var. schoetti(sic) Greenslade & Greenslade, 1987 ISS
             Syn.: Katianna oceanica var. schoetti(sic) Ireson, JE, 1990:83 ISS
Katianna oceanica var. schoetti(sic) from Tasmania
After Ireson JE, 1990 Fig.49

             Syn.: Katianna oceanica var. schoetti(sic) Ireson, 1993 ISS
             Syn.: Katianna schoetti(sic) Greenslade, P in Houston, WWK, 1994:126 ISS
             Syn.: Katianna schoetti(sic) Ardron, PA, 2009:14 ISS
Katianna schoetti(sic) from the UK
After Ardron PA, 2009 p.14

            Syn.: Katianna australis Ardron, PA, 2009:15
Katianna australis from the UK
After Ardron PA, 2009 p.15
Remark by Greenslade, P. 2013.11.19 il: However carefully you examine specimens from England and compare with published descriptions from Australia, you cannot determine species of Katianna with accuracy. The only desciptions of K. australis that exist were by Womersley in the, 1930s. His descriptions are notoriously brief and totally inadequate. We probably have over 50 undescribed species of Katianna in Australia and New Zealand, maybe more. Womersley's description of K. australis could apply to many of them.




Katianna nr schotti ♂? from the UK
2008.04.30 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2008.04.30 © Ardron, P.A.
Katianna nr schotti from the UK, mucrones
2008.04.30 © Ardron, P.A.
Katianna nr schotti from the UK
Dorsal macrosetae
2008.04.30 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2008.05.06 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2008.05.06 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2008.05.06 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2008.05.06 © Ardron, P.A.
Katianna nr schotti from the UK
Foot complex with tenent hairs
2008.05.06 © Ardron, P.A.
Katianna nr schotti ♀? from the UK
2008.12.09 © Ardron, P.A.
Katianna nr schotti ♀? from the UK
2008.12.11 © Ardron, P.A.
 
Katianna nr schotti ♀? from the UK
2008.12.19 © Ardron, P.A.
 
Katianna nr schotti ♂? & ♀? from the UK
2008.12.19 © Ardron, P.A.
Katianna nr schotti ♀? from the UK
2009.02.01 © Ardron, P.A.
Katianna nr schotti ♀? from the UK
2009.02.01 © Ardron, P.A.
Katianna nr schotti ♀? from the UK
2009.02.01 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2009.05.06 © Ardron, P.A.
 
Katianna nr schotti ♂? from the UK
2009.05.06 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
Grooming using everted collophore vesicle
2009.05.06 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2009.05.06 © Ardron, P.A.
Katianna nr schotti ♂? & ♀? from the UK
2009.06.06 © Ardron, P.A.
Katianna nr schotti ♀? & ♂? from the UK
2009.06.06 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2009.06.06 © Ardron, P.A.
Katianna nr schotti ♀? from the UK
2009.06.06 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2009.06.06 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2009.12.20 © Jones, A.
Katianna nr schotti ♂? from the UK
2011.12.04 © Jones, A.
Katianna nr schotti ♂? from the UK
2011.12.05 © Jones, A.
Katianna nr schotti ♂? from the UK
2011.12.05 © Jones, A.
Katianna nr schotti ♂? from the UK
2013.03.21 © Nurcombe, E.
Katianna nr schotti juv. from the UK
2013.03.21 © Nurcombe, E.
Katianna nr schotti ♀? from the UK
2013.04.13 © Henton-Jones, D.
Katianna nr schotti ♂? from the UK
2013.12.11 © Nurcombe, E.
Katianna nr schotti ♂? from the UK
2013.12.11 © Nurcombe, E.
Katianna nr schotti ♂? from the UK
2013.12.11 © Nurcombe, E.
Katianna nr schotti ♂? from the UK
2014.11.09 © Evans, M.
Katianna nr schotti ♂? from the UK
2014.11.09 © Evans, M.
Katianna nr schotti ♀? from the UK
2014.12.21 © Horton, M.
Katianna nr schotti ♀? from the UK
2014.12.21 © Horton, M.
Katianna nr schotti ♀? from the UK
2015.03.03 © Cunningham, T.
Katianna nr schotti ♂? from Holland
2015.11.18 © van Duinen, J.
Katianna nr schotti ♀? from Holland
2015.12.18 © van Duinen, J.
Katianna nr schotti ♀? from Holland
2015.12.18 © van Duinen, J.

             Syn.: Katianna schoetti(sic) Shaw, P, 2017:146 ISS
             Syn.: Katianna schoetti(sic) Shaw, P, 2017:148 ISS
             Syn.: Katianna schoetti(sic) Shaw, P, 2017:172 ISS
Katianna ?schotti? ♂? from Belgium
2018.10.08 © Huskens, M.-L.
Katianna ?schotti? ♂? from Belgium
2018.10.08 © Huskens, M.-L.
Katianna ?schotti? ♂? from the UK
2018.11.05 © Phillips, E.
Katianna ?schotti? ♂? from the UK
Forwards bent cephalic spikes (*)
2018.11.05 © Phillips, E.
Katianna ?schotti? from the UK
2020.10.28 © Phillips, E.
Katianna ?schotti? ♀? from the UK
2020.11.09 © Phillips, E.
Katianna ?schotti? ♀? from the UK
Footcomplex with many clavate tenent macrosetae
2020.11.09 © Phillips, E.
Katianna ?schotti? juv. ♂? from the UK
2021.01.29 © Phillips, E.
Katianna ?schotti? subadult ♂? from the UK
2021.01.29 © Phillips, E.
Katianna ?schotti? subadult ♂? from the UK
2021.01.29 © Phillips, E.
Katianna ?schotti? ♂? from the UK
Ant.4 distinctly black
2021.02.05 © Phillips, E.
Katianna ?schotti? from Australia
2021.04.15 © Hobern, D.
Katianna ?schotti? from Germany
2021.11.21 © Jäschke, J.
Katianna ?schotti? from Germany
2021.11.24 © Jäschke, J.
Katianna ?schotti? from Germany
2021.11.24 © Jäschke, J.





Species serrae Najt, 1967

Species serrata (Schäffer, 1897) Börner in Voeltzkow, 1907

Species serrata Womersley, 1936, nec Schäffer, 1897

Species steparia (Najt, 1967) Betsch, 1980

            Syn.: Metakatianna steparia de Izarra, DC, 1965:275 nn
            Syn.: Metakatianna steparia Rapoport & Najt, 1966 nn


Species uschuaiensis (Schäffer, 1897) Stach, 1956

Species viretorum Najt, 1967

Species viridis Delamare Deboutteville & Massoud, 1963:237

Species willincki (Delamare Deboutteville, C & Massoud, Z, 1963:269) Betsch, 1980

Species wygodzinskyi Delamare Deboutteville & Massoud, 1963:228

            Syn.: Katianna Wygodzinskyi(sic) Delamare Deboutteville & Massoud, 1963 IOS
Katianna nr wygodzinskyi from Peru
2009.01-03.dd © Contreras Valdez, A.



Species zebra (Womersley, H, 1942:30) Greenslade, P in Houston, WWK, 1994:126

Parakatianna zebra from Australia
After Womersley H, 1942 Fig.7A
Parakatianna zebra from Australia
After Womersley H, 1942 Fig.7B


Genus Polykatianna Salmon, JT, 1946:54

  Syn.: ... Millsurus Betsch, 1977


Polykatianna sp. from Brazil
After Cipola NG & al, 2019 Fig.16

Species aurea (Womersley, H, 1932:26) Bruzzese, 1980

Species cremea Salmon, JT, 1949:55

Species davidi (Tillyard, RJ, 1920:14) Greenslade P & Wise, 1986

            Syn.: Metakatianna gressitti Salmon, JT, 1964:317
            Syn.: Parakatianna litorea Salmon, JT, 1943:5
Polykatianna litorea ssp. litorea from New Zealand
2014 © Te Papa Museum

            Syn.: Parakatianna litorea ssp. luteaterga Salmon, JT, 1943:6
Parakatianna litorea ssp. luteaterga from New Zealand
After Salmon, J.T., 1943 Pl.1 Fig.6
Polykatianna litorea ssp. luteaterga from New Zealand
2014 © Te Papa Museum




             Syn.: Arrhopalites? Davidi(sic) Denis, J-R, 1933:290 ISS
             Syn.: Arrhopalites? Davidi(sic) Denis, J-R, 1933:295 ISS
Polykatianna davidi from Antarctica
After Davis, G. in Greenslade, P., 2006 Fig.78.



Species flammea Salmon, JT, 1946:55, t.t.

Species montana (Womersley, H, 1936:483) Greenslade P in Houston, 1994:128

Parakatianna montana from Australia
After Womersley H, 1936 Fig.6a

Species polygonia (Snider, 1978)

Species pulchella (Womersley, 1933) Greenslade P in Houston, 1994:128

Species quinseta (Salmon, 1954) Stach, 1956

             Syn.: Sminthurus quinqueseta(sic) Stach, J, 1956:206,is ISS

Species radicula (Maynard, EA, 1951:224) Arnett, 1970

Sminthurinus radiculus from the USA
After Maynard EA, 1951 Plate XXIII Fig.378

            Syn.: Sminthurinus radiculus f. pallens Maynard, EA, 1951:225
Sminthurinus radiculus f. pallens from the USA
After Maynard EA, 1951 Plate XXIII Fig.380

            Syn.: Sminthurinus radiculus f. pictus Maynard, EA, 1951:224
Sminthurinus radiculus f. pictus from the USA
After Maynard EA, 1951 Plate XXIII Fig.379




Polykatianna radicula from the USA
2012.04.26 © Leavengood, J.
Polykatianna radicula from the USA
2012.04.26 © Leavengood, J.

Species sminthurina (Mills, HB, 1934:99) Richards, 1968

             Syn.: Polykatianna sminthurinus(sic) Richards, 1968 ISS
Polykatianna sminthurina from the USA
2013.02.11 © Anderson, J.
Polykatianna sminthurina from the USA
2013.02.11 © Anderson, J.
?Polykatianna sminthurina? from Germany
Ant.4 subdivided; ant.3 with warth (*)
2014.10.26 © Landgraf, A.
Polykatianna sminthurina from the USA
2014.12.23 © Montgomery, G.
Polykatianna sminthurina from the USA
2019.02.03 © Coulter, Be.
Polykatianna sminthurina from the USA
2019.02.03 © Coulter, Be.
Polykatianna sminthurina from the USA
Ant.4 with long basal + 13 short apical subdivisions
2019.02.03 © Coulter, Be.
Polykatianna sminthurina from the USA
Dark antennae
Only ant.2 pale
2021.02.04 © Pearson, R.R.
Polykatianna sminthurina from the USA
2022.04.16 © Pavy, R.
Polykatianna sminthurina from the USA
2022.04.16 © Pavy, R.



Species utahensis (Snider, RJ, 1988:1)

Sminthurinus (Polykatianna) utahensis from the USA
After Snider RJ, 1988 Fig.1
Sminthurinus (Polykatianna) utahensis from the USA
After Snider RJ, 1988 Fig.2


Genus Sminthurinus Börner, C, 1901:99 key

  Syn.: Poduridæ(sic) Symnthurus(sic) Fitch, A, 1863:675 lapsus ISS
  Syn.: ... Symnthurus(sic) Fitch in Van Benthuysen, 1867 ISS
  Syn.: Sminthuridæ(sic) Sminthurus Schött, H, 1891:7 partim
  Syn.: Smynthuri(sic) Smynthurus(sic) Folsom, JW, 1896:448 partim IOS
  Syn.: ... Smynthurus(sic) Meinert, 1896 IOS
  Syn.: ... Smynthurus(sic) Iacobson, 1898 IOS
  Syn.: ... Smynthurus(sic) Wahlgren, 1900 IOS
  Syn.: ... Smynthurus(sic) Nielsen, 1909 IOS
  Syn.: Sminthuridae Smynthurus(sic) Gisin, H, 1960:280 IOS
  Syn.: Sminthuridae Symnthurus(sic) Barnes, JK, 1988:89 ISS
  Syn.: Sminthuridae Symnthurus(sic) Barnes, JK, 1988:90 ISS
  Syn.: Sminthurus Pilosi Schött, H, 1891:11 partim
  Syn.: Sminthurus Eusminthurus Börner, C, 1901:99 partim
  Syn.: Sminthurinus Inarticulati Börner, C, 1901:100
  Syn.: Sminthuridinae Smynthurella Houlbert, C, 1924:157
  Syn.: Sminthuridae Metakatianna Gisin, H, 1960:277
  Syn.: Symphypleona Metakatiana(sic) de Izarra, DC, 1965:275 ISS
  Syn.: Katiannidae Metakatianna Palacios-Vargas, JG et Vacaflores-Argandoña, MR, 2020:201 = first instar
  Syn.: Sminthuridae Stenognathellus Gisin, H, 1960:277
  Syn.: Katiannini Stenognathellus Richards, WR, 1968:23
  Syn.: Sminthuridae Katiannina Poole, RW in Poole, RW et Gentili, P, 1997:55
  Syn.: Sminthuridae Millsurus Poole, RW in Poole, RW et Gentili, P, 1997:55
  Syn.: Sminthuridae Neokatianna Poole, RW in Poole, RW et Gentili, P, 1997:55
  Syn.: Sminthuridae Polyclavella Poole, RW in Poole, RW et Gentili, P, 1997:55
  Syn.: Sminthuridae Polykatianna Poole, RW in Poole, RW et Gentili, P, 1997:55


  Syn.: Katiannini Smithurinus(sic) Stach, J, 1956:96 ISS
  Syn.: Katiannini Sminthurnius(sic) Yosii, R, 1977:164 ISS
  Syn.: Sminthurididae(sic) Sminthurimus(sic) Camousseight, A, 1999:6 ISS
Remark by ICZN. 2007:759: Placed on the Official List of Names in Zoology. Opinion 1027.


  Syn.: Collembola Simthurinus(sic) Chauvat, M, Trap, J, Perez, G, Delporte, P et Aubert, M, 2011:412 ISS
Sminthurinus henshawi from the USA
4th antennal segment not subdivided
2011.12.03 © Justis, S.
Sminthurinus sp. from the UK
Facial chaetotaxy
2012.11.09 © Murray, A.


  Syn.: Symphypleona Smynthurinus(sic) Shayanmehr, M, Yahyapour E, Kahrarian, M et Lafooraki EY, 2013:73 ISS
Sminthurinus sp. from the UK
Ventral aspect: furca and everted vesicles of collophore (*)
2013.11.07 © Phillips, E.
Sminthurinus aureus f. reticulatus from the UK
Footcomplex with clavate tenant setae (*)
2014.02.06 © Phillips, E.
Sminthurinus aureus from the UK
Ant.4 not subsegmented
2016.02.23 © Phillips, E.
Sminthurinus sp. from the USA
2016.03.09 © Abela, A.
Sminthurinus aureus f. atratus from Belgium
Ant.3 with wart (W)
2020.11.06 © Huskens, M.-L.
Sminthurinus sp. ♂ from Taiwan
Depositing spermatophore
2022.01.13 © Cheng, H.-J.
Sminthurinus henshawi from the USA
Feeding on fungal hyphae
2023.01.02 © Gruver, Be.
Sminthurinus sp. from Belgium
With aberrant white dorsal spots
2023.01.03 © Huskens, M.-L.
Sminthurinus sp. from Belgium
Grooming in tripod modus
Tripod stabilised using collophore vesicles
2024.04.04 © Huskens, M.-L.





Species sp.1 Bretfeld, 2000

Species sp.2 Zeppelini, D, Bellini, BC, Creao-Duarte, AJ et Hernández, MIM, 2009:1166

Species sp.3 Zeppelini, D, Bellini, BC, Creao-Duarte, AJ et Hernández, MIM, 2009:1166

Species sp.4 Hornung-Leoni, CT et Palacios-Vargas, JG, 2012:67

Species sp.5 Janssens F & Phillips E, 2017

Sminthurinus sp. ♀ from the UK
2017.10.09 © Phillips, E.
Sminthurinus sp. ♀ from the UK
Caudal aspect
2017.10.09 © Phillips, E.
Sminthurinus sp. juv. from the UK
2017.10.09 © Phillips, E.
Sminthurinus sp. from the UK
Abd.5 with middorsal spot and transverse stripe
2017.10.19 © Phillips, E.
Sminthurinus sp. from the UK
2017.10.19 © Phillips, E.
Sminthurinus sp. from the UK
2017.10.20 © Phillips, E.
Sminthurinus sp. from the UK
2017.10.20 © Phillips, E.
Sminthurinus sp. juv. from the UK
2017.10.25 © Phillips, E.
Sminthurinus sp. from the UK
2017.10.25 © Phillips, E.
Sminthurinus sp. from the UK
2017.10.25 © Phillips, E.
Sminthurinus sp. from the UK
2017.10.28 © Lewis, M.
Sminthurinus sp. from the UK
2017.10.28 © Lewis, M.
Sminthurinus sp. from the UK
2017.10.28 © Lewis, M.
Sminthurinus sp. from the UK
2017.11.14 © Phillips, E.
Sminthurinus sp. from the UK
2017.11.14 © Phillips, E.
Sminthurinus sp. from the UK
2017.11.14 © Phillips, E.

Species sp.6 Janssens F & van Duinen J, 2018

Sminthurinus sp. from France
2018.06.19 © van Duinen, J.
Sminthurinus sp. from France
2018.06.19 © van Duinen, J.

Species sp.7 Janssens F & Tiky S, 2018

Sminthurinus sp. nr reticulatus from France
2018.11.12 © Tiky, S.
Sminthurinus sp. nr reticulatus from France
2018.11.12 © Tiky, S.
Sminthurinus sp. nr reticulatus from France
2018.11.12 © Tiky, S.
Sminthurinus sp. nr reticulatus from France
2018.12.07 © Lebeaux, P.

Species sp.8 Janssens F & Miller T, 2021

Sminthurinus sp. from Canada
2021.05.21 © Miller, T.
Sminthurinus sp. from Canada
2021.05.21 © Miller, T.
Sminthurinus sp. from Canada
2021.05.21 © Miller, T.

Species albipes (Schött, 1896) Stach, 1956, i.s.

             Syn.: Sminthurinus albiceps(lapsus) Stach, 1956 lapsus ISS

Species alpinus Gisin, 1953

            Syn.: Sminthurinus alpinus ssp. bisetosus Ellis, WN, 1976:318


             Syn.: Sminthurinus niger f. alpina(sic) Gisin, H, 1960:280 ISS
Sminthurinus alpinus from Switzerland
Subapical anterior dental setae absent (*)
Proximal posterior setae: 1 outer, 1 median, 2 inner
Modified after Gisin H, 1963 Fig.7
Sminthurinus alpinus ♀ from Scandinavia
Sub-anal appendage with deeply incised apical branches
After Fjellberg A, 2007 Fig.115G
Sminthurinus alpinus from Corsica
After Schulz H-J, 2015 Abb.7
Sminthurinus alpinus from France
2018.11.10 © Tiky, S.
Sminthurinus alpinus from France
2018.11.10 © Tiky, S.
Sminthurinus alpinus from France
2018.11.10 © Tiky, S.
Sminthurinus alpinus from France
Facial pattern
2019.11.24 © Tiky, S.
Sminthurinus alpinus from France
Bipartite eyebrow
2019.11.24 © Tiky, S.
Sminthurinus alpinus from France
2019.12.06 © Tiky, S.
Sminthurinus alpinus from France
Caudal aspect
2019.12.06 © Tiky, S.
Sminthurinus alpinus from France
"Eyebrows"
2019.12.06 © Tiky, S.
Sminthurinus alpinus from France
2020.10.09 © Tiky, S.
Sminthurinus alpinus from France
2020.10.09 © Tiky, S.



Species antennalis (Rapoport, 1962) Delamare Deboutteville & Massoud, 1963:213

Species atrapallidus Snider, 1978

Sminthurinus atrapallidus from the USA
2006.02.25 © Schimming, L.
Sminthurinus atrapallidus from the USA
2009.02.07 © Justis, S.
Sminthurinus atrapallidus from the USA
2009.02.07 © Justis, S.
Sminthurinus atrapallidus from the USA
2009.02.07 © Justis, S.
Sminthurinus atrapallidus from the USA
2009.02.07 © Justis, S.
Sminthurinus atrapallidus from the USA
2009.02.07 © Justis, S.
Sminthurinus atrapallidus from the USA
2017.01.25 © Vitanza, S.
Sminthurinus atrapallidus from the USA
2022.03.23 © Charny, V.
Sminthurinus atrapallidus ♀ (up) & ♂ (low) from the USA
2022.03.23 © Charny, V.
Sminthurinus atrapallidus from the USA
2023.01.30 © Wang, G.

Species aureus (Lubbock, J, 1862:589) Börner, C, 1901:103, B

Smynthurus(sic) aureus from the UK
After Lubbock J, 1862 Pl.LIX Fig.1

            Syn.: Sminthurus ochraceus Koch, 1840
            Syn.: Sminthurinus signifer Denis, 1933
            Syn.: Smynthurus(sic) signifer Stach, J, 1956:241
            Syn.: Sminthurus quadrilineatus Tullberg, 1871
            Syn.: Sminthurus quadrilineatus var. Ochropus(sic) Reuter OM in Schött, H, 1893:32 IOS
            Syn.: Sminthurus quadrilineatus var. ochropus Reuter, 1890 u
            Syn.: Sminthurus 4-lineatus(sic) var. ochropus Reuter, OM, 1892:231 nn
            Syn.: Sminthurinus aureus var. ochropus Börner, C, 1901:104
            Syn.: Sminthurinus aureus ssp. ochropus Salmon, JT, 1964:578 partim Europan records
Sminthurinus aureus f. ochropus from the UK
2009.04.05 © Ardron, P.A.


            Syn.: Sminthurinus aureus Ardron, PA, 2009:19
Sminthurinus nr aureus from the UK
After Ardron PA, 2009 p.19

Sminthurinus aureus f. ochropus from the UK
2011.11.28 © Kent, J.J.
Sminthurinus aureus f. ochropus from the UK
2011.11.30 © Nurcombe, E.
Sminthurinus aureus f. ochropus from the UK
2012.04.06 © Nurcombe, E.
Sminthurinus aureus f. ochropus from the UK
2012.04.06 © Nurcombe, E.
Sminthurinus aureus f. ochropus from the UK
2012.07.05 © Nurcombe, E.
Sminthurinus aureus f. ochropus from the UK
2012.07.05 © Nurcombe, E.
Sminthurinus aureus f. ochropus from the UK
2012.07.05 © Nurcombe, E.
Sminthurinus aureus f. ochropus from the UK
2012.10.02 © Murray, A.
Sminthurinus aureus f. ochropus from the UK
2012.10.02 © Murray, A.
Sminthurinus aureus f. ochropus from the UK
2012.12.18 © Phillips, E.
Sminthurinus aureus f. ochropus from the UK
2012.12.18 © Phillips, E.
Sminthurinus aureus f. ochropus from the UK
2013.01.07 © Nurcombe, E.
Sminthurinus aureus f. ochropus juv. from the UK
2013.09.12 © Phillips, E.
Sminthurinus aureus f. ochropus juv. from the UK
2013.09.12 © Phillips, E.
Sminthurinus aureus f. ochropus juv. from the UK
2013.10.04 © Phillips, E.
Sminthurinus aureus f. ochropus juv. from the UK
2013.10.08 © Phillips, E.
Sminthurinus aureus f. ochropus juv. from the UK
2013.10.08 © Phillips, E.
Sminthurinus aureus f. ochropus juv. from the UK
2013.10.14 © Phillips, E.
Sminthurinus aureus f. ochropus juv. from the UK
2013.10.29 © Phillips, E.
Sminthurinus aureus f. ochropus from the UK
2013.10.29 © Phillips, E.
Sminthurinus aureus f. ochropus from the UK
2013.12.19 © Barton, T.
Sminthurinus aureus f. ochropus from the UK
2013.12.19 © Barton, T.
Sminthurinus aureus f. ochropus juv. from the UK
2014.08.14 © Phillips, E.
Sminthurinus aureus f. ochropus from the UK
2014.08.15 © Nurcombe, E.
Sminthurinus aureus f. ochropus from the UK
2014.08.22 © Phillips, E.
Sminthurinus aureus f. ochropus from Germany
2014.10.05 © Jäschke, J.
Sminthurinus aureus f. ochropus from the UK
2014.11.04 © Phillips, E.
Sminthurinus aureus f. ochropus juv. from the UK
2014.11.22 © Phillips, E.
Sminthurinus aureus f. ochropus juv. from the UK
2014.11.22 © Phillips, E.
Sminthurinus aureus f. ochropus from the UK
2014.11.22 © Phillips, E.
Sminthurinus aureus f. ochropus juv. from the UK
2014.11.29 © Phillips, E.
Sminthurinus aureus f. ochropus from the UK
2016.11.12 © Phillips, E.
Sminthurinus aureus f. ochropus from Belgium
2016.12.18 © Huskens, M.-L.
Sminthurinus aureus f. ochropus juv. from France
Ant.3 wart with 4 papilla (*)
2017.12.10 © Picard, J.
Sminthurinus aureus f. ochropus from Belgium
Everted collophore vesciles (*)
2017.12.27 © Huskens, M.-L.
Sminthurinus aureus f. ochropus from Belgium
2018.08.19 © Huskens, M.-L.
Sminthurinus aureus f. ochropus juv. from the UK
2018.10.03 © Phillips, E.
Sminthurinus aureus f. ochropus juv. from Belgium
Aberration with 2 bilateral white thoracic spots
2018.11.01 © Huskens, M.-L.
Sminthurinus aureus f. ochropus adult + juv. from Belgium
2019.10.06 © Huskens, M.-L.
Sminthurinus aureus f. ochropus ♀ from Belgium
Midanal seta bifurcate (**)
Sub-anal appendages on papillae (*)
2019.10.24 © Huskens, M.-L.
Sminthurinus aureus f. ochropus ♀ from Belgium
Midanal seta bifurcate (v)
Sub-anal appendages on papillae (*)
2020.06.20 © Huskens, M.-L.
Sminthurinus aureus f. ochropus ♀ from Belgium
Deposition of egg
1. Begin of production of egg
2020.07.17 © Huskens, M.-L.
Sminthurinus aureus f. ochropus ♀ from Belgium
Deposition of egg
2. End of production of egg
Chorion being hardened
2020.07.17 © Huskens, M.-L.
Sminthurinus aureus f. ochropus ♀ from Belgium
Deposition of egg
3. Egg covered with faeces
2020.07.17 © Huskens, M.-L.
Sminthurinus aureus f. ochropus ♀ from Belgium
Deposition of egg
4. Dried faecal cover
2020.07.17 © Huskens, M.-L.
Sminthurinus aureus f. ochropus ♀ from Belgium
Deposition of egg
5. Egg deposited at substrate
2020.07.17 © Huskens, M.-L.
Sminthurinus aureus f. ochropus juv. from Belgium
Patchy dorsal pattern
Preparing to moult
2020.07.17 © Huskens, M.-L.
Sminthurinus aureus f. ochropus ♂ from Belgium
Genital papilla (*)
2021.04.28 © Huskens, M.-L.
Sminthurinus aureus f. ochropus ♂ from Belgium
2021.06.02 © Huskens, M.-L.
Sminthurinus aureus f. ochropus ♀ from Belgium
Egg production
2023.09.18 © Huskens, M.-L.
Sminthurinus aureus f. ochropus ♀ from Belgium
Egg produced via gonopore
Blown-up like balloon
2023.09.18 © Huskens, M.-L.
Sminthurinus aureus f. ochropus ♀ from Belgium
Egg transferred to anus
2023.09.18 © Huskens, M.-L.
Sminthurinus aureus f. ochropus ♀ from Belgium
Egg deposited at substrate
2023.09.18 © Huskens, M.-L.


            Syn.: Sminthurus aureus var. fusca(sic) Krausbauer, 1898 IOS
            Syn.: Sminthurus (Eusminthurus) aureus var. fusca(sic) Börner, 1900 IOS
            Syn.: Sminthurinus aureus var. fusca(sic) Börner, C, 1901:104 =? ochropus IOS
            Syn.: Sminthurinus aureus var. fusca(sic) Börner, C, 1901:138 = ochropus IOS
            Syn.: Sminthurinus aureus var. fusca(sic) Voigts, 1903 IOS
            Syn.: Sminthurus aureus var. fusca(sic) Linnaniemi, WM, 1912:287 =? ochropus IOS
            Syn.: Sminthurinus aureus var. fusca(sic) Stach, J, 1956:122 ab. IOS
            Syn.: Sminthurus aureus var. fusca(sic) Salmon, JT, 1964:578 partim = ochropus IOS
            Syn.: Sminthurinus aureus f. fusca(sic) Stach, 1964 IOS
Sminthurinus aureus f. fuscus from Holland
2012.03.02 © van Duinen, J.
Sminthurinus aureus f. fuscus from the UK
2012.06.18 © Murray, A.
Sminthurinus aureus f. fuscus from the UK
2013.11.05 © Nurcombe, E.
Sminthurinus aureus f. fuscus from France
With pale labrum
2017.11.12 © Picard, J.
Sminthurinus aureus f. fuscus from Belgium
2018.09.20 © Huskens, M.-L.
Sminthurinus aureus f. fuscus juv. from the UK
2020.08.24 © Phillips, E.
Sminthurinus aureus f. fuscus from the UK
2020.09.22 © Phillips, E.
Sminthurinus aureus f. fuscus from the UK
2020.09.22 © Phillips, E.
Sminthurinus aureus f. fuscus from France
2020.10.08 © Dejoux, P.

            Syn.: Sminthurus albus Schäffer, 1900
             Syn.: Sminthurinus aureus var. alba(sic) Börner, 1901 ISS
             Syn.: Sminthurus aureus var. alba(sic) Krausbauer, 1901 ISS
             Syn.: Sminthurinus aureus f. alba(sic) Handschin, E in Dahl, F, 1929:113 ISS
             Syn.: Sminthurinus aureus f. alba(sic) Marlier, GJ, 1942:10,B ISS
             Syn.: Sminthurinus aureus var. alba(sic) Bretfeld, G, 1999:110 ISS
Sminthurinus aureus var. albus from France
= juvenile
2011.08.29 © Lebeaux, P.
Sminthurinus aureus var. albus from the UK
= juvenile
2014.01.08 © Phillips, E.

            Syn.: Sminthurinus aureus var. atrata(sic) Börner, C, 1901:104 IOS
            Syn.: Sminthurinus aureus var. atrata(sic) Börner, 1901 nn
             Syn.: Sminthurinus aureus var. atrata(sic) Axelson, 1904 ISS
             Syn.: Sminthurinus aureus var. atrata(sic) Linnaniemi, WM, 1912:288 ISS
             Syn.: Sminthurinus aureus f. atrata(sic) Handschin, E in Dahl, F, 1929:114 ISS
             Syn.: Sminthurinus aureus var. atrata(sic) Stach, J, 1956:122 ISS
            Syn.: Sminthurinus aureus ssp. atratus Salmon, JT, 1964:578
            Syn.: Sminthurus aureus var. fusca(sic) Salmon, JT, 1964:578 partim = atratus IOS
             Syn.: Sminthurinus aureus var. atrata(sic) Ellis, WN, 1974:131 ISS
             Syn.: Sminthurinus aureus var. atrata(sic) Bretfeld, G, 1999:110 ISS
Sminthurinus aureus f. atratus from the UK
With dark legs
2013.11.05 © Nurcombe, E.
Sminthurinus aureus f. atratus from the UK
With dark legs
2014.11.21 © Phillips, E.
Sminthurinus aureus f. atratus from Holland
2018.09.06 © van Duinen, J.
Sminthurinus aureus f. atratus from Hungary
With dark appendages
2018.09.28 © Ujvári, Z.
Sminthurinus aureus f. atratus from the UK
With dark appendages
2018.10.16 © Phillips, E.
Sminthurinus aureus f. atratus from Belgium
Tripartite eyebrow
Dark/yellow/white
2020.07.17 © Huskens, M.-L.
Sminthurinus aureus f. atratus from the UK
With dark appendages
2020.09.01 © Phillips, E.
Sminthurinus aureus f. atratus from Belgium
2020.11.06 © Huskens, M.-L.
Sminthurinus aureus f. atratus from Belgium
Grooming
2020.11.06 © Huskens, M.-L.
Sminthurinus aureus f. atratus from Belgium
Ant.3 with wart (W)
2020.11.06 © Huskens, M.-L.
Sminthurinus aureus f. atratus from Belgium
Ocellus C indistinct, D small
2020.11.06 © Huskens, M.-L.
Sminthurinus aureus f. atratus from Belgium
Eyebrow tripartite
2021.06.01 © Huskens, M.-L.
Sminthurinus aureus f. atratus from Belgium
2021.11.24 © Huskens, M.-L.
Sminthurinus aureus f. atratus from the UK
2022.10.25 © Phillips, E.
Sminthurinus aureus f. atratus from the UK
Eyebrow with dark transverse suture at ocellus H
2022.10.25 © Phillips, E.
Sminthurinus aureus f. atratus from Belgium
Lowest part of eyebrow dark
2022.10.25 © Huskens, M.-L.
Sminthurinus aureus f. atratus juv. from the UK
Eyebrow with dark transverse suture at ocellus H
2022.11.28 © Phillips, E.
Sminthurinus aureus f. atratus from the UK
2022.11.28 © Phillips, E.

            Syn.: Sminthurus aureus var. maculata(sic) Krausbauer, 1901 IOS
             Syn.: Sminthurinus aureus var. maculata(sic) Börner, C, 1901:138 ISS
             Syn.: Sminthurinus aureus var. maculata(sic) Linnaniemi, 1907 ISS
             Syn.: Sminthurus aureus var. maculata(sic) Linnaniemi, WM, 1912:285 ISS
             Syn.: Sminthurinus aureus var. maculata(sic) Linnaniemi, WM, 1912:285 ISS
             Syn.: Sminthurinus aureus var. maculata(sic) Denis, 1922 ISS
             Syn.: Sminthurinus aureus f. maculata(sic) Handschin, E in Dahl, F, 1929:114 ISS
             Syn.: Sminthurinus aureus var. maculata(sic) Stach, J, 1956:122 ISS
             Syn.: Sminthurus aureus ssp. maculata(sic) Salmon, JT, 1964:578 = atratus ISS
             Syn.: Sminthurinus aureus ssp. maculata(sic) Salmon, JT, 1964:578 = atratus ISS
             Syn.: Sminthurinus aureus f. maculata(sic) Stach, J, 1964:58 ISS
Sminthurinus aureus f. maculatus from France
2008.03.24 © Lebeaux, P.
Sminthurinus aureus f. maculatus from France
2008.03.24 © Lebeaux, P.
Sminthurinus aureus f. maculatus from the UK
2012.07.22 © Murray, A.
Sminthurinus aureus f. maculatus from the UK
2012.07.22 © Murray, A.
Sminthurinus aureus f. maculatus from Holland
Preparing to moult
2012.11.12 © van Duinen, J.
Sminthurinus aureus f. maculatus from the UK
2012.12.21 © Phillips, E.
Sminthurinus aureus f. maculatus from Holland
2013.10.23 © van Duinen, J.
Sminthurinus aureus f. maculatus from Holland
2013.10.23 © van Duinen, J.
Sminthurinus aureus f. maculatus from the UK
Freeing itself from a silk fiber
2014.01.17 © Phillips, E.
Sminthurinus aureus f. maculatus from Germany
2014.11.01 © Kluck, A.
Sminthurinus aureus f. maculatus from Germany
2014.11.01 © Kluck, A.
Sminthurinus aureus f. maculatus from Germany
2014.11.01 © Kluck, A.
Sminthurinus aureus f. maculatus from Holland
2014.11.17 © van Duinen, J.
Sminthurinus aureus f. maculatus from Holland
Colour form inbetween f. maculatus and f. reticulatus
2014.11.21 © van Duinen, J.
Sminthurinus aureus f. maculatus from Holland
Colour form inbetween f. maculatus and f. reticulatus
2014.11.21 © van Duinen, J.
Sminthurinus aureus f. maculatus from Belgium
2016.12.09 © Huskens, M.-L.
Sminthurinus aureus f. maculatus from Germany
2017.11.20 © Kluck, A.
Sminthurinus aureus f. maculatus from the UK
2018.12.11 © Cann, A.J.
Sminthurinus aureus f. maculatus from the UK
2018.12.11 © Cann, A.J.
Sminthurinus aureus f. maculatus from Belgium
2018.12.14 © Huskens, M.-L.
Sminthurinus aureus f. maculatus from Belgium
Caudal aspect
2018.12.14 © Huskens, M.-L.
Sminthurinus aureus f. maculatus from the UK
2020.10.14 © Phillips, E.
Sminthurinus aureus f. maculatus from the UK
Caudal aspect
2020.10.14 © Phillips, E.
Sminthurinus aureus f. maculatus from Belgium
2022.12.22 © Huskens, M.-L.
Sminthurinus aureus f. maculatus from Belgium
Caudal mimicry
2022.12.22 © Huskens, M.-L.
Sminthurinus aureus f. maculatus from Belgium
2022.12.28 © Huskens, M.-L.
Sminthurinus aureus f. maculatus from Belgium
2022.12.28 © Huskens, M.-L.


            Syn.: Sminthurinus aureus var. punctata Börner, C, 1901:103
            Syn.: ... pustulata Krausbauer, 1902
            Syn.: Sminthurinus niger Linnaniemi, WM, 1912:289
            Syn.: Sminthurinus aureus var. punctiformis Latzel, R, 1917:249
            Syn.: ... anglicana Salmon, 1955
            Syn.: Sminthurinus igniceps da Gama, 1959
            Syn.: Sminthurinus aureus f. reticulatus Janssens F & Phillips E, 2013
Sminthurinus aureus f. reticulatus from the UK
2012.11.19 © Phillips, E.
Sminthurinus aureus f. reticulatus from the UK
2012.12.03 © Phillips, E.
Sminthurinus aureus f. reticulatus from the UK
2012.12.04 © Phillips, E.
Sminthurinus aureus f. reticulatus from Switzerland
2012.12.06 © Murray, A.
Sminthurinus aureus f. reticulatus from the UK
2012.12.21 © Phillips, E.
Sminthurinus aureus f. reticulatus from the UK
2013.01.28 © Phillips, E.
Sminthurinus aureus f. reticulatus from the UK
2013.02.12 © Phillips, E.
Sminthurinus aureus f. reticulatus juv. from the UK
2013.11.30 © Nurcombe, E.
Sminthurinus aureus f. reticulatus juv. from the UK
Mottled appearance: ready to moult
2013.11.30 © Nurcombe, E.
Sminthurinus aureus f. reticulatus ♂ from the UK
Raised genital papilla (*)
2013.11.30 © Nurcombe, E.
Sminthurinus aureus f. reticulatus from the UK
2014.01.17 © Phillips, E.
Sminthurinus aureus f. reticulatus from the UK
2014.01.25 © Phillips, E.
Sminthurinus aureus f. reticulatus from the UK
Feeding instar
2014.02.06 © Phillips, E.
Sminthurinus aureus f. reticulatus from Holland
2014.10.31 © van Duinen, J.
Sminthurinus aureus f. reticulatus from Holland
2014.11.23 © van Duinen, J.
Sminthurinus aureus f. reticulatus from Belgium
2016.12.09 © Huskens, M.-L.
Sminthurinus aureus f. reticulatus juv. from Belgium
2016.12.20 © Huskens, M.-L.
Sminthurinus aureus f. reticulatus ♂ from Belgium
2018.10.18 © Huskens, M.-L.
Sminthurinus aureus f. reticulatus from the UK
Preparing to moult
2018.12.31 © Phillips, E.
Sminthurinus aureus f. reticulatus from Belgium
Camouflage in autumn
2019.10.05 © Huskens, M.-L.
Sminthurinus aureus f. reticulatus juv. from Holland
2019.10.21 © Kamsteeg, G.
Sminthurinus aureus f. reticulatus from Holland
2019.11.23 © Kamsteeg, G.
Sminthurinus aureus f. reticulatus from Holland
2019.11.23 © Kamsteeg, G.


            Syn.: Sminthurinus aureus f. apigmentatus Janssens F & Nurcombe E, 2013
Sminthurinus aureus f. apigmentatus ♂ from the UK
Form without black pigment having red eyes
2013.11.24 © Nurcombe, E.
Sminthurinus aureus f. apigmentatus ♂ from the UK
Form without black pigment having red eyes
2013.11.24 © Nurcombe, E.
Sminthurinus aureus f. apigmentatus from Holland
2019.10.20 © Kamsteeg, G.
Sminthurinus aureus f. apigmentatus from Holland
Grooming tripod posture
2019.10.20 © Kamsteeg, G.

            Syn.: Sminthurinus aureus f. cyaneus Janssens F & Huskens M-L, 2018
Sminthurinus aureus f. cyaneus from Holland
Abdominal bothriotricha (A-D)
2014.10.09 © van Duinen, J.
Sminthurinus aureus f. cyaneus from Holland
2014.10.09 © van Duinen, J.
Sminthurinus aureus f. cyaneus from Belgium
2018.08.16 © Huskens, M.-L.
Leg.: Janssens, F.
Sminthurinus aureus f. cyaneus from Belgium
2018.08.16 © Huskens, M.-L.
Leg.: Janssens, F.
Sminthurinus aureus f. cyaneus from Belgium
2018.08.16 © Huskens, M.-L.
Leg.: Janssens, F.
Sminthurinus aureus f. cyaneus juv. from Belgium
2018.08.16 © Huskens, M.-L.
Leg.: Janssens, F.
Sminthurinus aureus from Belgium
Reticulate skin suggests specimen is preparing to moult
2018.08.16 © Huskens, M.-L.
Sminthurinus aureus f. cyaneus ♂ & ♀ from Belgium
2018.09.17 © Huskens, M.-L.
Sminthurinus aureus f. cyaneus from Belgium
2018.08.18 © Huskens, M.-L.
Sminthurinus aureus f. cyaneus from Belgium
2018.09.16 © Huskens, M.-L.
Sminthurinus aureus f. cyaneus ♂ from Belgium
2018.09.17 © Huskens, M.-L.
Sminthurinus aureus f. cyaneus from Belgium
2019.05.19 © Huskens, M.-L.
Sminthurinus aureus f. cyaneus from Belgium
2019.05.19 © Huskens, M.-L.
Sminthurinus aureus f. cyaneus from Belgium
2019.11.04 © Huskens, M.-L.
Sminthurinus aureus f. cyaneus from Belgium
2019.11.04 © Huskens, M.-L.
Sminthurinus aureus f. cyaneus from Belgium
Preparing to moult
2019.11.04 © Huskens, M.-L.
Sminthurinus aureus f. cyaneus from Belgium
2020.10.25 © Huskens, M.-L.
Sminthurinus aureus f. cyaneus from Belgium
2020.10.26 © Huskens, M.-L.
Sminthurinus aureus f. cyaneus from Belgium
Abd.5 with lateral bothriotrichum D on papilla
2020.10.26 © Huskens, M.-L.
Sminthurinus aureus f. cyaneus from Belgium
2021.01.12 © Huskens, M.-L.
Sminthurinus aureus f. cyaneus from Belgium
2021.01.12 © Huskens, M.-L.
Sminthurinus aureus f. cyaneus from Belgium
2021.01.12 © Huskens, M.-L.
Sminthurinus aureus f. cyaneus from Belgium
2021.06.02 © Huskens, M.-L.
Sminthurinus aureus ?f. cyaneus? from Belgium
2024.03.15 © Huskens, M.-L.
Sminthurinus aureus ?f. cyaneus? from Belgium
2024.03.20 © Huskens, M.-L.




Smynthurus(sic) aureus from the UK
After Lubbock J, 1873 Pl.7
Sminthurus aureus from the UK
After Lubbock J in Collinge WE & Shoebotham JW, 1909 Pl.III Fig.2
Sminthurinus aureus from the UK
Ventral subapical dental setae absent
2004 © Hopkin, S.P.
Sminthurinus aureus
2006 © Boldsystems.org
 
Sminthurinus aureus from France
2007.03.17 © Lebeaux, P.
Sminthurinus aureus from the Netherlands
2009.01.26 © Heemskerk, R.
Sminthurinus aureus from France
2009.09.dd © Lebeaux, P.
Sminthurinus aureus from France
2009.09.dd © Lebeaux, P.
Sminthurinus aureus from France
2009.09.dd © Lebeaux, P.
Sminthurinus aureus from France
2009.09.16 © Lebeaux, P.
Sminthurinus aureus from Czechia
2010.11.25 © Fiala, M.
Sminthurinus aureus from Germany
2011.05.04 © Marquard, F.
Sminthurinus aureus from France
2011.11.13 © Lebeaux, P.
Sminthurinus aureus from France
2011.11.13 © Lebeaux, P.
Sminthurinus aureus from France
2011.11.14 © Lebeaux, P.
Sminthurinus aureus (bar=2µm)
Hexagonal epicuticular ultrastructure
After Nickerl J & al, 2012 Fig.6C
Sminthurinus aureus from the UK
2012.01.06 © Jones, A.
Sminthurinus aureus from the UK
2012.01.06 © Jones, A.
Sminthurinus aureus from the UK
2012.01.29 © Nurcombe, E.
Sminthurinus aureus juv. from the UK
2012.01.29 © Nurcombe, E.
Sminthurinus aureus from the UK
2012.04.10 © Nurcombe, E.
Sminthurinus aureus from the UK
2012.06.14 © Murray, A.
Sminthurinus aureus from the UK
2012.02.16 © Nurcombe, E.
Sminthurinus aureus from the UK
2012.06.18 © Murray, A.
Sminthurinus aureus juv. from the UK
2012.08.01 © Murray, A.
Sminthurinus aureus from the UK
Lateral abdominal bothriotricha
2012.10.26 © Nurcombe, E.
Sminthurinus aureus from the UK
4th antennal segment not subdivided
2012.10.26 © Nurcombe, E.
Sminthurinus aureus from the UK
2012.10.29 © Jones, A.
Sminthurinus aureus from the UK
2012.11.15 © Kent, J.J.
Sminthurinus aureus from the UK
2012.11.25 © Nurcombe, E.
Sminthurinus aureus from the UK
2012.11.26 © Kent, J.J.
Sminthurinus aureus from the UK
2012.11.26 © Phillips, E.
Sminthurinus aureus from Switzerland
2012.12.06 © Murray, A.
Sminthurinus aureus from the UK
2012.12.07 © Phillips, E.
Sminthurinus aureus-group s.l. from the UK
Forma aureus (1,2,4,6,8,10), maculatus (12), reticulatus (3,5,7,9,11)
2012.12.24 © Phillips, E.
Sminthurinus aureus from Holland
2013.02.16 © van Duinen, J.
Sminthurinus aureus from the UK
2013.04.30 © Murray, A.
Sminthurinus aureus from the UK
2013.04.30 © Murray, A.
Sminthurinus aureus from the UK
Feeding instar with dark gut contents
2013.10.04 © Phillips, E.
Sminthurinus aureus from the UK
Dark spots mark positions of 4 abdominal bothriotricha (A-D)
2013.10.08 © Phillips, E.
Sminthurinus aureus from France
2014.01.04 © Sandra.
Sminthurinus aureus from Germany
2014.11.15 © Kluck, A.
Sminthurinus aureus from Germany
Walking in a row
2015.01.07 © Landgraf, A.
Sminthurinus aureus juv. from Belgium
Distinctly separate thoracic segments
2016.01.26 © Huskens, M.-L.
Sminthurinus aureus from the UK
Bothriotricha A-B-C-D
2016.03.10 © Phillips, E.
Sminthurinus aureus ♂ from Belgium
2017.11.20 © Huskens, M.-L.
Sminthurinus aureus ♀ from Holland
Ventral aspect
2017.12.28 © Kamsteeg, G.
Sminthurinus aureus from Belgium
2018.05.05 © Kamsteeg, G.
Sminthurinus aureus from Switzerland
2018.12.15 © Willi, R.
Sminthurinus aureus from the UK
Ocelli C+D reduced in size
2018.12.26 © Cann, A.J.
Sminthurinus aureus from France
2019.02.06 © Garcelon, P.
Sminthurinus aureus from Japan
Preparing to moult
2019.02.22 © Hara, Y.
Sminthurinus aureus from France
Walking
Front- and hindleg lifted up
2021.01.02 © Fiacre, R.
Sminthurinus aureus juv. from Belgium
2021.01.11 © Huskens, M.-L.
Sminthurinus aureus juv. from Belgium
2021.01.11 © Huskens, M.-L.
Sminthurinus aureus adult from Belgium
2021.01.11 © Huskens, M.-L.
Sminthurinus aureus adult from Belgium
2021.01.11 © Huskens, M.-L.
Sminthurinus aureus from Belgium
Dorsolateral fat tissue
2021.01.19 © Huskens, M.-L.
Sminthurinus aureus from Belgium
Dorsolateral fat tissue
2021.01.19 © Huskens, M.-L.
Sminthurinus aureus ♀ & ♂ from Belgium
2023.02.15 © Huskens, M.-L.
Sminthurinus aureus from France
Knuckle walking
Foot with clavate tenent setae
2023.11.19 © Clerc, O.

  Subspecies purpureus Salmon, JT, 1941:410



Species aureussetosus Bretfeld, 2002

Species australis Najt, 1973

Species bimaculatus Axelson, 1902

            Syn.: Sminthurinus igniceps var. bimaculata(sic) Axelson, 1902 IOS
            Syn.: Sminthurus biflavopunctatus Linnaniemi, 1912
            Syn.: Sminthurinus aureus f. biflavopunctata Handschin, E in Dahl, F, 1929:114
            Syn.: Sminthurinus biflavopunctatus Womersley, 1930
            Syn.: Sminthurinus biflavopunctatus Davies, 1934
            Syn.: Sminthurinus biflavopunctatus Kloet & Hincks, 1934
            Syn.: Sminthurinus aureus f. biflavopunctatus Uchida, 1957
            Syn.: Sminthurus biflavopunctatus Hopkin, 2000
            Syn.: Sminthurinus aureus var. circumfasciata Stach, 1922
            Syn.: Sminthurinus aureus f. fenestratus Handschin, E in Dahl, F, 1929:114



            Syn.: Sminthurinus igniceps var. bimaculata(sic) Collinge, WE et Shoebotham, JW, 1909:10 IOS
            Syn.: Sminthurinus aureus var. bimaculata(sic) Schött, H, 1917:55 IOS
            Syn.: Sminthurinus aureus f. bimaculata(sic) Handschin, E in Dahl, F, 1929:114 IOS
Sminthurinus bimaculatus from France
2005 © Losacco, S.
Sminthurinus bimaculatus from France
2005 © Losacco, S.
Sminthurinus bimaculatus from France
2005 © Losacco, S.
Sminthurinus bimaculatus from the UK
2008.06.06 © Barton, T.
Sminthurinus bimaculatus from Holland
2012.07.11 © van Duinen, J.
Sminthurinus bimaculatus from Holland
2012.07.11 © van Duinen, J.
Sminthurinus bimaculatus from the UK
2012.07.15 © Murray, A.
Sminthurinus bimaculatus juv. from the UK
2012.07.15 © Murray, A.

            Syn.: Sminthurinus igniceps var. bimaculata(sic) Mehrafrooz Mayvan, M, Sadeghi-Namaghi, H et Shayanmehr, M, 2021:7 IOS
Sminthurinus nr bimaculatus juv. from Taiwan
2021.11.10 © Cheng, H.-J.
Sminthurinus nr bimaculatus juv. from Taiwan
Grooming
2021.11.13 © Cheng, H.-J.
Sminthurinus nr bimaculatus from Taiwan
Pressing collophore vesicles in surface waterfilm
2021.11.15 © Cheng, H.-J.
Sminthurinus nr bimaculatus from Taiwan
2021.11.16 © Cheng, H.-J.
Sminthurinus nr bimaculatus from Taiwan
2021.11.16 © Cheng, H.-J.
Sminthurinus nr bimaculatus from Taiwan
2021.11.16 © Cheng, H.-J.
Sminthurinus nr bimaculatus from Taiwan
Facial pattern
2021.11.16 © Cheng, H.-J.
Sminthurinus nr bimaculatus from India
After Mandal P & al, 2024 Fig.6A



Species bisetosus Ellis, 1976

Species bullai Najt, 1972

Species cantonensis Rusek, 1971

Species castagninoi Najt, 1969

Species cingulatus Bagnall, 1921

             Syn.: Sminthurinus cingula(sic) Gisin, 1960 sd ISS

Species conchyliatus Snider, 1978

Sminthurinus conchyliatus? from the USA
After Dukes CD & al, 2022 Fig.5F

Species concolor (Meinert, 1896) Jorgensen, 1934

            Syn.: Sminthurinus aureus Fjellberg, 1984
            Syn.: Sminthurinus niger Fjellberg, 1984


Sminthurinus concolor from Groenland
2 subapical anterior dental setae present (*)
Proximal posterior setae: 2 outer, 3 inner
Modified after Gisin H, 1963 Fig.4
Sminthurinus concolor from the UK
Two setae on retinaculum
2004 © Hopkin, S.P.
Sminthurinus concolor from Luxembourg
After Stomp N & Weiner WM, 2005 Fig.5
Sminthurinus concolor from Norway
2011 © Fjellberg, A.
Sminthurinus concolor from the UK
2012.08.21 © Murray, A.
Det.: Shaw, P.
Sminthurinus concolor from the UK
2012.08.21 © Murray, A.
Det.: Shaw, P.
Sminthurinus concolor juv. from the UK
2012.08.22 © Murray, A.
Det.: Shaw, P.
Sminthurinus concolor from Norway
2015 © Fjellberg, A.
Sminthurinus concolor from the UK
2018.04.08 © Robson, S.
Sminthurinus concolor from France
2018.12.29 © Picard, J.
Sminthurinus concolor from France
Eyebrow as long as ocelli C to G
Eyebrow not touching ocellus C
2018.12.29 © Picard, J.
Sminthurinus concolor from France
Inter eyebrows distance less than 2x width of eyebrow
2018.12.29 © Picard, J.
Sminthurinus concolor ♀ from the UK
2021.02.06 © Robson, S.
Sminthurinus concolor ♀ from the UK
Elongate eyebrows
2021.02.06 © Robson, S.
Sminthurinus concolor ♀ from the UK
Ratio dens:mucro = 1.8
2021.02.06 © Robson, S.
Sminthurinus concolor ♀ from the UK
Sub-anal appendage (saa)
2021.02.06 © Robson, S.
Sminthurinus concolor from the UK
2 dental anterior subapical setae
2021.05.28 © McCulloch, J.
Leg.: Robson, S.
Sminthurinus concolor from France
2021.12.31 © Usami, R.
Sminthurinus concolor from France
2021.12.31 © Usami, R.
Sminthurinus concolor from Holland
Pointed eyebrow
not touching ocellus C
2022.02.08 © Kamsteeg, G.
Sminthurinus concolor from Holland
2022.02.08 © Kamsteeg, G.

Species discordipes Salmon, JT, 1949:49

Species dispar Babenko, AB, 2007:375

Species distinctus Kang J-S et Lee B-H in Kang, J-S, Lee, B-H et Park, K-H, 2005:240

Sminthurinus distinctus from Korea
After Kang J-S, Lee B-H & Park K-H, 2005 Fig.4A

Species domesticus Gisin, H, 1963:84

Sminthurinus domesticus from Portugal
2 subapical anterior dental setae present (*)
Proximal posterior setae: 1 outer, 1 median, 2 inner
Modified after Gisin H, 1963 Fig.5
Sminthurinus domesticus from Portugal
Ocellus D smaller
1 seta inbetween ocelli H and G
After Gisin H, 1963 Fig.5

            Syn.: Sminthurinus niger da Gama, 1959 partim
            Syn.: Sminthurinus niger da Gama, 1961 partim
            Syn.: Sminthurinus niger Gisin, H, 1963:80 partim
            Syn.: Sminthurinus domesticus f. apigmentatus Janssens F & van Duinen J, 2012
Sminthurinus domesticus f. apigmentatus from Holland
Form without black pigment having red eyes
2012.07.11 © van Duinen, J.
Sminthurinus domesticus f. apigmentatus from Holland
Form without black pigment having red eyes
2013.08.18 © van Duinen, J.
Sminthurinus domesticus f. apigmentatus from Holland
Form without black pigment having red eyes
2013.09.03 © van Duinen, J.
Sminthurinus domesticus f. apigmentatus from Holland
Form without black pigment having red eyes
2013.09.03 © van Duinen, J.




Sminthurinus domesticus from the UK
Two subapical ventral dental setae
2004 © Hopkin, S.P.
Sminthurinus domesticus from the UK, juvenile
2007.09.29 © Valentine, Br.
Sminthurinus domesticus from the UK
2007.10.13 © Valentine, Br.
Sminthurinus domesticus from the UK
2007.10.13 © Valentine, Br.
Sminthurinus domesticus from the UK
2007.10.18 © Valentine, Br.
Sminthurinus domesticus from the UK
2007.10.24 © Valentine, Br.
Sminthurinus domesticus from the UK
2007.10.25 © Valentine., B
Sminthurinus domesticus from the UK
2007.10.25 © Valentine, Br.
Sminthurinus domesticus from the UK
2007.11.15 © Valentine, Br.
Sminthurinus domesticus from the UK
2008.08.27 © Valentine, Br.
Sminthurinus domesticus from the UK
2008.10.16 © Valentine, Br.
Sminthurinus domesticus from the UK
2008.10.16 © Valentine, Br.
Sminthurinus domesticus from the UK
2008.11.18 © Valentine., B
Sminthurinus domesticus from the UK
2008.11.18 © Valentine, Br.
Sminthurinus domesticus from the UK
2008.12.05 © Valentine, Br.
Sminthurinus domesticus from the UK
2008.12.05 © Valentine, Br.
Sminthurinus domesticus from the UK
After Ardron PA, 2009 p.18
Sminthurinus domesticus from the UK
2009.11.07 © Valentine, Br.
Sminthurinus domesticus from the UK
2009.11.07 © Valentine, Br.
Sminthurinus domesticus from the UK
Premoult reticulate skin
2010.01.19 © Jones, A.
Sminthurinus domesticus from Holland
2011.09.11 © van Duinen, J.
Sminthurinus domesticus from Holland
2011.11.08 © van Duinen, J.
Sminthurinus domesticus from Holland
2011.11.08 © van Duinen, J.
Sminthurinus domesticus from Holland
2011.11.08 © van Duinen, J.
Sminthurinus ?domesticus? from the UK
2012.06.13 © Murray, A.
Sminthurinus ?domesticus? from the UK
2012.06.13 © Murray, A.
Sminthurinus ?domesticus? from the UK
2012.06.13 © Murray, A.
Sminthurinus domesticus from Holland
2012.07.11 © van Duinen, J.
Sminthurinus domesticus from Holland
Pale colour form
2012.07.11 © van Duinen, J.
Sminthurinus domesticus from the UK
2012.08.30 © Murray, A.
Sminthurinus domesticus from the UK
Face
2012.08.30 © Murray, A.
Sminthurinus domesticus from the UK
Pale colour form
2012.08.30 © Murray, A.
Sminthurinus domesticus early instars from Portugal
2012.10.11 © Murray, A.
Sminthurinus domesticus juv. from Portugal
2012.10.11 © Murray, A.
Sminthurinus domesticus from Portugal
2012.10.11 © Murray, A.
Sminthurinus domesticus from the UK
2013.01.04 © Murray, A.
Sminthurinus domesticus from the UK
Asymmetric aberration in frontocellar area
2013.01.04 © Murray, A.
Sminthurinus domesticus from Morocco
2014.04.21 © Beck, K.
Sminthurinus domesticus juv. from Morocco
2014.04.28 © Beck, K.
Sminthurinus domesticus from Morocco
2014.04.28 © Beck, K.
Sminthurinus domesticus from Morocco
2014.04.28 © Beck, K.
Sminthurinus domesticus from Morocco
Circumanal setae with expanded base (*)
2014.04.28 © Beck, K.
Sminthurinus domesticus from Morocco
Circumanal median seta bifid (*)
2014.04.28 © Beck, K.
Sminthurinus domesticus from Morocco
Secreting a drop of saliva
2014.04.28 © Beck, K.
Sminthurinus domesticus from Morocco
Picking up drop of saliva with proclaw
2014.04.28 © Beck, K.
Sminthurinus domesticus from Morocco
Cleaning antenna with drop of saliva
2014.04.28 © Beck, K.
Sminthurinus domesticus from Morocco
2014.04.29 © Beck, K.
Sminthurinus domesticus from Morocco
2014.04.30 © Beck, K.
Sminthurinus domesticus from Morocco
2014.04.30 © Beck, K.
Sminthurinus domesticus juv. from Germany
2016.02.06 © Gutekunst, V.
Sminthurinus domesticus juv. from Germany
2016.02.06 © Gutekunst, V.
Sminthurinus domesticus juv. from Germany
2016.02.06 © Gutekunst, V.
Sminthurinus domesticus from Germany
2017.01.28 © Melzer, L.
Sminthurinus domesticus juv. from Belgium
Premoult reticulate skin
2019.01.17 © Huskens, M.-L.
Sminthurinus domesticus juv. from Belgium
2019.01.20 © Huskens, M.-L.
Sminthurinus domesticus juv. from Belgium
Pale colour form
2019.01.20 © Huskens, M.-L.
Sminthurinus domesticus juv. from Belgium
Bipartite + bicoloured 'eyebrow'
2019.01.20 © Huskens, M.-L.
Sminthurinus domesticus juv. from Belgium
Bipartite + bicoloured 'eyebrow'
2019.01.20 © Huskens, M.-L.
Sminthurinus domesticus from Belgium
2019.01.21 © Huskens, M.-L.
Sminthurinus domesticus from Belgium
2019.01.21 © Huskens, M.-L.
Sminthurinus domesticus from Belgium
2019.01.24 © Huskens, M.-L.
Sminthurinus domesticus ♀ from Belgium
Sub-anal appendages (*)
2019.01.24 © Huskens, M.-L.
Sminthurinus domesticus ♂ from Belgium
2019.01.26 © Huskens, M.-L.
Sminthurinus domesticus from Belgium
2019.02.26 © Huskens, M.-L.
Sminthurinus domesticus from Belgium
2019.02.26 © Huskens, M.-L.
Sminthurinus domesticus from Belgium
Bipartite bicoloured 'eyebrow'
Eyepatch with 8 ocelli; D smallest
2019.02.26 © Huskens, M.-L.
Sminthurinus domesticus ♂ from Belgium
Depositing spermatophore
2019.02.27 © Huskens, M.-L.
Sminthurinus domesticus ♀ from Belgium
2019.02.28 © Huskens, M.-L.
Sminthurinus domesticus ♀ & ♂ from Belgium
Failed sperm transfer
1. Male approaches female
2019.03.02 © Huskens, M.-L.
Sminthurinus domesticus ♀ & ♂ from Belgium
Failed sperm transfer
2. Male deposits spermatophore near to female
2019.03.02 © Huskens, M.-L.
Sminthurinus domesticus ♀ & ♂ from Belgium
Failed sperm transfer
3. Female is not interested...
2019.03.02 © Huskens, M.-L.
Sminthurinus domesticus from Belgium
2019.03.13 © Huskens, M.-L.
Sminthurinus domesticus from Belgium
2019.03.13 © Huskens, M.-L.
Sminthurinus domesticus from the UK
2019.10.24 © Valentine, Br.
Sminthurinus domesticus from Portugal
2020.01.07 © Viana, J.
Sminthurinus domesticus from Portugal
2020.01.07 © Viana, J.
Sminthurinus domesticus from Portugal
Ant.3 with wart (*)
2020.01.07 © Viana, J.
Sminthurinus domesticus from Portugal
Grooming
2020.01.07 © Viana, J.
Sminthurinus domesticus from the UK
Eyebrow bicoloured
2020.10.10 © Valentine, Br.
Sminthurinus domesticus from the UK
2021.03.10 © David W.
Sminthurinus domesticus from the UK
Eyebrow bipartite
Upper part white, lower part reddish
2021.03.10 © David W.
Sminthurinus domesticus ♀ from Belgium
2023.08.31 © Huskens, M.-L.
Sminthurinus domesticus ♀ from Belgium
Bothriotrichum A
2023.08.31 © Huskens, M.-L.
Sminthurinus domesticus ♀ from Belgium
Third footcomplex with clavate tenent macrosetae
2023.08.31 © Huskens, M.-L.
Sminthurinus domesticus ♀ from Belgium
Circumanal setae
Median seta bifurcate
2023.08.31 © Huskens, M.-L.
Sminthurinus domesticus ♀ from Belgium
Sub-anal appendage on basal papilla
2023.08.31 © Huskens, M.-L.
Sminthurinus domesticus from Belgium
2023.09.01 © Huskens, M.-L.
Sminthurinus domesticus ♂ from Belgium
2023.09.02 © Huskens, M.-L.
Sminthurinus domesticus ♀ from Belgium
Ready to jump
With everted collophore vesicles
2023.09.03 © Huskens, M.-L.
Sminthurinus domesticus from Belgium
2023.09.06 © Huskens, M.-L.
Sminthurinus domesticus ♀ from Belgium
Bipartite bicoloured eyebrows
2023.09.07 © Huskens, M.-L.
Sminthurinus domesticus ♀ from Belgium
With eggs covered by faeces
2023.09.14 © Huskens, M.-L.
Sminthurinus domesticus from Belgium
2023.09.25 © Huskens, M.-L.
Sminthurinus domesticus from Belgium
Specimen with distinctly pale head
2023.10.13 © Huskens, M.-L.
Sminthurinus domesticus from Belgium
2023.10.15 © Huskens, M.-L.

Species duplicatus Salmon, JT, 1941:411

Species elegans (Fitch, A, 1863:674) Folsom, 1928, B

            Syn.: Symnthurus(sic) elegans var. a Fitch, A, 1863:674
            Syn.: Symnthurus(sic) elegans var. b Fitch, A, 1863:674
            Syn.: Smynthurus(sic) signifer Schött, H, 1891:13 =?
            Syn.: Sminthurus quadrilineatus Tullberg, 1871, partim
             Syn.: Sminthurus 4-lineatus(sic) Reuter, OM in Lindberg, SO, 1881:204 ISS
             Syn.: Sminthurus qvadrilineatus(sic) Reuter, OM, 1892:17 ISS
Sminthurus quadrilineatus from Sweden
After Schött H, 1893 Taf.I Fig.18

            Syn.: Sminthurus (Eusminthurus) aureus var. ornata(sic) Börner, 1900 IOS
             Syn.: Sminthurinus aureus var. ornata(sic) Börner, C, 1901:104 ISS
             Syn.: Sminthurinus aureus var. ornata(sic) Linnaniemi, WM, 1912:285 ISS
            Syn.: Sminthurinus elegans Maynard, EA, 1951:237
Sminthurinus elegans from the USA
After Maynard EA, 1951 Plate XXVIII Fig.520
=? ornatus Janssens F, 2017

             Syn.: Sminthurinus quadrilineata(sic) ab. ornata(sic) Stach, J, 1956:122 ISS
             Syn.: Sminthurinus aureus var. ornata(sic) Stach, J, 1956:122 ISS
             Syn.: Sminthurinus aureus var. quadrilineata(sic) ab. ornata(sic) Stach, J, 1956:122 ISS
             Syn.: Sminthurinus elegans var. ornata(sic) Stach, J, 1956:127 ISS
             Syn.: Sminthurus aureus var. ornata(sic) Gisin, H, 1960:278 ISS
             Syn.: Sminthurus aureus ssp. ornata(sic) Salmon, JT, 1964:579 ISS
             Syn.: Sminthurinus aureus ssp. ornata(sic) Salmon, JT, 1964:579 ISS
             Syn.: Sminthurinus aureus ssp. ornata(sic) Salmon, JT, 1964:580 ISS
             Syn.: Sminthurus aureus ssp. ornatous(sic) Salmon, JT, 1964:580 ISS
             Syn.: Sminthurinus aureus ssp. ornata(sic) Salmon, JT, 1964:582 ISS
Sminthurinus elegans f. ornatus from the UK
2008.12.09 © Valentine., B
Sminthurinus elegans f. ornatus from the UK
2008.12.09 © Valentine., B
Sminthurinus elegans f. ornatus from the UK
2008.12.09 © Valentine., B
Sminthurinus elegans f. ornatus from the USA
2009.11.25 © Gledhill, L.
Sminthurinus elegans f. ornatus juv. from the UK
2012.06.10 © Murray, A.
Sminthurinus elegans f. ornatus from the UK
2012.06.11 © Murray, A.
Sminthurinus elegans f. ornatus from the UK
2012.06.11 © Murray, A.
Sminthurinus elegans f. ornatus juv. from the UK
2012.06.13 © Murray, A.
Sminthurinus elegans f. ornatus from the UK
2012.06.14 © Murray, A.
Sminthurinus elegans f. ornatus from the UK
2012.06.14 © Murray, A.
Sminthurinus elegans f. ornatus from the UK
2012.06.14 © Murray, A.
Sminthurinus elegans f. ornatus from the UK
2012.06.14 © Murray, A.
Sminthurinus elegans f. ornatus from the UK
2012.06.14 © Murray, A.
Sminthurinus elegans f. ornatus from the UK
Grooming
2012.08.16 © Nurcombe, E.
Sminthurinus elegans f. ornatus from the UK
2012.08.16 © Nurcombe, E.
Sminthurinus elegans f. ornatus from the UK
2012.08.16 © Nurcombe, E.
Sminthurinus elegans f. ornatus from the UK
2012.08.17 © Nurcombe, E.
Sminthurinus elegans f. ornatus from Portugal
2012.10.16 © Murray, A.
Sminthurinus elegans f. ornatus from Portugal
2012.10.16 © Murray, A.
Sminthurinus elegans f. ornatus from the UK
2012.10.29 © Jones, A.
Sminthurinus elegans f. ornatus from the UK
Dorsal aspect
2012.11.06 © Murray, A.
Sminthurinus elegans f. ornatus from the UK
Facial aspect
2012.11.06 © Murray, A.
Sminthurinus elegans f. ornatus from the UK
Abdomen lateral aspect
2012.11.06 © Murray, A.
Sminthurinus elegans f. ornatus from the UK
6th abdominal segment
2012.11.06 © Murray, A.
Sminthurinus elegans f. ornatus from the UK
2013.09.18 © Nurcombe, E.
Sminthurinus elegans f. ornatus from the UK
2013.11.02 © Phillips, E.
Sminthurinus elegans f. ornatus from the UK
Note spermatophore of Isotomidae
2013.11.12 © Phillips, E.
Sminthurinus elegans f. ornatus from the UK
2014.08.21 © Phillips, E.
Sminthurinus elegans f. ornatus from the UK
2014.08.21 © Phillips, E.
Sminthurinus elegans f. ornatus from Holland
2014.11.23 © van Duinen, J.
Sminthurinus elegans f. ornatus from the UK
2016.01.21 © Phillips, E.
Sminthurinus elegans f. ornatus from the UK
2016.10.26 © Phillips, E.
Sminthurinus elegans f. ornatus from France
2018.01.05 © Tiky, S.
Sminthurinus elegans f. ornatus from France
2018.01.05 © Tiky, S.
Sminthurinus elegans f. ornatus from France
2018.01.05 © Tiky, S.
Sminthurinus elegans f. ornatus from France
2018.01.07 © Tiky, S.
Sminthurinus elegans f. ornatus juv. from France
2018.01.07 © Tiky, S.
Sminthurinus elegans f. ornatus from Holland
2018.06.17 © Kamsteeg, G.
Sminthurinus elegans f. ornatus from the UK
2018.10.23 © Smart, J.
Sminthurinus elegans f. ornatus from the UK
2018.10.23 © Smart, J.
Sminthurinus elegans f. ornatus from the UK
2019.08.15 © Smart, J.
Sminthurinus elegans f. ornatus from Belgium
2020.10.09 © Huskens, M.-L.
Sminthurinus elegans f. ornatus ♂ from the UK
2021.06.16 © Cann, A.J.
Sminthurinus elegans f. ornatus ♂ from the UK
2021.06.16 © Cann, A.J.
Sminthurinus elegans f. ornatus from the USA
2021.10.16 © Pavy, R.
Sminthurinus elegans f. ornatus from the USA
2022.10.20 © Loarie, D.


            Syn.: Sminthurinus aureus var. dorsalis Axelson, WM, 1905:794
            Syn.: Sminthurinus elegans var. atratus Mills, HB, 1934:96
             Syn.: Sminthurinus aureus var. atrata(sic) Christiansen, K et Bellinger, P, 1981:1116 ISS
             Syn.: Sminthurinus aureus var. atrata(sic) Christiansen, K et Bellinger, P, 1998:1268 ISS
            Syn.: Sminthurinus elegans f. cancellus Maynard, EA, 1951:238
Sminthurinus elegans f. cancellus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.522
Sminthurinus elegans f. cancellus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.523

             Syn.: Sminthurinus cancellatus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS
Sminthurinus elegans f. cancellus from the USA
2009.05.21 © Cowen, R.
Sminthurinus elegans f. cancellus from the USA
2012.04.18 © Kouri, J.
Sminthurinus elegans f. cancellus from the USA
2012.04.18 © Kouri, J.
Sminthurinus elegans f. cancellus from the USA
2017.01.26 © Dobbs, Br.
Sminthurinus elegans f. cancellus from the USA
2017.11.30 © Dobbs, Br.
Sminthurinus elegans nr f. cancellus ♂ from Belgium
2020.10.10 © Huskens, M.-L.
Sminthurinus elegans nr f. cancellus ♂ from Belgium
2020.10.10 © Huskens, M.-L.
Sminthurinus elegans nr f. cancellus ♂ from Belgium
Spermatophore deposited
2020.10.10 © Huskens, M.-L.
Sminthurinus elegans nr f. cancellus from Portugal
2021.01.19 © Viana, J.
Sminthurinus elegans nr f. cancellus from Holland
2023.04.17 © Huskens, M.-L.
Sminthurinus elegans nr f. cancellus from Holland
2023.04.17 © Huskens, M.-L.
Sminthurinus elegans nr f. cancellus from Holland
2023.04.17 © Huskens, M.-L.
Sminthurinus elegans nr f. cancellus from Holland
Dorso-caudal aspect
2023.04.17 © Huskens, M.-L.


            Syn.: Sminthurinus elegans var. ochropus Folsom, 1934
Sminthurinus elegans f. ochropus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.524
Sminthurinus elegans f. ochropus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.525
Sminthurinus elegans f. ochropus from the USA
2009.02.08 © Justis, S.
Sminthurinus elegans f. ochropus from the USA
2009.02.08 © Justis, S.
Sminthurinus elegans f. ochropus from the USA
2012.02.15 © Kouri, J.
Sminthurinus elegans f. ochropus from the USA
2012.02.15 © Kouri, J.
Sminthurinus elegans f. ochropus from the USA
2012.02.24 © Mallory, C.
Sminthurinus elegans f. ochropus from the USA
Egg cluster
2012.04.09 © Zungri, D.
Sminthurinus elegans f. ochropus from the USA
2016.12.23 © Dobbs, Br.
Sminthurinus elegans f. ochropus from the USA
2017.11.12 © Dobbs, Br.
Sminthurinus elegans f. ochropus from the USA
With pale abd.6
2017.11.17 © Dobbs, Br.
Sminthurinus elegans f. ochropus from the USA
With pale abd.6
2017.11.19 © Dobbs, Br.
Sminthurinus elegans f. ochropus from Canada
2018.11.12 © Barbin, T.
Sminthurinus elegans f. ochropus from Canada
2018.11.12 © Barbin, T.
Sminthurinus elegans nr f. ochropus ♀ from Belgium
2020.07.05 © Huskens, M.-L.
Sminthurinus elegans nr f. ochropus ♀ from Belgium
2020.07.05 © Huskens, M.-L.
Sminthurinus elegans nr f. ochropus ♀ from Belgium
Dorso-caudal aspect
2020.07.05 © Huskens, M.-L.
Sminthurinus elegans nr f. ochropucs ♀ from Belgium
With 'dented' eyebrows
2020.07.05 © Huskens, M.-L.
Sminthurinus elegans ?f. ochropus? from Canada
2023.02.05 © Rennert, M.
Sminthurinus elegans f. ochropus from Canada
2024.01.26 © Chan, J.
Sminthurinus elegans f. ochropus from Canada
2024.01.26 © Chan, J.


            Syn.: Sminthurinus similitortus Maynard, EA, 1951:232
Sminthurinus similitortus from the USA
After Maynard EA, 1951 Plate XXIV Fig.413
 

            Syn.: Sminthurinus aureus ssp. transversalis Salmon, JT, 1964:579



Sminthurinus elegans from the USA
After Maynard EA, 1951 Plate XXIV Fig.424
Sminthurinus elegans from Madeira
After Delamare Deboutteville C & Bassot J-M, 1957 Fig.1
Sminthurinus elegans from Australia
After Greenslade P & Ireson JE, 1986 Fig.2
Sminthurinus elegans
Rare form with lateral white spots
2004 © Baquero, E.
Sminthurinus elegans from the UK
Female sub-anal appendages
2004 © Hopkin, S.P.
Sminthurinus elegans from the USA
2006 © McClarin, J.
Sminthurinus elegans from Sweden
2006 © Fjellberg, A.
Sminthurinus elegans from the USA
2008.05.16 © Thombs, D.
Sminthurinus elegans from the USA
2009.02.14 © Justis, S.
Sminthurinus elegans from the USA
2009.02.14 © Justis, S.
Sminthurinus elegans from the USA
2009.02.14 © Justis, S.
Sminthurinus elegans from the USA
2009.02.14 © Justis, S.
Sminthurinus elegans from the USA
2010.01.05 © Justis, S.
Sminthurinus elegans from the UK
2011.12.31 © Jones, A.
Sminthurinus elegans from the USA
2012.01.29 © Kouri, J.
Sminthurinus elegans from the USA
2012.02.24 © Mallory, C.
Sminthurinus elegans juv. from the UK
2012.06.10 © Murray, A.
Sminthurinus elegans from the UK
2012.08.08 © Nurcombe, E.
Sminthurinus elegans from the UK
2012.08.16 © Nurcombe, E.
Sminthurinus elegans from Iran
After Falahati Hossein Abad A & al, 2013 Fig.4
Sminthurinus elegans from the USA
2013.04.04 © Kouri, J.D.
Sminthurinus elegans from the UK
Ant.3 with wart (*)
2013.08.25 © Henton-Jones, D.
Sminthurinus elegans from the UK
2013.09.18 © Nurcombe, E.
Sminthurinus elegans from the UK
2013.10.06 © Phillips, E.
Sminthurinus elegans juv. from the UK
Dark form
2013.11.25 © Phillips, E.
Sminthurinus elegans juv. from the UK
2014.08.28 © Phillips, E.
Sminthurinus elegans juv. from Holland
Ready to moult
2014.11.22 © van Duinen, J.
Sminthurinus elegans from Belgium
Very dark colour form
2017.01.31 © Huskens, M.-L.
Sminthurinus elegans ♀ from France
Bifurcate median dorsal circumanal seta (*) and sub-anal appendages (s.a.a.)
2017.08.20 © Picard, J.
Sminthurinus elegans juv. from Belgium
2017.10.17 © Huskens, M.-L.
Sminthurinus elegans from France
Dark form
2017.12.02 © Picard, J.
Sminthurinus elegans from Belgium
Eyepatch with 8 ocelli (C & D reduced)
2017.12.21 © Huskens, M.-L.
Sminthurinus elegans from Belgium
Very dark colour form
2018.10.01 © Huskens, M.-L.
Sminthurinus elegans from France
Near to colour form ornatus
2019.11.05 © Garcelon, P.
Sminthurinus elegans ♂ from France
Spermatophore deposition
2020.05.07 © Reinach, Br.
Sminthurinus elegans juv. from Belgium
With pathogenic fungal infection
2020.09.22 © Huskens, M.-L.
Sminthurinus elegans from Belgium
2021.05.20 © Huskens, M.-L.
Sminthurinus elegans from Holland
Abd.5 fused with great abdomen
2023.12.13 © Kamsteeg, G.
Sminthurinus elegans from Belgium
Very dark specimen
2024.03.17 © Huskens, M.-L.



Species exiguus (Nicolet, H, 1847:360)

Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Species gamae Gisin, H, 1963:88

Sminthurinus gamæ(sic) ♀ from Madeira
After Gisin H, 1963 Fig.9
Sminthurinus gamæ(sic) from Madeira
1 subapical anterior dental seta present (*)
Proximal posterior setae: 1 outer, 1 median, 1 inner
Modified after Gisin H, 1963 Fig.10

            Syn.: Sminthurinus igniceps da Gama, 1959 cf



             Syn.: Sminthurinus gamæ(sic) Gisin, H, 1963:89 ISS

Species gisini da Gama, 1965

            Syn.: Sminthurinus carpathicus Rusek, 1966
             Syn.: Sminthurinus carphathicus(sic) Baquero, E, Jordana, R et Ortuño, VM, 2021:7 ISS



Sminthurinus glaucus from New Zealand
After Salmon, J.T. 1943 Pl.1 Fig.8.
Species glaucus Salmon, JT, 1943:2


Species gloriosus (Womersley, H, 1937:157) Greenslade & Mayer, 1993 cf

            Syn.: Arrhopalites gloriosa(sic) Womersley, H, 1937:157 IOS
Arrhopalites gloriosa(sic) from Australia
After Womersley H, 1937 Fig.1m





            Syn.: Arrhopalites gloriosa(sic) Womersley, 1937 IOS
            Syn.: Arrhopalites gloriosa(sic) Womersley, H, 1939:225 IOS
            Syn.: Arrhopalites gloriosa(sic) Womersley, H, 1939:228 IOS
            Syn.: Arrhopalites gloriosa(sic) Womersley, H, 1939:272 IOS
            Syn.: Arrhopalites gloriosa(sic) Stach, J, 1956:72 IOS
            Syn.: Arrhopalites gloriosa(sic) Salmon, JT, 1964:573 IOS
            Syn.: NotArrhopalites gloriosa(sic) Betsch, J-M, 1980:130 IOS
            Syn.: Sminthurinus gloriosa(sic) Greenslade & Mayer, 1993 cf IOS
            Syn.: Sminthurinus gloriosa(sic) Greenslade, P in Houston, WWK, 1994:131 IOS

Species granulatus Salmon, JT, 1946:34

Species granulosus Enderlein, G, 1908:409

Species handschini Folsom, 1934

Species harryius Palacios-Vargas, 2017

Species henshawi (Folsom, JW, 1896:449) Maynard, EA, 1951:233

            Syn.: Smynthurus(sic) henshawii(sic) Folsom, JW, 1896:449 IOS
            Syn.: Sminthurinus aureus Folsom, 1898
            Syn.: Sminthurus aureus Guthrie, JE, 1903:43
            Syn.: Smynthurus(sic) aureus Mills, 1930
            Syn.: Sminthurus aureus Maynard, 1931
            Syn.: Sminthurinus aureus James, 1933
            Syn.: Sminthurinus aureus Mills, HB, 1934:95
            Syn.: Sminthurinus aureus Folsom, 1934
            Syn.: Sminthurinus aureus Brimley, 1938
            Syn.: Sminthurinus aureus Maynard, EA, 1951:233
Sminthurinus aureus from the USA
After Maynard EA, 1951 Plate XXIII Fig.410
Sminthurinus henshawi f. aureus from the USA
2009.03.10 © Cowen, R.
Sminthurinus henshawi f. aureus from the USA
2009.03.10 © Cowen, R.
Sminthurinus henshawi f. aureus from the USA
2009.03.10 © Cowen, R.
Sminthurinus henshawi f. aureus from the USA
2009.11.21 © Gledhill, L.
Sminthurinus henshawi f. aureus from the USA
Feeding instar
2009.11.22 © Gledhill, L.
Sminthurinus henshawi f. aureus from the USA
Feeding instar
2009.11.25 © Gledhill, L.
Sminthurinus henshawi f. aureus from the USA
2011.11.19 © Justis, S.
Sminthurinus henshawi f. aureus from the USA
2013.04.14 © Uhey, D.
Sminthurinus henshawi f. aureus from the USA
2015.04.17 © Dankowicz, E.
Sminthurinus henshawi f. aureus from the USA
Feeding instar with dark gut
2015.11.20 © Schrayer, G.
Sminthurinus henshawi f. aureus from the USA
2020.01.23 © Hendrix, S.
Sminthurinus henshawi f. aureus from Canada
2020.02.04 © Strickland, O.
Sminthurinus henshawi f. aureus from the USA
2020.11.15 © Kemper, Y.
Sminthurinus henshawi f. aureus from the USA
2022.12.31 © Hendrix, S.
Sminthurinus henshawi f. aureus from the USA
2022.12.31 © Hendrix, S.
Sminthurinus henshawi f. aureus from the USA
2024.04.10 © Hendrix, S.
Sminthurinus henshawi f. aureus from the USA
2024.04.10 © Hendrix, S.


            Syn.: Sminthurinus aureus var. novus Mills, HB, 1934:96
            Syn.: Sminthurinus megoculatus Maynard, EA, 1951:233
Sminthurinus megoculatus from the USA
After Maynard EA, 1951 Plate XXIX Fig.542
Sminthurinus megoculatus from the USA
After Maynard EA, 1951 Plate XXIX Fig.544
Sminthurinus henshawi f. megoculatus from the USA
2007.03.26 © Reed, D.E.
Sminthurinus henshawi f. megoculatus from the USA
2008.01.26 © Montgomery, G.
Sminthurinus henshawi f. megoculatus from the USA
2008.01.26 © Montgomery, G.
Sminthurinus henshawi f. megoculatus from the USA
2012.01.29 © Kouri, J.
Sminthurinus henshawi f. megoculatus from the USA
2012.01.29 © Kouri, J.
Sminthurinus henshawi f. megoculatus from the USA
2012.01.29 © Kouri, J.


            Syn.: Sminthurinus similitortus Maynard, EA, 1951:232
Sminthurinus similitortus from the USA
After Maynard EA, 1951 Plate XXIV Fig.414
Sminthurinus similitortus from the USA
After Maynard EA, 1951 Plate XXIX Fig.537
Sminthurinus similitortus from the USA
After Maynard EA, 1951 Plate XXIX Fig.538
Sminthurinus elegans from the USA
After Maynard EA, 1951 Plate XXVIII Fig.521
Sminthurinus henshawi f. similitortus from the USA
2009.02.05 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.02.05 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.02.07 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.02.07 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.02.07 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.02.07 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.04.08 © Justis, S.
Sminthurinus henshawi f. similitortus from the USA
2011.01.21 © Gerber, G.K.
Sminthurinus henshawi f. similitortus from the USA
2011.01.21 © Gerber, G.K.
Sminthurinus henshawi f. similitortus from the USA
2011.12.24 © Justis, S.
Sminthurinus henshawi f. similitortus from the USA
2011.12.24 © Justis, S.
Sminthurinus henshawi f. similitortus from the USA
2012.01.29 © Kouri, J.
Sminthurinus henshawi f. similitortus from the USA
2012.02.05 © Kouri, J.
Sminthurinus henshawi f. similitortus from the USA
2017.12.01 © Guevara, E.
Sminthurinus henshawi f. similitortus from Canada
2020.04.04 © Spencer, Br.
Sminthurinus henshawi f. similitortus from the USA
2021.02.05 © Shahan, T.
Sminthurinus henshawi f. similitortus from the USA
Ocelli C+D smaller
2021.03.12 © Shahan, T.
Sminthurinus henshawi f. similitortus from the USA
Ocelli C+D smaller
2021.03.12 © Shahan, T.
Sminthurinus henshawi f. similitortus from the USA
2021.03.15 © Shahan, T.
Sminthurinus henshawi f. similitortus from the USA
2022.01.01 © Charny, V.
Sminthurinus henshawi f. similitortus from the USA
Pattern similar to S. reticulatus
2022.12.29 © Shahan, T.
Sminthurinus henshawi f. similitortus from the USA
Abd.5 not fused with great abdomen
2022.12.29 © Shahan, T.
Sminthurinus henshawi f. similitortus from the USA
Caudal aspect
2022.12.29 © Shahan, T.
Sminthurinus henshawi f. similitortus from the USA
Ocellus B distinctly large
2022.12.29 © Shahan, T.
Sminthurinus henshawi f. similitortus from the USA
Eating spermatophore
2023.01.19 © Gruver, Be.
Sminthurinus henshawi f. similitortus from the USA
Depositing spermatophore
2023.01.19 © Gruver, Be.




             Syn.: Sminthurinus henshawii(sic) Stach, 1956 ISS
Sminthurinus (Sminthurinus) henshawi from the USA
After Snider RJ in Christiansen KA & Bellinger PF, 1998 Fig.914A
Sminthurinus henshawi from the USA
2006 © McClarin, J.
Sminthurinus henshawi from the USA
2009.11.25 © Gledhill, L.
Sminthurinus henshawi from the USA
2012.03.03 © Montgomery, G.
Sminthurinus henshawi from Canada
Feeding instar; with phoretic nematode
2013.01.15 © Kyron, T.F.
Sminthurinus henshawi from Canada
2013.01.15 © Kyron, T.F.
Sminthurinus henshawi from Canada
2013.01.15 © Kyron, T.F.
Sminthurinus henshawi from the USA
2013.04.14 © Uhey, D.
Sminthurinus henshawi juv. from Canada
2014.01.23 © Hochhalter, R.
Sminthurinus henshawi from the USA
2014.04.20 © Dankowicz, E.
Sminthurinus henshawi from the USA
2018.12.27 © Duncan, W.L.
Sminthurinus henshawi from the USA
2019.01.23 © Duncan, W.L.
Sminthurinus henshawi from the USA
Grooming its back with drop of recycled urine secreted from labium or collophore
2021.01.08 © Duncan, W.L.
Sminthurinus henshawi from the USA
With fungal infection
2023.01.29 © Gruver, Be.
Sminthurinus henshawi from the USA
2024.04.10 © Hendrix, S.
Sminthurinus henshawi from the USA
2024.04.10 © Hendrix, S.





Species hygrophilus Bretfeld, 2000

             Syn.: Sminthurinus hydrophilus(sic) Babenko, AB, 2007:377 ISS

Species igniceps (Reuter, OM in Lindberg, SO, 1881:203) Börner, C, 1901:102

            Syn.: Sminthurus rex Uzel, 1891


Sminthurus igniceps from Finland
After Reuter OM, 1892 Tab.I Fig.2
Sminthurus igniceps from Finland
Head frontal aspect
After Reuter OM, 1892 Tab.I Fig.2a
Sminthurus igniceps from Sweden
After Schött H, 1893 Taf.II Fig.17
Sminthurinus igniceps ♀ from Poland
After Stach J, 1956 Plate XV Fig.1
Sminthurinus igniceps ♂ from Poland
After Stach J, 1956 Plate XV Fig.2
             Syn.: Sminthurinus ignipes(sic) Grinbergs, 1960 ISS
             Syn.: Sminthurinus ignipes(sic) Busmachiu, 2010 ISS
Sminthurinus ?igniceps? from France
2010 © Porco, D.
Sminthurinus igniceps from the UK
2015.07.27 © Boardman, P.
Sminthurinus igniceps from Switzerland
2017.03.19 © Lorenzo, R.
Sminthurinus igniceps from Switzerland
2017.03.19 © Lorenzo, R.
Sminthurinus igniceps from Belgium
Abd.5 not fused with great abdomen
2020.04.19 © Huskens, M.-L.
Sminthurinus igniceps from Belgium
2020.04.19 © Huskens, M.-L.
Sminthurinus ?igniceps? from the UK
2020.11.07 © Lewis, M.P.
Sminthurinus ?igniceps? from the UK
2020.11.07 © Lewis, M.P.
Sminthurinus ?igniceps? from Portugal
2021.01.03 © Viana, J.
Sminthurinus ?igniceps? from Portugal
2021.01.03 © Viana, J.

             Syn.: Sminthurinus ignipes(sic) Sinchuk, AV, Logachev, MA et Sinchuk, NV, 2023:285 ISS


Species imperialis Palacios-Vargas, JG, Hornung-Leoni, CT et Garrido, I, 2012:60

Sminthurinus imperialis from Mexico
After Palacios-Vargas JG & al, 2012 Fig.1A
             Syn.: Sminthurinus Imperialis(sic) Palacios-Vargas, JG, Hornung-Leoni, CT et Garrido, I, 2012:60 ISS


Species inexcussus Najt & Rapoport, 1965

Species insularis Najt, 1972

Species jonesi Wise, KAJ, 1970:203

Species kaha Christiansen, K et Bellinger, P, 1992:349

Species kerguelensis Salmon, JT, 1964:314

Species latimaculosus Maynard, EA, 1951:231

Sminthurinus latimaculosus from the USA
After Maynard EA, 1951 Plate XXIII Fig.389
Sminthurinus latimaculosus from the USA
After Maynard EA, 1951 Plate XXIII Fig.390




             Syn.: Sminthurinus latimaculatus(sic) Zeppelini, 1996 ISS
             Syn.: Sminthurinus latimaculosos(sic) Palacios-Vargas, JG et Castaño-Meneses, G in Basset, Y, Novotny, V, Miller, SE et Kitching, RL, 2003:169 ISS
Sminthurinus latimaculosus from the USA
2012.06.15 © Anderson, J.



Species lawrencei Gisin, H, 1963:87, B

Sminthurinus lawrencei from the UK
Subapical anterior dental setae absent (*)
Proximal posterior setae: 2 outer, 1 median, 1 inner
Modified after Gisin H, 1963 Fig.8
Sminthurinus lawrencei from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus lawrencei from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus lawrencei from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus lawrencei from the UK
2007.12.22 © Barton, T.
Sminthurinus lawrencei from the UK
2007.12.22 © Barton, T.
Sminthurinus lawrencei ♀ from the UK
2008.04.08 © Valentine., B
Sminthurinus lawrencei ♂ from the UK
2008.04.08 © Valentine., B
Sminthurinus lawrencei from the UK
2008.11.28 © Valentine., B
Sminthurinus lawrencei from the UK
2008.12.25 © Corley, P.
Sminthurinus lawrencei from the UK
2009.01.04 © Corley, P.
Sminthurinus lawrencei from the UK
2009.03.10 © Valentine., B
Sminthurinus lawrencei from the UK
2009.03.10 © Valentine., B
Sminthurinus lawrencei from the UK
2009.04.12 © Barton, T.
Sminthurinus lawrencei from the UK
2009.12.06 © Barton, T.
Sminthurinus lawrencei from the UK
2009.12.28 © Jones, A.
Sminthurinus lawrencei from the UK
2011.12.11 © Nurcombe, E.
Sminthurinus lawrencei from the UK
2011.12.11 © Nurcombe, E.
Sminthurinus lawrencei from the UK
2012.12.07 © Nurcombe, E.
Sminthurinus lawrencei from the UK
Wart on 3rd antennal segment
2012.12.13 © Valentine., B
Sminthurinus lawrencei from the UK
Speckled specimen: ready to moult
2013.01.04 © Murray, A.
Sminthurinus lawrencei juv. from the UK
2013.01.12 © Nurcombe, E.
Sminthurinus lawrencei from Canada
Ant.3 with papilla (*)
2013.01.23 © Kyron, T.F.
Sminthurinus lawrencei from Canada
2013.01.23 © Kyron, T.F.
Sminthurinus lawrencei from Canada
2013.01.23 © Kyron, T.F.
Sminthurinus lawrencei from Canada
2013.01.23 © Kyron, T.F.
Sminthurinus lawrencei from Canada
Dental anterior chaetotaxy
Subapical setae absent (*)
2013.01.23 © Kyron, T.F.
Sminthurinus lawrencei from Canada
Dental posterior chaetotaxy
Proximal setae : 2 outer, 1 median, 1 inner
2013.01.23 © Kyron, T.F.
Sminthurinus lawrencei from Canada
Hind foot with 4 tenent setae
2013.01.24 © Kyron, T.F.
Sminthurinus lawrencei ♀ from Canada
Female palmate sub-anal appendage (s.a.a) on papilla (*)
2013.01.24 © Kyron, T.F.
Sminthurinus lawrencei ♀ from Canada
Circumanal setae: dorsal median bifid seta (*), setae with expanded base (**), s.a.a. (***)
2013.01.24 © Kyron, T.F.
Sminthurinus lawrencei juv. from the UK
Speckled : preparing to moult
2013.02.06 © Murray, A.
Sminthurinus lawrencei from the UK
Cuticular tubercles
2013.02.06 © Murray, A.
Sminthurinus lawrencei from the UK
Footcomplex with unguiculus (u), unguis (U) and clavate tenent macrosetae (*)
2013.12.09 © Phillips, E.
Sminthurinus lawrencei from the UK
2013.12.10 © Phillips, E.
Sminthurinus lawrencei from the UK
Trapped in sticky spider web
2015.02.05 © Garlick, T.
Sminthurinus lawrencei from the UK
2015.02.05 © Garlick, T.
Sminthurinus lawrencei from the UK
Speckled specimen: ready to moult
2015.02.05 © Garlick, T.
Sminthurinus lawrencei from the UK
Short white "eyebrows"
2015.02.15 © Phillips, E.
Sminthurinus lawrencei juv. from the UK
Speckled specimen: ready to moult
2015.11.10 © Jeffery, J.
Sminthurinus lawrencei from the UK
Genital abdominal segment not fused with great abdomen
2016.01.06 © Phillips, E.
Sminthurinus lawrencei from Belgium
2016.02.02 © Huskens, M.-L.
Sminthurinus lawrencei juv. from Belgium
Preparing to moult
2016.03.15 © Huskens, M.-L.
Sminthurinus lawrencei ♀ from the UK
Abd.5 not fused with large abdomen
Sub-anal appendage (*)
2017.01.14 © David W.
Sminthurinus lawrencei from France
Ant.3 with wart (*)
2017.10.15 © Picard, J.
Sminthurinus lawrencei from Holland
2017.11.21 © Kamsteeg, G.
Sminthurinus lawrencei from Holland
7 large ocelli (D not visible)
Short white "eyebrow"
2017.11.21 © Kamsteeg, G.
Sminthurinus lawrencei ♂ from Belgium
2017.11.25 © Huskens, M.-L.
Sminthurinus lawrencei from Belgium
3 large inner ocelli BCH in straight line
White antennal base
2017.11.25 © Huskens, M.-L.
Sminthurinus lawrencei from Holland
Cuticular tubercles
2017.11.26 © Kamsteeg, G.
Sminthurinus lawrencei from Belgium
Collophore vesicles eversed just before escape jump
2017.11.28 © Huskens, M.-L.
Sminthurinus lawrencei juv. from Holland
2018.01.15 © Wieringa, P.
Sminthurinus lawrencei from Holland
Distinctly speckled = ready to moult
2018.01.26 © Kamsteeg, G.
Sminthurinus lawrencei from France
Ant.3 base with wart (*)
2018.02.01 © Garcelon, P.
Sminthurinus lawrencei from the UK
Abd.5 not included in great abdomen (*)
2018.12.13 © Uncle, F.
Sminthurinus lawrencei from the UK
Dens with long mucro; m/d ratio 0.6
2018.12.13 © Uncle, F.
Sminthurinus lawrencei from the UK
Footcomplex with inner unguiculus (u), outer unguis (U) and outer clavate tenent setae (t)
2018.12.13 © Uncle, F.
Sminthurinus lawrencei from the UK
Ant.3 with inner basal wart (*)
2018.12.13 © Uncle, F.
Sminthurinus lawrencei from France
2019.02.03 © Nogues, Be.
Sminthurinus lawrencei from the UK
Speckled specimen: ready to moult
2019.03.08 © Marquis, A.
Sminthurinus lawrencei from the UK
With phoretic Nematoda
2020.01.25 © Valentine, Br.
Sminthurinus lawrencei juv. from Belgium
2020.12.17 © Huskens, M.-L.
Sminthurinus lawrencei from the UK
2021.01.20 © McCulloch, J.
Sminthurinus lawrencei from the UK
Abd.5 separated from great abdomen
2021.01.20 © McCulloch, J.
Sminthurinus lawrencei from the UK
Subapical anterior dental setae absent
2021.01.20 © McCulloch, J.
Sminthurinus lawrencei from the UK
Eyebrow white and short
2021.01.20 © McCulloch, J.
Sminthurinus lawrencei from the UK
Ant.3 with elongate undivided basal wart (*)
2021.01.20 © McCulloch, J.
Sminthurinus lawrencei from the UK
Eyepatch with depigmented ocelli?
2021.01.20 © McCulloch, J.
Sminthurinus lawrencei from the UK
Subapical anterior dental setae absent
2021.01.21 © McCulloch, J.
Sminthurinus lawrencei from the UK
Mucro with both serrate edges
2021.01.21 © McCulloch, J.
Sminthurinus lawrencei from the UK
4 proximal posterior dental setae
2 outer (o), 1 long median (m), 1 inner (i)
2021.01.22 © McCulloch, J.
Sminthurinus lawrencei ♀ from the UK
Sub-anal appendage with about 6 short branches
2021.01.22 © McCulloch, J.
Sminthurinus lawrencei from the UK
Ant.3 with elongate undivided basal wart
2021.01.25 © McCulloch, J.
Sminthurinus lawrencei from Belgium
Preparing to moult
Tibiotarsus apically white
2021.01.26 © Huskens, M.-L.
Sminthurinus lawrencei from Belgium
Preparing to moult
Appearance spotted
2021.01.26 © Huskens, M.-L.
Sminthurinus lawrencei ♀ from Holland
2021.02.27 © van Leeuwen, J.
Sminthurinus lawrencei ♀ from Holland
Short eyebrow
2021.02.27 © van Leeuwen, J.
Sminthurinus lawrencei ♀ from Holland
Unguiculus with filament as long as unguis
2021.02.28 © van Bezouw, R.
Sminthurinus lawrencei ♀ from Holland
Bipartite eyebrow
Anterior part colourless
2021.02.28 © van Bezouw, R.
Sminthurinus lawrencei ♀ from Holland
Ant.3 wart undivided
2021.02.28 © van Bezouw, R.
Sminthurinus lawrencei ♀ from Holland
Dens without anterior subapical setae (*)
2021.02.28 © van Bezouw, R.
Sminthurinus lawrencei ♀ from Holland
Dens with 2 lateral subapical outer setae
2021.02.28 © van Bezouw, R.
Sminthurinus lawrencei ♀ from Holland
Dens with 1 long posterior proximal inner seta
2021.02.28 © van Bezouw, R.
Sminthurinus lawrencei ♀ from Holland
Dens with 1 long posterior proximal median seta
And 3 subapical median setae
2021.02.28 © van Bezouw, R.
Sminthurinus lawrencei ♀ from Holland
Dens with 2 posterior proximal outer setae
2021.02.28 © van Bezouw, R.
Sminthurinus lawrencei ♀ from Holland
Sub-anal appendage in lateral aspect
2021.02.28 © van Bezouw, R.
Sminthurinus lawrencei ♀ from Holland
Mucro with both edges serrated
2021.02.28 © van Bezouw, R.
Sminthurinus lawrencei ♀ from Holland
Apex of sub-anal appendage with 6 branches (*)
2021.03.01 © van Bezouw, R.
Sminthurinus lawrencei ♀ from Belgium
Distinctly short eyebrows
Clypeo-labral area without spots
2021.03.31 © Huskens, M.-L.
Sminthurinus lawrencei ♀ from Belgium
Facial chaetotaxy
Negative image
2021.03.31 © Huskens, M.-L.
Sminthurinus ?lawrencei? from Holland
Preparing to moult
2021.04.05 © Kamsteeg, G.
Sminthurinus lawrencei ♀ from Belgium
Furca
2021.04.11 © Huskens, M.-L.
Sminthurinus lawrencei from Belgium
2 ocular setae
2021.04.18 © Huskens, M.-L.
Sminthurinus lawrencei from Belgium
Distance between eyebrows > 2x width eyebrow
With upside down spermatophore
2021.05.07 © Huskens, M.-L.
Sminthurinus lawrencei from Belgium
4 equidistant white spots form square
2021.05.07 © Huskens, M.-L.
Sminthurinus lawrencei from Belgium
Eye with 2 ocullar setae :
oc2 inbetween G and H; oc3 inbetween F and H
2021.05.12 © Huskens, M.-L.
Sminthurinus lawrencei from Holland
Retinaculum with anterior lobe (al), posterior lobe (pl) and ramus (r) bearing 3 teeth
2021.08.09 © van Bezouw, R. Leg.: Berg, M.
Sminthurinus lawrencei from Belgium
Inter-eyebrows distance relatively large
2021.08.12 © van Bezouw, R.
Leg.: Huskens, M.-L.
Sminthurinus lawrencei from Belgium
Small eyebrow not reaching frontocellar suture
2021.08.12 © van Bezouw, R.
Leg.: Huskens, M.-L.
Sminthurinus lawrencei from the UK
After McCulloch J, 2021 Fig.1
Sminthurinus lawrencei from Belgium
2021.10.13 © Huskens, M.-L.
Sminthurinus lawrencei juv. from Belgium
Spotted appearance
Preparing to moult
2021.10.13 © Huskens, M.-L.
Sminthurinus lawrencei from Belgium
2021.10.23 © Huskens, M.-L.
Sminthurinus lawrencei from Belgium
Distinctly small eyebrow
not reaching frontocular suture
2021.11.09 © Huskens, M.-L.
Sminthurinus lawrencei from the UK
2021.11.28 © Valentine, Br.
Sminthurinus lawrencei from the UK
2021.12.09 © Valentine, Br.
Sminthurinus lawrencei from Holland
Facial aspect
Distinctly small eyebrows
2022.02.08 © Kamsteeg, G.
Sminthurinus lawrencei from Belgium
Eyebrow short
with two inner indentations (*)
2022.02.22 © Huskens, M.-L.
Sminthurinus lawrencei from Belgium
Mid-anal seta bifurcate
2022.02.22 © Huskens, M.-L.
Sminthurinus lawrencei from Belgium
Head tilted with mouth opening forwards
Occiput not visible
2022.10.14 © Huskens, M.-L.
Sminthurinus lawrencei from Holland
Eyebrows distinctly small
Not reaching ocular suture
2023.04.17 © Kamsteeg, G.
Sminthurinus lawrencei from Canada
2023.12.02 © Chans, J.
Sminthurinus lawrencei from Belgium
2024.03.06 © Huskens, M.-L.
Sminthurinus lawrencei from Belgium
Head tilted with mouth opening backwards
Occiput appears as narrow rim around eyes
2024.03.06 © Huskens, M.-L.

Species lichenatus Salmon, JT, 1943:3

Species lineatus Bretfeld in Bretfeld, Poliakov & Broza, 2000

Species maculosus Snider, 1978

Sminthurinus (Sminthurinus) maculosus from the USA
After Christiansen K & Bellinger P, 1998 Fig.943A

             Syn.: Sminthurinus maculosos(sic) Palacios-Vargas, JG, Hornung-Leoni, CT et Garrido, I, 2012:61 ISS
Sminthurinus maculosus from the USA
2017.01.28 © Abela, A.
Sminthurinus maculosus from the USA
2017.04.05 © Johnson, S.T.
Sminthurinus maculosus from the USA
Pale form
2017.04.05 © Johnson, S.T.



Species mime Börner in Voeltzkow, 1907

            Syn.: Sminthurus niger Parona, 1895
            Syn.: Sminthurinus niger Börner in Voeltzkow, 1907
            Syn.: Sminthurinus terrestris Womersley, 1931


Sminthurinus mime? from South Africa
2015.09.24 © Adkock, R.
Sminthurinus mime? from South Africa
2015.09.24 © Adkock, R.

Species minutus (MacGillivray, AD, 1894:109) Maynard, EA, 1951:234

Sminthurinus minutus from the USA
After Maynard EA, 1951 Plate XXIII Fig.402
Sminthurinus minutus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.535
Sminthurinus minutus from the USA
2008.02.09 © Montgomery, G.
Sminthurinus minutus from the USA
2009.12.29 © Zungri, D.
Sminthurinus minutus from the USA
2010.01.01 © Zungri, D.
Sminthurinus minutus juv. from the USA
2010.01.24 © Zungri, D.
Sminthurinus minutus from the USA
2010.02.07 © Zungri, D.
Sminthurinus minutus from the USA
2010.03.23 © Montgomery, G.
Sminthurinus minutus from the USA
2010.03.23 © Montgomery, G.
Sminthurinus nr minutus from the UK
2014.11.24 © Nurcombe, E.
Sminthurinus nr minutus from the UK
2014.11.24 © Nurcombe, E.
Sminthurinus nr minutus from the UK
2014.11.27 © Nurcombe, E.
Sminthurinus nr minutus from the UK
2014.11.27 © Nurcombe, E.
Sminthurinus nr minutus from the UK
2014.11.27 © Nurcombe, E.
Sminthurinus nr minutus from the UK
2015.01.04 © Nurcombe, E.
Sminthurinus nr minutus juv. from the UK
2015.01.04 © Nurcombe, E.
Sminthurinus nr minutus from the UK
2015.02.08 © Phillips, E.
Sminthurinus nr minutus from France
2017.05.07 © Picard, J.
Sminthurinus nr minutus from France
2017.05.29 © Picard, J.
Sminthurinus nr minutus from France
2017.05.29 © Picard, J.
Sminthurinus nr minutus from France
2017.11.17 © Tiky, S.
Sminthurinus nr minutus from France
2017.11.17 © Tiky, S.
Sminthurinus nr minutus from France
2017.11.17 © Tiky, S.
Sminthurinus minutus from the USA
2017.12.03 © Dobbs, Br.
Sminthurinus minutus from the USA
2017.12.03 © Dobbs, Br.
Sminthurinus minutus from the USA
2017.12.03 © Dobbs, Br.
Sminthurinus nr minutus from Holland
2018.01.14 © de Haas, M.
Sminthurinus nr minutus from Holland
2019.01.02 © de Haas, M.
Sminthurinus nr minutus from Holland
2019.01.02 © de Haas, M.
Sminthurinus nr minutus from Holland
2019.11.03 © de Haas, M.
Sminthurinus minutus from the USA
2020.03.22 © Engel, V.
Sminthurinus ?minutus? from the USA
2021.09.09 © Caterino, M.
Sminthurinus minutus from the USA
2022.02.18 © Charny, V.
Sminthurinus minutus from the USA
2022.02.18 © Charny, V.
Sminthurinus minutus from the USA
Abd.6 pattern
2022.02.18 © Charny, V.
Sminthurinus minutus from the USA
2023.11.30 © Schneider, J.
Sminthurinus minutus from the USA
2023.11.30 © Schneider, J.
Sminthurinus minutus from the USA
2023.11.30 © Schneider, J.

Species modestus Yosii, R, 1970:20

Sminthurinus modestus from Japan
After Yosii R, 1970 Fig.13A
Sminthurinus modestus from Japan
After Yosii R, 1970 Fig.13B
Sminthurinus modestus from Japan
2014.01.13 © Komatsu, T.
Sminthurinus modestus juv. from Japan
2019.01.20 © Hara, Y.
Sminthurinus modestus juv. from Japan
2019.01.20 © Hara, Y.
Sminthurinus nr modestus from Taiwan
2022.02.13 © Cheng, H.-J.
Sminthurinus nr modestus from Taiwan
2022.02.13 © Cheng, H.-J.
Sminthurinus nr modestus from Taiwan
2022.02.13 © Cheng, H.-J.
Sminthurinus nr modestus ♂ from Taiwan
2022.05.06 © Cheng, H.-J.
Sminthurinus nr modestus ♂ from Taiwan
2022.05.06 © Cheng, H.-J.
Sminthurinus nr modestus ♂ from Taiwan
2022.05.06 © Cheng, H.-J.

Species molinai Arlé, 1940

Species mountfordi (Womersley, H, 1939:242) Greenslade P in Houston, 1994:132

Species muscophilus Salmon, JT, 1946:32

Species niger (Lubbock, 1862) Börner, C, 1901:101, B

            Syn.: Sminthurus bimaculatus Tullberg, 1871
            Syn.: ... niger f. ochracea Axelson, 1902
            Syn.: Sminthurinus albifrons Bagnall, 1921 =?
            Syn.: Sminthurinus albifrons Jackson, 1928 =?
            Syn.: Sminthurinus albifrons Hopkin, SP, 2007:178 =?
             Syn.: Sminthurinus aureus var. alba(sic) Stach, J, 1956:122 ISS
             Syn.: Sminthurinus aureus var. alba(sic) Stach, J, 1956:237 ISS
            Syn.: Smynthurus(sic) concolor Gisin, H, 1960:280
            Syn.: Sminthurinus concolor Scott, 1963


Smynthurus(sic) niger from the UK
After Lubbock J, 1870 Pl.XXI Fig.11
Smynthurus(sic) niger from the UK
Distinctly big eyebrows
After Lubbock J, 1870 Pl.XXI Fig.11
Smynthurus(sic) niger from the UK
Saltatory appendage
After Lubbock J, 1870 Pl.XXI Fig.12
Smynthurus(sic) niger from the UK
After Lubbock J, 1873 Pl.6

             Syn.: Sminthurinus Niger(sic) Denis, J-R, 1933:280 ISS
Sminthurinus niger from the UK
1 subapical anterior dental seta present (*)
Proximal posterior setae: 1 outer, 3 inner
Modified after Gisin H, 1963 Fig.1
Sminthurinus niger from the UK
Ocellus D smaller
2 setae inbetween ocelli H and G
Modified after Gisin H, 1963 Fig.2
Sminthurinus niger from the UK
2007.03.04 © Cornwall, N.J.
 
Sminthurinus niger from the UK
2007.03.18 © Cornwall, N.J.
Sminthurinus niger from France
2011.08.29 © Lebeaux, P.
Sminthurinus niger from France
2011.08.29 © Lebeaux, P.
Sminthurinus ?niger? from France
Distinctly speckled = ready to moult
2018.02.01 © Garcelon, P.
Sminthurinus niger from France
2018.03.02 © Garcelon, P.
Sminthurinus niger from France
Eyebrows triangular
2018.03.02 © Garcelon, P.
Sminthurinus niger from Portugal
Distinctly speckled = ready to moult
2018.11.10 © Almeida, J.
Sminthurinus niger from Spain
2020.03.08 © Zeeders, M.
Sminthurinus niger from France
2020.11.05 © Garcelon, P.
Sminthurinus niger from Holland
Abd.5 with bothriotrichum D (*)
2020.12.05 © Kamsteeg, G.
Sminthurinus niger from Holland
Eyebrows triangular
2020.12.05 © Kamsteeg, G.
Sminthurinus niger from Holland
Eyebrow suddenly more narrow
2021.03.08 © Kamsteeg, G.
Sminthurinus niger juv. from Holland
Eyebrow with elongated narrow branch
2021.03.08 © Kamsteeg, G.
Sminthurinus niger juv. from Holland
Eyebrow triangular
Pre-moult instar
2021.03.08 © Kamsteeg, G.
Sminthurinus niger from Holland
Eyebrows broad and long
2021.03.14 © Kamsteeg, G.
Sminthurinus niger from Holland
2 ocular setae
2021.03.17 © Kamsteeg, G.
Sminthurinus niger from Holland
2021.05.21 © Kamsteeg, G.
Sminthurinus niger from Holland
Eyebrow broad and triangular
circumventing eyepatch
2021.05.21 © Kamsteeg, G.
Sminthurinus niger from Holland
Mucro with 2 edges serrated
2021.05.23 © van Bezouw, R.
Sminthurinus niger from Holland
1 dental anterior subapical seta (*)
2021.05.23 © van Bezouw, R.
Sminthurinus niger from Holland
Dental posterior proximal setae :
1 outer (o), 1 median (m), 2 inner (i)
2021.05.23 © van Bezouw, R.
Sminthurinus niger from Holland
2022.02.10 © Kamsteeg, G.
Sminthurinus niger from Holland
Eye with 2 ocular setae
2022.02.10 © Kamsteeg, G.
Sminthurinus niger from Holland
Head dorsal profile
Similar to Stenognathellus
2022.02.10 © Kamsteeg, G.
Sminthurinus niger from Holland
Spotted
During moult
2022.02.10 © Kamsteeg, G.
Sminthurinus niger juv. from Holland
2022.10.28 © Kamsteeg, G.
Sminthurinus niger from Holland
Bothriotrichum D setaceous
2022.11.01 © Kamsteeg, G.



Species nigrafuscus Salmon, JT, 1941:409

Species nigrescens Schött, H, 1917:54

Species nunezi Rapoport in Delamare Deboutteville & Massoud, 1963:210

            Syn.: Sminthurinus Nunezi(sic) Rapoport in Delamare Deboutteville & Massoud, 1963:210 IOS
             Syn.: Sminthurinus nuñezi(sic) de Izarra, DC, 1965:275 ISS
Sminthurinus nunezi from Peru
2009.01-03.dd © Contreras Valdez, A.





Species obscurus Salmon, JT, 1944:173

Species oculatus Schött, H, 1917:53

Sminthurinus oculatus typus I from Australia
After Schött H, 1917 Taf.4 Fig.1
Sminthurinus oculatus typus I from Australia
After Schött H, 1917 Taf.4 Fig.2
Sminthurinus oculatus typus II from Australia
After Schött H, 1917 Taf.4 Fig.3
Sminthurinus oculatus typus III from Australia
After Schött H, 1917 Taf.4 Fig.4

Species oiskiyensis Bretfeld, G, 2010:305

Sminthurinus oiskyensis from Russia
After Bretfeld G, 2010 Fig.1

Species operosus Najt & Rapoport, 1965

Species orientalis Stach, J, 1964:21

Sminthurinus orientalis from China
After Stach J, 1964 Plate XI Fig.1
Sminthurinus orientalis from China
After Stach J, 1964 Plate XI Fig.2

Species ovatulus (Koch, CL & Berendt, GC in Berendt, GC, 1854:121, -) -

Smynthurus(sic) ovatulus from Baltic amber
After Koch CL & Berendt GC, 1845 Tab.XV Fig.132

Species packardi Denis, J-R, 1933:274

            Syn.: Sminthurinus Packardi(sic) Denis, J-R, 1933:274 IOS
Sminthurinus Packardi(sic) from Costa Rica
After Denis J-R, 1933 Fig.62

            Syn.: Sminthurinus Packardi(sic) Denis, J-R, 1933:275 IOS
            Syn.: Sminthurinus Packardi(sic) Denis, J-R, 1933:276 IOS
            Syn.: Sminthurinus Packardi(sic) Denis, J-R, 1933:315 IOS
            Syn.: Sminthurinus Packardi(sic) Denis, J-R, 1933:320 IOS


Species pallescens Yosii, R, 1970:22

Species pallidus Womersley, 1931

Species patagonicus Najt, 1971

Species pekinensis Stach, J, 1964:22

Sminthurinus pekinensis from China
After Stach J, 1964 Plate XII Fig.1
Sminthurinus pekinensis from China
After Stach J, 1964 Plate XII Fig.2

Species planasiensis Dallai, 1969

Species proceraseta Salmon, JT, 1946:32

Species pullus Bretfeld in Bretfeld, Poliakov & Broza, 2000

Species quadratus Bretfeld in Bretfeld, Poliakov & Broza, 2000

Sminthurinus quadratus from Germany
2014.05.05 © Beck, K.
Sminthurinus quadratus from Holland
2017.10.17 © Wieringa, P.
Sminthurinus quadratus from France
2017.11.18 © Tiky, S.
Sminthurinus quadratus from France
2017.11.18 © Tiky, S.
Sminthurinus quadratus from Belgium
2018.12.11 © Huskens, M.-L.
Sminthurinus quadratus from Denmark
2018.12.12 © Schlütter, P.
Sminthurinus quadratus from France
2019.01.01 © Picard, J.
Sminthurinus quadratus from Israel
2019.05.07 © Schlütter, P.
Det.: Schlütter, P.
Sminthurinus quadratus from Israel
2019.05.07 © Schlütter, P.
Det.: Schlütter, P.
Sminthurinus quadratus from Israel
Dark form
2019.05.07 © Schlütter, P.
Det.: Schlütter, P.
Sminthurinus quadratus from Sweden
2019.11.26 © Neergaard, R.
Sminthurinus quadratus from Sweden
2019.12.02 © Neergaard, R.
Sminthurinus quadratus from Belgium
2020.07.01 © Huskens, M.-L.
Sminthurinus quadratus from the UK
Dark form
2020.09.06 © Phillips, E.
Sminthurinus quadratus from the UK
Dark form
2020.09.06 © Phillips, E.
Sminthurinus quadratus from the UK
Dark form
2020.09.06 © Phillips, E.
Sminthurinus quadratus juv. from Belgium
2022.12.26 © Huskens, M.-L.
Sminthurinus quadratus juv. from Belgium
2022.12.26 © Huskens, M.-L.
Sminthurinus quadratus juv. from Belgium
2022.12.26 © Huskens, M.-L.
Sminthurinus quadratus from Belgium
Dark form
2022.12.26 © Huskens, M.-L.
Sminthurinus quadratus from Belgium
Dark form
2022.12.26 © Huskens, M.-L.
Sminthurinus quadratus from Holland
2023.11.06 © Kamsteeg, G.
Sminthurinus quadratus juv. from Holland
2023.11.06 © Kamsteeg, G.

Species quadrimaculatus (Ryder, 1878) Denis, 1933

            Syn.: Smynthurus(sic) quadrimaculata(sic) Ryder, 1878 IOS

            Syn.: Smynthurus(sic) 6-maculata(sic) Harvey, 1892 IOS
            Syn.: Smynthurus(sic) sexmaculatus MacGillivray, AD, 1894:109
             Syn.: Sminthurinus Sexmaculatus(sic) Denis, J-R, 1933:293 ISS
            Syn.: Smynthurus(sic) minutus MacGillivray, 1894
            Syn.: Sminthurus (Pilosi) niger Schött, H, 1891:12
Sminthurus niger from the USA
After Schött H, 1891 Taf.II Fig.1

            Syn.: Smynthurus(sic) niger Christiansen, K et Bellinger, P, 1981:1117
            Syn.: Smynthurus(sic) niger Christiansen, K et Bellinger, P, 1998:1269
Sminthurinus quadrimaculatus f. "niger" from the USA
2008.01.21 © Moorehead, C.
Sminthurinus quadrimaculatus f. "niger" from the USA
2009.04.08 © Zungri, D.
Sminthurinus quadrimaculatus f. "niger" from the USA
2010.02.07 © Zungri, D.
Sminthurinus quadrimaculatus f. "niger" from the USA
2017.12.01 © Dobbs, Br.
Sminthurinus quadrimaculatus f. "niger" from the USA
Distinctly speckled = ready to moult
2017.12.01 © Dobbs, Br.
Sminthurinus quadrimaculatus f. "niger" from Canada
2023.10.21 © Lypkie, P.
Sminthurinus quadrimaculatus f. "niger" from Canada
2023.10.21 © Lypkie, P.
Sminthurinus quadrimaculatus f. "niger" from Canada
2023.10.21 © Lypkie, P.

            Syn.: Smynthurus(sic) remotus Folsom, JW, 1896:448
            Syn.: Sminthurinus concolor Hammer, 1953 =?
            Syn.: Sminthurus concolor Christiansen, K et Bellinger, P, 1981:1117 =?
            Syn.: Sminthurinus concolor Christiansen, K et Bellinger, P, 1981:1117 =?
            Syn.: Sminthurus concolor Christiansen, K et Bellinger, P, 1998:1269 =?
            Syn.: Sminthurinus concolor Christiansen, K et Bellinger, P, 1998:1269 =?
            Syn.: Sminthurinus concolor Babenko, A, Stebaeva, S et Turnbull, MS, 2019:99 =?
            Syn.: Sminthurinus quadrimaculatus f. bimaculatus Maynard, EA, 1951:230
Sminthurinus quadrimaculatus f. bimaculatus from the USA
After Maynard EA, 1951 Plate XXVI Fig.490
Sminthurinus quadrimaculatus f. bimaculatus from the USA
After Maynard EA, 1951 Plate XXVI Fig.491

             Syn.: Sminthurinus quadriomaculatus(sic) Richards, WR, 1968:23 ISS
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2003 © Schoenherr, J.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2007.05.17 © Pilon, C.
Sminthurinus quadrimaculatus f. bimaculatus from Canada
2007.05.17 © Pilon, C.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2007.12.16 © Moorehead, C.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2008.06.03 © Schoenherr, J.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2008.06.04 © Schoenherr, J.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
Spotted: ready to moult
2008.09.08 © Murray, T.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2011.12.03 © Justis, S.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2011.12.03 © Justis, S.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2011.12.03 © Justis, S.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2012.01.07 © Justis, S.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2012.01.07 © Justis, S.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2012.01.07 © Justis, S.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2014.11.05 © Rosenfeld, J.
Sminthurinus quadrimaculatus f. bimaculatus from Canada
2021.01.14 © Rennert, M.
Sminthurinus quadrimaculatus f. bimaculatus from Canada
2021.01.14 © Rennert, M.


            Syn.: Sminthurinus minutus f. bipunctatus Maynard, EA, 1951:235
Sminthurinus minutus f. bipunctatus from the USA
After Maynard EA, 1951 Plate XXIII Fig.403
Sminthurinus minutus f. bipunctatus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.536
Sminthurinus quadrimaculatus f. bipunctatus from the USA
2008.02.07 © Gross, J.
Sminthurinus quadrimaculatus f. bipunctatus from the USA
2008.02.07 © Gross, J.
Sminthurinus quadrimaculatus f. bipunctatus from the USA
2008.02.07 © Gross, J.
Sminthurinus quadrimaculatus f. bipunctatus from the USA
2013.07.13 © Israel, D.
Sminthurinus quadrimaculatus f. bipunctatus from the USA
Spotted: ready to moult
2016.04.02 © Justis, S.
Sminthurinus quadrimaculatus f. bipunctatus from the USA
2017.11.14 © Dobbs, Br.
Sminthurinus quadrimaculatus f. bipunctatus from Canada
2021.01.14 © Rennert, M.


            Syn.: Sminthurinus brunneus Maynard, EA, 1951:226
Sminthurinus brunneus from the USA
After Maynard EA, 1951 Plate XXIII Fig.400
Sminthurinus brunneus from the USA
After Maynard EA, 1951 Plate XXIII Fig.401


            Syn.: Sminthurinus downsi Maynard, EA, 1951:225
Sminthurinus downsi from the USA
After Maynard EA, 1951 Plate XXII Fig.377

             Syn.: Sminthurinus dowsi(sic) Stach, J, 1956:102 ISS

            Syn.: Sminthurinus quadrimaculatus f. latipictus Maynard, EA, 1951:231
Sminthurinus quadrimaculatus f. latipictus from the USA
After Maynard EA, 1951 Plate XXIII Fig.391
Sminthurinus quadrimaculatus f. latipictus from the USA
After Maynard EA, 1951 Plate XXIII Fig.392
Sminthurinus quadrimaculatus from the USA
Pale form
2006 © McClarin, J.


            Syn.: Sminthurinus quadrimaculatus ssp. maynardi Snider, RJ, 1967:223



Sminthurinus quadrimaculatus from the USA
After Maynard EA, 1951 Plate XXIII Fig.393
Sminthurinus quadrimaculatus from the USA
After Maynard EA, 1951 Plate XXIII Fig.394
Sminthurinus quadrimaculatus from the USA
After Maynard EA, 1951 Plate XXVI Fig.489
             Syn.: Sminthurines(sic) cuadrimaculatus(sic) Palacios-Vargas, 1979 ISS
             Syn.: Sminthurus quadrimaaculatus(sic) Vázquez & Palacios-Vargas, 1987 ISS
Sminthurinus quadrimaculatus from the USA
2007 © McClarin, J.
 
Sminthurinus quadrimaculatus from the USA
First instar
2007.04.12 © Murray, T.
Sminthurinus quadrimaculatus from the USA
Second instar
2007.04.15 © Murray, T.
Sminthurinus quadrimaculatus from the USA
Second instar
2007.04.15 © Murray, T.
Sminthurinus quadrimaculatus from the USA
2007.10.23 © McClarin, J.
Sminthurinus quadrimaculatus from the USA
2008.10.04 © Murray, T.
Sminthurinus quadrimaculatus juv. from Canada
2016.08.28 © Levac, G.
Sminthurinus quadrimaculatus from the USA
2016.09.20 © Rosenfeld, J.
Sminthurinus quadrimaculatus from the USA
2021.05.17 © Coogler, J.
Sminthurinus quadrimaculatus from the USA
2021.05.17 © Coogler, J.
Sminthurinus quadrimaculatus from the USA
2021.08.23 © Cowles, J.H.





Species quadripunctatus (Edinger, 1937) Christiansen & Bellinger, 1981

Species reticulatus Cassagnau, P, 1964:481

Sminthurinus reticulatus from France
After Cassagnau P, 1964

            Syn.: Sminthurinus aureus var. transversalis Hüther, 1961
            Syn.: Sminthurinus aureus Ellis, 1976



Sminthurinus reticulatus from Sweden
After Fjellberg A, 2006 ©
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from France
2007.09.08 © Lebeaux, P.
Sminthurinus reticulatus from France
2008.12.07 © Jacquemin, G.
Sminthurinus reticulatus from France
2011.11.13 © Lebeaux, P.
Sminthurinus reticulatus from France
2011.11.13 © Lebeaux, P.
Sminthurinus reticulatus from the UK
2012.03.30 © Nurcombe, E.
Sminthurinus reticulatus from the UK
2012.03.30 © Nurcombe, E.
Sminthurinus reticulatus from the UK
2012.04.01 © Nurcombe, E.
Sminthurinus reticulatus from the UK
2012.04.11 © Nurcombe, E.
Sminthurinus reticulatus from the UK
2012.07.20 © Murray, A.
Sminthurinus reticulatus from the UK
2012.07.22 © Murray, A.
Sminthurinus reticulatus juv. from the UK
2012.07.22 © Murray, A.
Sminthurinus reticulatus from the UK
2012.09.13 © Nurcombe, E.
Sminthurinus reticulatus from the UK
2012.09.26 © Nurcombe, E.
Sminthurinus reticulatus from the UK
2012.12.09 © Shaw, P.
Sminthurinus reticulatus from the UK
2012.12.09 © Shaw, P.
Sminthurinus reticulatus from the UK
2012.12.09 © Shaw, P.
Sminthurinus reticulatus from the UK
2012.12.23 © Phillips, E.
Sminthurinus reticulatus from the UK
2013.01.04 © Barton, T.
Sminthurinus reticulatus from the UK
2013.02.12 © Phillips, E.
Sminthurinus reticulatus from the UK
2013.02.15 © Phillips, E.
Sminthurinus reticulatus from the UK
2013.07.01 © Phillips, E.
Sminthurinus reticulatus from the UK
2013.11.04 © Phillips, E.
Sminthurinus reticulatus from the UK
2014.01.10 © Phillips, E.
Sminthurinus reticulatus from the UK
2014.01.25 © Phillips, E.
Sminthurinus reticulatus from the UK
2014.08.20 © Phillips, E.
Sminthurinus reticulatus juv. from Germany
2014.11.01 © Kluck, A.
Sminthurinus reticulatus from Holland
2015.10.09 © van Duinen, J.
Sminthurinus reticulatus from Belgium
2016.12.02 © Huskens, M.-L.
Sminthurinus reticulatus from Belgium
Grooming upside-down
2016.12.02 © Huskens, M.-L.
Sminthurinus reticulatus from Belgium
2017.02.04 © Huskens, M.-L.
Sminthurinus reticulatus from Belgium
2017.02.04 © Huskens, M.-L.
Sminthurinus reticulatus from Belgium
2017.02.04 © Huskens, M.-L.
Sminthurinus reticulatus juv. from Belgium
2017.10.17 © Huskens, M.-L.
Sminthurinus reticulatus from Belgium
Eyepatch with 7 ocelli (C absent, D reduced)
2017.12.21 © Huskens, M.-L.
Sminthurinus cf. reticulatus from Belgium
Everted collophore vesicles
2017.12.24 © Huskens, M.-L.
Sminthurinus reticulatus juv. from Belgium
2018.05.12 © Kamsteeg, G.
Sminthurinus reticulatus from Belgium
2018.10.18 © Huskens, M.-L.
Sminthurinus reticulatus ♂ from Belgium
2018.11.03 © Huskens, M.-L.
Sminthurinus reticulatus juv. from Holland
2019.12.17 © Kamsteeg, G.
Sminthurinus reticulatus from France
2020.01.18 © Joachim, J.
Sminthurinus reticulatus juv. from France
2020.01.18 © Joachim, J.
Sminthurinus reticulatus from France
2020.01.18 © Joachim, J.
Sminthurinus reticulatus from France
2020.01.18 © Joachim, J.
Sminthurinus reticulatus from Belgium
Caudal aspect mimicry
2020.07.01 © Huskens, M.-L.
Sminthurinus reticulatus from Belgium
2021.12.30 © Huskens, M.-L.
Sminthurinus nr reticulatus from Holland
2024.04.08 © Kamsteeg, G.
Sminthurinus nr reticulatus from Holland
2024.04.08 © Kamsteeg, G.

Species salti Delamare Deboutteville, C, 1953:828

Species sayi Denis, J-R, 1933:276

            Syn.: Sminthurinus Sayi(sic) Denis, J-R, 1933:276 IOS
            Syn.: Sminthurinus Sayi(sic) Denis, J-R, 1933:277 IOS
            Syn.: Sminthurinus Sayi(sic) Denis, J-R, 1933:278 IOS
            Syn.: Sminthurinus Sayi(sic) Denis, J-R, 1933:279 IOS
            Syn.: Sminthurinus Sayi(sic) Denis, J-R, 1933:280 IOS
            Syn.: Sminthurinus Sayi(sic) Denis, J-R, 1933:283 IOS
            Syn.: Sminthurinus Sayi(sic) Denis, J-R, 1933:315 IOS
            Syn.: Sminthurinus Sayi(sic) Denis, J-R, 1933:320 IOS
            Syn.: Sminthurinus Sayi(sic) Delamare Deboutteville, C et Massoud, Z, 1963:205 IOS

Species signatus (Krausbauer, 1898 partim) Börner, 1900

            Syn.: Sminthurus aureus var. signata(sic) Krausbauer, 1898 partim IOS
            Syn.: Sminthurinus aureus var. signata(sic) Börner, 1901 IOS
            Syn.: Sminthurus aureus var. signata(sic) Krausbauer, 1901 IOS
            Syn.: Sminthurinus aureus var. signata(sic) Axelson, 1904 IOS
            Syn.: Sminthurinus aureus var. signata(sic) Axelson, WM, 1905:794 IOS
            Syn.: Sminthurinus aureus var. signata(sic) Linnaniemi, WM, 1912:283 IOS
            Syn.: Sminthurinus aureus f. signata(sic) Handschin, E in Dahl, F, 1929:113 IOS
Sminthurinus aureus f. signatus from Japan
After Uchida H, 1954 Fig.1A
Sminthurinus aureus f. signatus from Japan
After Uchida H, 1954 Fig.1B

            Syn.: Sminthurinus aureus var. signata(sic) Stach, J, 1956:122 IOS
            Syn.: Sminthurinus quadrilineata ab. signata(sic) Stach, J, 1956:122 IOS
            Syn.: Sminthurinus elegans var. signata(sic) Stach, J, 1956:127 IOS
            Syn.: Sminthurinus aureus var. signata(sic) Gisin, H, 1960:279 IOS
Sminthurinus signatus from Denmark
After Fjellberg A, 2007 Fig.13.4
Sminthurinus signatus from France
2016.12.07 © Legrand, T.
Sminthurinus signatus from France
2016.12.07 © Legrand, T.
Sminthurinus signatus from France
2016.12.07 © Legrand, T.
Sminthurinus signatus from France
2016.12.07 © Legrand, T.
Sminthurinus signatus from France
2017.11.05 © Picard, J.
Sminthurinus signatus from France
2017.11.05 © Picard, J.
Sminthurinus signatus from France
2017.11.05 © Picard, J.
Sminthurinus signatus from France
2017.11.05 © Picard, J.
Sminthurinus signatus from France
2017.11.18 © Tiky, S.
Sminthurinus signatus from France
With distinct middorsal pattern
2017.12.10 © Picard, J.
Sminthurinus signatus from France
2018.01.09 © Garcelon, P.
Sminthurinus signatus from France
2018.06.19 © van Duinen, J.
Sminthurinus signatus from France
2019.02.13 © Garcelon, P.
Sminthurinus signatus from France
2019.12.06 © Garcelon, P.

            Syn.: Sminthurus aureus var. signata(sic) Mehrafrooz Mayvan, M, Sadeghi-Namaghi, H et Shayanmehr, M, 2021:8 IOS


Species signifer (Fitch, A, 1863:675)

            Syn.: Smynthurus(sic) niger Schött, H, 1891:12 =?


             Syn.: Sminthurus signifier(sic) Salmon, JT, 1964:599 ISS

Species speciosus Yosii, R, 1970:19

            Syn.: Sminthurinus (niger-group) specious(sic) Yosii, R, 1970:19 IOS
Sminthurinus speciosus from Japan
After Yosii R, 1970 Fig.12A



Species splendidus (Womersley, 1933) Greenslade, P in Houston, WWK, 1994:133

Parakatianna splendida from Australia
After Womersley H, 1939 Pl.1 Fig.4

             Syn.: Sminthurinus splendida(sic) Greenslade, P in Houston, WWK, 1994:133 ISS
             Syn.: Sminthurinus splendida(sic) Greenslade, P in Houston, WWK, 1994:159 ISS
Sminthurinus splendidus from Australia
2023.07.05 © Cassidy, E.
Sminthurinus splendidus from Australia
2023.07.05 © Cassidy, E.



Species subalpinus Itoh, 2000

Species suborientalis Stach, J, 1965:367

Species timorensis Yoshii, R et Suhardjono, YR, 1992:92

Species transversalis Axelson, WM, 1905:794

            Syn.: Sminthurinus reticulatus Fjellberg, A, 2007:197 =?


Sminthurinus transversalis from Finland
Det. Axelson, W.M., 1902
2006 © Fjellberg, A.
Sminthurinus transversalis ♀ from Denmark
After Fjellberg A, 2007 Fig.13.6.left
Sminthurinus transversalis ♂ from Denmark
After Fjellberg A, 2007 Fig.13.6.right

             Syn.: Sminthurinus transvernalis(sic) Falahati HosseinAbad, A, Kheirodin, A et Bretfeld, G, 2013:208 ISS
             Syn.: Smynthurinus(sic) transvernalis(sic) Shayanmehr, M, Yahyapour E, Kahrarian, M et Lafooraki EY, 2013:73 ISS
Sminthurinus nr transversalis from the UK
2021.01.15 © Uncle, F.
Sminthurinus nr transversalis from the UK
2021.01.15 © Uncle, F.
Sminthurinus nr transversalis from the UK
2021.01.15 © Uncle, F.
Sminthurinus nr transversalis from the UK
2021.01.15 © Uncle, F.

             Syn.: Sminthurinus transvernalis(sic) Mehrafrooz Mayvan, M, Greenslade, P et Sadeghi-Namaghi, H, 2023:57 ISS


Species tricolor Schött, H, 1917:54

Sminthurinus tricolor from Australia
After Schött H, 1917 Taf.4 Fig.5

Species trinotatus Axelson, WM, 1905:794, B

            Syn.: Sminthurinus igniceps var. trinotata(sic) Axelson, WM, 1905:794 IOS
            Syn.: Sminthurus biflavopunctatus Collinge, WE et Shoebotham, JW, 1909:9
Sminthurus biflavopunctatus from the UK
After Collinge WE & Shoebotham JW, 1909 Pl.III Fig.1

             Syn.: Sminthurinus bifavopunctatus(sic) Salmon, JT, 1964:582 ISS
             Syn.: Sminthurinus aureus var. bimaculata(sic) Carpenter H in Collinge WE & Shoebotham JW, 1909:10 ISS
            Syn.: Sminthurinus fenestratus Börner, C, 1909:124
            Syn.: Sminthurinus fenestratus Uchida, 1952
            Syn.: Sminthurinus fenestratus Yosii, R, 1977:164
             Syn.: Sminthurinus trinotatus f. incompleta(sic) Stach, J, 1965:367 ISS



             Syn.: Sminthurinus igniceps f. trinotata(sic) Handschin, E in Dahl, F, 1929:113 ISS
             Syn.: Sminthurinus igniceps var. trinotata(sic) Buitendijk, AM, 1933:66 ISS
            Syn.: Sminthurus igniceps var. trinotata(sic) Itoh, R et Zhao, L-J, 2000:175 IOS
Sminthurinus trinotatus from Germany
2006.12.29 © Kohl, F.
Sminthurinus trinotatus from Belgium
2008.06.09 © Vuijlsteke, M.
Sminthurinus trinotatus from the UK
2008.07.22 © Shaw, P.
Sminthurinus trinotatus juv. from France
2008.12.07 © Jacquemin, G.
Sminthurinus trinotatus from France
2008.12.07 © Jacquemin, G.
Sminthurinus trinotatus from the UK
After Ardron PA, 2009 p.17
Sminthurinus trinotatus from the UK
2009.09.01 © Thomason, R.
Sminthurinus trinotatus from the UK
2010.10.23 © Jones, A.
Sminthurinus trinotatus from Holland
Distinct abdominal segments 5 & 6
2011.07.08 © van Duinen, J.
Sminthurinus trinotatus from Germany
2012.04.14 © Schmitt, R.
Sminthurinus trinotatus from the UK
2012.10.03 © Murray, A.
 
Sminthurinus trinotatus from the UK
2012.10.03 © Murray, A.
Sminthurinus trinotatus from the UK
2012.10.03 © Murray, A.
Sminthurinus trinotatus from the UK
2012.10.31 © Jones, A.
Sminthurinus trinotatus from Holland
2013.08.19 © van Duinen, J.
Sminthurinus trinotatus from Holland
2013.08.19 © van Duinen, J.
Sminthurinus trinotatus from the UK
2013.11.15 © Horton, M.
Sminthurinus trinotatus juv. from the UK
2013.12.11 © Nurcombe, E.
Sminthurinus trinotatus juv. from the UK
2013.12.11 © Nurcombe, E.
Sminthurinus trinotatus from Germany
2014.05.06 © Beck, K.
Sminthurinus trinotatus juv. from Germany
First instar
2014.05.06 © Beck, K.
Sminthurinus trinotatus from Japan
2014.05.15 © Hara, Y.
Sminthurinus trinotatus from Germany
2014.10.28 © Beck, K.
Sminthurinus trinotatus from Germany
Dorsal aspect mimicry
2014.11.02 © Beck, K.
Sminthurinus trinotatus juv. from Germany
2014.11.02 © Beck, K.
Sminthurinus trinotatus from Germany
2014.11.02 © Beck, K.
Sminthurinus trinotatus from the UK
2015.07.27 © Boardman, P.
Sminthurinus trinotatus from Germany
2015.09.01 © Jäschke, J.
Sminthurinus trinotatus from the UK
2015.11.16 © Horton, M.
Sminthurinus trinotatus from the UK
Abdominal bothriotricha in red cups
2015.11.16 © Horton, M.
Sminthurinus trinotatus from the UK
Caudal aspect
2015.11.16 © Horton, M.
Sminthurinus trinotatus from the UK
Broad white "eyebrows"
2015.11.16 © Horton, M.
Sminthurinus trinotatus from the UK
Facial aspect
2015.11.16 © Horton, M.
Sminthurinus trinotatus from Belgium
2016.01.06 © Huskens, M.-L.
Sminthurinus trinotatus from Japan
2016.07.21 © Hara, Y.
Sminthurinus trinotatus from the UK
Caudal aspect
2017.09.25 © Phillips, E.
Sminthurinus trinotatus from Holland
2017.10.26 © Wieringa, P.
Sminthurinus trinotatus ♂ from Belgium
Grooming
2018.04.26 © Huskens, M.-L.
Sminthurinus trinotatus from Belgium
2018.08.06 © Huskens, M.-L.
Sminthurinus trinotatus from Holland
Abd.5 not included in great abdomen
2019.01.09 © Kamsteeg, G.
Sminthurinus trinotatus from Belgium
Gap between ocellus G and H
2019.08.29 © Huskens, M.-L.
Sminthurinus trinotatus ♀ from Belgium
Anal valves with sub-anal appendage (s.a.a.) + bifurcate macroseta (*)
2019.09.05 © Huskens, M.-L.
Sminthurinus trinotatus from Holland
2019.11.05 © de Haas, M.
Sminthurinus trinotatus from Holland
2019.11.05 © de Haas, M.
Sminthurinus trinotatus from Holland
Dorsal cephalic pattern
2019.11.05 © de Haas, M.
Sminthurinus trinotatus from Belgium
2019.11.19 © Huskens, M.-L.
Sminthurinus trinotatus ♂ from Belgium
2020.04.01 © Huskens, M.-L.
Sminthurinus trinotatus juv. from Belgium
2020.04.01 © Huskens, M.-L.
Sminthurinus trinotatus juv. from Belgium
2020.07.01 © Huskens, M.-L.
Sminthurinus trinotatus from Belgium
Caudal aspect mimicry of face
2020.07.01 © Huskens, M.-L.
Sminthurinus trinotatus juv. from Holland
2021.03.06 © Kamsteeg, G.
Sminthurinus trinotatus ♀ & ♂ from Belgium
2021.10.05 © Huskens, M.-L.
Sminthurinus trinotatus ♂ from Belgium
Spermatophore deposited
2021.10.05 © Huskens, M.-L.
Sminthurinus trinotatus from Taiwan
After Cheng H-J & al, 2022 Fig.4C
Sminthurinus trinotatus ♂ from Belgium
♂ guards his spermatophore
2022.09.09 © Huskens, M.-L.
Sminthurinus trinotatus ♂ & ♂ from Belgium
♂'s fighting to protect/destroy spermatophore
2022.09.09 © Huskens, M.-L.
Sminthurinus trinotatus ♂ & ♀ from Belgium
♂ steps back from his spermatophore to free space for ♀ passing by
2022.09.10 © Huskens, M.-L.
Sminthurinus trinotatus ♂ & ♂ from Belgium
♂ defends his spermatophore against intruder
2022.09.11 © Huskens, M.-L.
Sminthurinus trinotatus ♂ & ♂ from Belgium
♂'s fight eachother to defend/destroy spermatophore
2022.09.11 © Huskens, M.-L.





Species tuberculatus Delamare Deboutteville, C et Massoud, Z, 1963:201

            Syn.: Sminthurinus sp. tuberculatus group King, Greenslade P & Hutchinson, 1985


Species tucumanensis Delamare Deboutteville & Massoud, 1963:205

Species tunicatus Salmon, JT, 1954:216

Sminthurinus tunicatus from New Zealand
2009 © Te Papa Museum

Species victorius Nguyen, 2001

Species wakhanicus Yosii, 1966

Species woringeri Delamare Deboutteville, C et Massoud, Z, 1963:207

            Syn.: Sminthurinus Woringeri(sic) Delamare Deboutteville, C et Massoud, Z, 1963:207 IOS





Genus Borgesminthurinus Palacios-Vargas JG in Palacios-Vargas, JG et Vacaflores-Argandoña, MR, 2020:202

Species andinus Palacios-Vargas, JG et Vacaflores-Argandoña, MR, 2020:207, t.t.

Borgesminthurinus andinus from Bolivia
After Palacios-Vargas JG & Vacaflores-Argandoña, 2020 Fig.19
Borgesminthurinus andinus from Bolivia
After Palacios-Vargas JG & Vacaflores-Argandoña, 2020 Fig.20


Genus Pseudokatianna Salmon, JT, 1944:179

  Syn.: Katiannini Metakatianna Richards, WR, 1968:22 partim


Species campbellensis Salmon, JT, 1949:52

            Syn.: Pseudokatianna cambellensis(sic) Salmon, JT, 1949:52 IOS

Species fagophila Salmon, JT, 1946:50

Species fasciata (Salmon, JT, 1944:178)

Species livida (Salmon, JT, 1943:2) Salmon, JT, 1944:181

Species lutea Salmon, JT, 1946:51

Species minuta Salmon, JT, 1946:53

Species nigretalba Salmon, JT, 1944:179, t.t.

  Subspecies aurea Salmon, JT, 1944:180

Species niveovata Salmon, JT, 1946:48

Pseudokatianna niveovata from New Zealand
After Salmon JT, 1946 Fig.42

  Subspecies nigra Salmon, JT, 1946:50

Species triclavata Salmon, JT, 1949:53

Species triverrucata Salmon, JT, 1944:181

Species umbrosalata Salmon, JT, 1946:52

Species zebra Salmon, JT, 1946:47

Pseudokatianna zebra from New Zealand
After Salmon JT, 1946 Fig.36


Genus Stenognathellus Stach, J, 1956:95

  Syn.: Sminthuridinae Stenognathellus Cassagnau, P, 1953:226


Remark by Ellis, W.N. & Bellinger, P.F. 1973:54: Stenognathellus Cassagnau, 1953 is an unavailable name (art. 13b): type species not designated.

Sminthurinus (left) versus Stenognathellus (right) from Holland
Swollen occiput (v)
2021.06.03 © van Bezouw, R.

Species cassagnaui Yosii, 1966

Stenognathellus cassagnaui from the UAE
After Schulz H-J & van Harten A, 2013 Pl.4

Species denisi Cassagnau, P, 1953:227, t.t.

            Syn.: Stenognathellus Denisi(sic) Cassagnau, P, 1953:227,tt IOS
Stenognathellus Denisi(sic) from France
After Cassagnau P, 1953 p.227 Fig.A
Stenognathellus Denisi(sic) from France
After Cassagnau P, 1953 p.227 Fig.B


            Syn.: Sminthurinus sp. nov. Denis, J-R, 1933:293
            Syn.: Sminthurinus sp. nov. Cassagnau, P, 1953:226 =?




Stenognathellus denisi from Belgium
2017.10.19 © Huskens, M.-L.
Stenognathellus denisi from Belgium
Ocelli C+D indistinct
2017.10.19 © Huskens, M.-L.
Stenognathellus denisi from France
2017.10.23 © Picard, J.
Stenognathellus denisi from Belgium
2017.11.09 © Huskens, M.-L.
Stenognathellus denisi from Belgium
2017.11.09 © Huskens, M.-L.
Stenognathellus denisi from Belgium
2017.11.09 © Huskens, M.-L.
Stenognathellus denisi from Belgium
Bipartite gray eyebrows
Ocelli C and D indistinct
2017.11.09 © Huskens, M.-L.
Stenognathellus denisi from Belgium
2018.10.11 © Huskens, M.-L.
Stenognathellus denisi from Belgium
Producing egg
2018.10.12 © Huskens, M.-L.
Stenognathellus denisi from Belgium
Chorion of egg hardens after 10 min.
2018.10.12 © Huskens, M.-L.
Stenognathellus denisi from Belgium
Egg deposited on substrate
2018.10.12 © Huskens, M.-L.
Stenognathellus denisi from Belgium
Ant.3 with wart (*)
2019.07.21 © Huskens, M.-L.
Det.: van Bezouw, R.
Stenognathellus denisi juv. from Belgium
2019.07.22 © Huskens, M.-L.
Stenognathellus denisi from Belgium
2019.08.02 © Huskens, M.-L.
Stenognathellus denisi from Belgium
2019.08.02 © Huskens, M.-L.
Stenognathellus denisi from Belgium
Ocelli C and D indistinct
2019.08.02 © Huskens, M.-L.
Stenognathellus denisi from Belgium
Abd.5 not fused with great abdomen
2019.08.02 © Huskens, M.-L.
Stenognathellus denisi from Belgium
Triangular head
2020.06.17 © Huskens, M.-L.
Stenognathellus denisi from Belgium
Ocelli C and D indistinct
2020.06.17 © Huskens, M.-L.
Stenognathellus denisi from Belgium
Abd.5 not fused with great abdomen
2020.09.27 © Huskens, M.-L.
Stenognathellus denisi from Belgium
Ocelli C and D indistinct
2020.09.27 © Huskens, M.-L.
Stenognathellus denisi from Belgium
Ocelli C+D indistinct
2020.11.02 © Huskens, M.-L.
Stenognathellus denisi from Belgium
Abd.5 marked by 2 sutures
2020.11.02 © Huskens, M.-L.
Stenognathellus denisi from Belgium
Abd.5 marked by 2 sutures
2020.11.02 © Huskens, M.-L.
Stenognathellus denisi from Belgium
2020.11.03 © Huskens, M.-L.
Stenognathellus denisi from Holland
Mouthcone
2021.05.31 © van Bezouw, R.
Stenognathellus denisi from Australia
Abd.5 not included in large abdomen
2021.06.13 © Hobern, D.
Stenognathellus denisi from Australia
Ocelli C+D distinctly smaller
2021.06.13 © Hobern, D.



Species polygonalis Hüther, 1967

Species stenognathus (Börner, 1907) Betsch, 1980

            Syn.: Sminthurinus stenognathus ssp. similis Najt & Rapoport, 1965


Stenognathellus stenognathus from South Africa
© Biological Museum Lund University, 2019


Genus Katiannellina Delamare Deboutteville, C et Massoud, Z, 1963:217

Species patagonica Delamare Deboutteville, C et Massoud, Z, 1963:217, t.t.


Genus Betschurinus Dallai, R et Martinozzi, I, 1980:31

Species farmae Dallai, R et Martinozzi, I, 1980:33, t.t.


Genus Dalianus Cassagnau, 1969

Species galae Cassagnau, 1969


Genus Katiannina Maynard EA et Downs WL in Maynard, EA, 1951:242

  Syn.: ... Deuterosminthurus Wray, 1948
  Syn.: ... Polyclavella Stach, 1955
  Syn.: Katiannini Metakatianna Richards, WR, 1968:22 partim
  Syn.: ... Metakatianna Denis, 1933 u


  Syn.: Bourletiellini Katiannia(sic) Snider, RJ, 1967:225 ISS
Katiannina sp. from the USA
2014.01.13 © Philonatura, N.



Species sp.1 Janssens F & Kyron TF, 2013

Katiannina? sp.1 from Canada
With distinctly reddish eyepatches
2013.01.17 © Kyron, T.F.
Katiannina? sp.1 from Canada
With distinctly reddish eyepatches
2013.01.17 © Kyron, T.F.
Katiannina? sp.1 ♀ from Canada
Facial chaetotaxy
2013.01.21 © Kyron, T.F.
Katiannina? sp.1 ♀ from Canada
Female palmate sub-anal appendages (s.a.a.)
2013.01.21 © Kyron, T.F.
Katiannina? sp.1 ♀ from Canada
Anogenital abdominal segments
2013.01.21 © Kyron, T.F.
Katiannina? sp.1 ♀ from Canada
Ocelli C & H subequal in size but distinctly different in colour
2013.01.21 © Kyron, T.F.
Katiannina? sp.1 ♀ from Canada
Fourth antennal segment not subdivided
2013.01.21 © Kyron, T.F.
Katiannina? sp.1 from the USA
2018.12.14 © Pearson, R.R.
Katiannina? sp.1 from the USA
2018.12.14 © Pearson, R.R.
Katiannina? sp.1 from the USA
2018.12.26 © Pearson, R.R.
Katiannina? sp.1 from the USA
2019.01.10 © Pearson, R.R.
Katiannina? sp.1 from the USA
2021.01.03 © Pearson, R.R.
Katiannina? sp.1 from the USA
Dark colour form
2021.01.07 © Pearson, R.R.
Katiannina? sp.1 from Canada
2021.03.15 © Hochhalter, R.
Katiannina? sp.1 from Canada
2021.03.15 © Hochhalter, R.
Katiannina? sp.1 from Canada
Abdominal pattern
2021.03.15 © Hochhalter, R.
Katiannina? sp.1 from Canada
Small abdomen
2021.03.15 © Hochhalter, R.
Katiannina? sp.1 from Canada
Foot with long unguiculus
2021.03.15 © Hochhalter, R.
Katiannina? sp.1 from Canada
2024.02.18 © Lypkie, P.
Katiannina? sp.1 from Canada
Ocelli D+H reddish
Ocellus C black
2024.02.18 © Lypkie, P.

Species macgillivrayi (Banks, 1897) Maynard, EA, 1951:242, t.t.

            Syn.: Deuterosminthurus altamontus Wray, 1948
            Syn.: Deuterosminthurus batrachos Wray, 1948
            Syn.: Polykatianna batchi Snider, 1969


Katiannina macgillivrayi from the USA
After Maynard EA Plate XXII Fig.377a
Katiannina macgillivrayi from the USA
After Maynard EA Plate XXIX Fig.552
Katiannina macgillivrayi from the USA
After Maynard EA Plate XXIX Fig.553
Katiannina macgillivrayi from the USA
2006 © Bernard, E.C.
Katiannina macgillivrayi from the USA
2007.05.15 © Murray, T.
Katiannina macgillivrayi from the USA
2008.12.14 © Fleming, K.
Katiannina macgillivrayi from the USA
2009.05.08 © Loser, I.
Katiannina macgillivrayi from the USA
2009.11.26 © Justis, S.
Katiannina macgillivrayi from the USA
2009.11.26 © Justis, S.
Katiannina macgillivrayi from the USA
2009.11.26 © Justis, S.
Katiannina macgillivrayi from the USA
2009.11.26 © Justis, S.
Katiannina macgillivrayi from the USA
2009.11.26 © Justis, S.
Katiannina macgillivrayi from the USA
2009.11.26 © Justis, S.
Katiannina macgillivrayi ♀ from the USA
2009.11.26 © Justis, S.
Katiannina macgillivrayi from the USA
2010.04.17 © Skvarla, M.
Katiannina macgillivrayi juv. from the USA
2011.04.21 © Eiseman, C.
Katiannina macgillivrayi from the USA
2012.02.20 © Fleming, K.
Katiannina macgillivrayi juv. from the USA
2012.03.27 © Eiseman, C.
Katiannina macgillivrayi from the USA
Red eyepatch
2012.04.21 © Eiseman, C.
Katiannina macgillivrayi from the USA
2012.04.21 © Eiseman, C.
Katiannina macgillivrayi from the USA
2015.01.19 © Fleming, K.
Katiannina macgillivrayi from the USA
2018.01.22 © Fleming, K.
Katiannina macgillivrayi from the USA
2024.02.12 Van Epps, R.
Katiannina macgillivrayi from the USA
2024.02.12 Van Epps, R.

Species subalpina Itoh, 2000


Genus Neokatianna Snider, RJ, 1989:541

Species acantholaema (Snider, RJ, 1989:542) Bretfeld, G, 1999:99, t.t.

            Syn.: Sminthurinus (Neokatianna) acantholaema(sic) Snider, RJ, 1989:542,tt IOS
Sminthurinus (Neokatianna) acantholaema(sic) from the USA
After Snider RJ, 1989 Fig.1
Sminthurinus (Neokatianna) acantholaema(sic) from the USA
After Snider RJ, 1989 Fig.2

            Syn.: Sminthurinus (Neokatianna) acantholaema(sic) Christiansen, K et Bellinger, P, 1998:1266 IOS
Neokatianna acantholaema from the USA
2009.04.12 © Justis, S.
Neokatianna acantholaema from the USA
2009.04.12 © Justis, S.
Neokatianna acantholaema from the USA
Bothriotricha ABC forming large obtuse angle
Bothriotrichum D present
2009.04.12 © Justis, S.
Neokatianna acantholaema from the USA
Ant.4 weakly annulated with 10-11 subdivisions
2009.04.12 © Justis, S.
Neokatianna acantholaema from the USA
2012.01.29 © Kouri, J.
Neokatianna acantholaema from the USA
2012.01.29 © Kouri, J.
?Neokatianna acantholaema? from the USA
2017.03.27 © Dobbs, Br.
Neokatianna acantholaema from the USA
2020.02.27 © Solomon.
Neokatianna acantholaema from the USA
2020.05.03 © Solomon.
Neokatianna acantholaema from the USA
2021.05.05 © Pavy, R.
Neokatianna nr acantholaema from Poland
2021.06.14 © Antosik-Zoladek, J.
Neokatianna nr acantholaema from Poland
2021.06.15 © Antosik-Zoladek, J.
Neokatianna nr acantholaema from Poland
2021.06.15 © Antosik-Zoladek, J.
Neokatianna nr acantholaema from Poland
Dark interantennal spot
2021.06.15 © Antosik-Zoladek, J.
Neokatianna nr acantholaema from Poland
Ant.4 subdivided
2021.06.15 © Antosik-Zoladek, J.
Neokatianna nr acantholaema from Poland
Ocelli C+D smaller
2021.06.15 © Antosik-Zoladek, J.
Neokatianna nr acantholaema from Poland
Bothriotricha ABC form obtuse angle
2021.06.20 © Antosik-Zoladek, J.
Neokatianna nr acantholaema from Poland
Abd.5 with bothriotrichum D
2021.06.22 © Antosik-Zoladek, J.
Neokatianna acantholaema from the USA
2024.02.21 © AVNJ.




Genus Rusekianna Betsch, J-M, 1977:61

Rusekianna from Norway
2009.06.03 © Fjellberg, A.
Pores at base of tibiotarsus

Species albifrons (Tullberg, 1871) Fjellberg, A, 2007:il

            Syn.: Rusekianna sibirica Bretfeld, 2002 =?


             Syn.: Sminthurus albifronti(sic) Reuter, OM, 1876:85 ISS
             Syn.: Sminthurus albifronti(sic) Reuter, OM, 1895:12 ISS
             Syn.: Sminthurinus Albifrons(sic) Denis, J-R, 1933:279 ISS


Sminthurinus albifrons from Norway
2006 © Fjellberg, A.
Rusekianna albifrons from France
2007.03.17 © Lebeaux, P.
Rusekianna albifrons from France
2007.03.17 © Lebeaux, P.
Rusekianna albifrons from Portugal
2009.04.21 © Marcelino, J.
Rusekianna albifrons juv. from France
2020.09.23 © Reinach, Br.
Rusekianna albifrons juv. from France
2020.09.23 © Reinach, Br.
Rusekianna albifrons juv. from France
2020.09.23 © Reinach, Br.

Species bescidica Smolis, A et Skarzynski, D, 2006:317

Species intermedia (Snider, 1978) Babenko, A, Stebaeva, S & Turnbull, MS, 2019:99

Species mongolica Betsch, J-M, 1977:62, t.t.

Rusekianna mongolica from Mongolia
After Betsch J-M, 1977 Fig.2A

Species sibirica Bretfeld, 2002


Genus Zebulonia Betsch, J-M, 1970:69

Species massoudi Betsch, J-M, 1970:52


Genus Gisinianus Betsch, 1977

  Syn.: ... Gisianus(sic) Bushmakiu & al, 2000 ISS
Gisinianus flammeolus from Denmark
Abd.5 not fused with great abdomen
2019.02.24 © Schlütter, P.
Gisinianus flammeolus from Denmark
Abd.5+6 dorsal aspect
2019.02.26 © Schlütter, P.



Species flammeolus (Gisin, 1957) Betsch, 1977, t.t.

Sminthurinus flammeolus from Europe
Ocelli C+D smaller
2 setae inbetween ocelli H and G
Modified after Gisin H, 1963 Fig.3
             Syn.: Sminthurinus flameolus(sic) Kovacevic, 1967 ISS
             Syn.: Sminthurinus flameolus(sic) Kovacevic & Pagliarini, 1967 ISS
Gisinianus flammeolus from Denmark
2008.12.21 © Bildstrup, O.
Det.: Petersen, H.
Gisinianus flammeolusfrom Germany
2015.01.08 © Mattew, M.
Gisinianus flammeolus from Holland
Abd.5 not fused with great abdomen
2015.02.03 © van Duinen, J.
Gisinianus flammeolus from Holland
Abd.5 not fused with great abdomen
2015.02.03 © van Duinen, J.
Gisianus flammeolus from Belgium
Ant.1:2:3:4 ratio 1:2:2.5:5
2018.01.26 © Huskens, M.-L.
Gisianus flammeolus from Belgium
Circum-anal setae basally not expanded
a0 not furcate
2018.01.26 © Huskens, M.-L.
Gisianus flammeolus from Belgium
2018.01.26 © Huskens, M.-L.
Gisinianus flammeolus from Denmark
2019.02.24 © Schlütter, P.
Gisinianus flammeolus from Denmark
Abd.5 not fused with great abdomen
2019.02.24 © Schlütter, P.
Gisinianus flammeolus from Holland
Abd.5 not fused with great abdomen
2019.02.28 © van Duinen, J.
Gisinianus flammeolus from Holland
Abd.5 not fused with great abdomen
2019.02.28 © van Duinen, J.
Gisinianus flammeolus ♀ from Belgium
2020.01.23 © Huskens, M.-L.
Gisinianus flammeolus from Belgium
2021.01.11 © Huskens, M.-L.
Gisinianus flammeolus from Belgium
Footcomplex with more than 1 clavate tenent seta (*)
2021.01.11 © Huskens, M.-L.
Gisinianus flammeolus from Belgium
Abd.5 middorsally very short
2021.01.11 © Huskens, M.-L.
Gisinianus flammeolus from Holland
Preparing to moult
2022.01.11 © Kamsteeg, G.
Gisinianus flammeolus from Holland
Preparing to moult
Abd.5 part of small abdomen
2022.01.11 © Kamsteeg, G.
Gisinianus flammeolus from Holland
Preparing to moult
Caudal aspect
2022.01.11 © Kamsteeg, G.
Gisinianus flammeolus from Holland
Preparing to moult
Abd.5 not fused with large abdomen
2022.01.13 © Kamsteeg, G.
Gisinianus flammeolus from Holland
Ocelli C+D distinctly smaller
2022.01.13 © Kamsteeg, G.
Gisinianus flammeolus ♀ from Belgium
Grey colour variant
2022.02.02 © Huskens, M.-L.
Gisinianus flammeolus ♀ from Belgium
Grey colour variant
Ocellus C+D distinctly smaller
2022.02.02 © Huskens, M.-L.
Gisinianus flammeolus ♀ from Belgium
Grey colour variant
Abd.5 not fused with great abdomen
2022.02.02 © Huskens, M.-L.
Gisinianus flammeolus ♀ from Belgium
Grey colour variant
Sub-anal appendages apically broadly branched
2022.02.02 © Huskens, M.-L.




Genus Papirinus Yosii, 1954

  Syn.: Katiannidae Papirius(sic) Budha, PB et Shrestha, P, 2021:29 ISS

Species ankaratrensis Betsch, J-M, 1974:530

Species ieti Yosii, 1966

            Syn.: Papirius(sic) ieti Budha, PB et Shrestha, P, 2021:29

Species leleupi Delamare Deboutteville, C et Massoud, Z, 1963:227

            Syn.: Papirinus Leleupi(sic) Delamare Deboutteville, C et Massoud, Z, 1963:227 IOS

Species prodigiosus Yosii, 1954 tt

            Syn.: Papirinus prodigiosum(sic) Yosii, 1954 tt IOS
             Syn.: Papirinus prodigiosum(sic) Delamare Deboutteville, C et Massoud, Z, 1963:225 ISS
            Syn.: Papirius(sic) prodigiosus Budha, PB et Shrestha, P, 2021:29

Tribus Vesicephalini Richards, 1968:28






Genus Vesicephalus Richards WR in Delamare Deboutteville C & Massoud Z, 1964:75 key

Remark by Delamare Deboutteville, C. & Massoud, Z. 1964:75: A French translation of the original generic diagnosis of Vesicephalus Richards, W.R. (1956:79, unpublished thesis) was published.
  Syn.: Vesicephalini Vesicephalus Richards, 1956 u
Remark by Bretfeld, G. 1999:98: The new monophylum Katiannida Breteld, 1986 contains the Katiannidae Börner, 1913 sensu Betsch, 1980 and the genera incertae sedis sensu Betsch, 1980 and Bretfeld, 1986, including Vesicephalus Richards in Delamare Deboutteville & Massoud, 1964, based on two diagnostic apomorphies: 1. the presence of 2 additional outer setae Ka and Kp distally at the tibiotarsus; 2. the presence of spatulate setae distally at the tibiotarsus.
Remark by Fauna Europaea 2005: Family placement is uncertain. Vesicephalus is intermediate between Katiannidae (anal appendage in females) and Sminthuridae (dorsal chaetotaxy).
Vesicephalus sp. from the USA
Specimen caught in spider web
2007.12.02 © Ten Eyck, C.
Vesicephalus occidentalis from the USA
Fourth antennal segment not subdivided
2010.01.17 © Ten Eyck, C.
Vesicephalus occidentalis from the USA
Blunt dorsal macrosetae
2016.11.06 © Johnson, S.T.
Vesicephalus occidentalis from the USA
Clear swollen interocular vesicles
2016.11.06 © Johnson, S.T.
Vesicephalus occidentalis from the USA
Fourth antennal segment not subdivided
2016.11.06 © Johnson, S.T.



Species sp.1 Richards, 1968

Species sp.2 Snider & Janssens, 2008

Vesicephalus sp. from the USA
2008.03.04 © Fleming, K.R.
Vesicephalus sp. from the USA
2008.03.04 © Fleming, K.R.
Vesicephalus sp. from the USA
Reddish eyepatch
2017.02.03 © Fleming, K.R.
Vesicephalus sp. ♂ subadult from the USA
2022.02.20 © Charny, V.
Vesicephalus sp. ♂ subadult from the USA
2022.02.20 © Charny, V.
Vesicephalus sp. ♂ subadult from the USA
2022.02.20 © Charny, V.
Vesicephalus sp. ♂ subadult from the USA
Undeveloped male genital papilla (*)
2022.02.20 © Charny, V.
Vesicephalus sp. ♂ subadult from the USA
Caudal aspect
2022.02.20 © Charny, V.
Vesicephalus sp. from the USA
2022.03.25 © Charny, V.
Vesicephalus sp. from the USA
2022.03.25 © Charny, V.
Vesicephalus sp. from the USA
2022.03.25 © Charny, V.

Species sp.3 Janssens F & Cheng H-J, 2022

Vesicephalus sp. from Taiwan
2022.05.06 © Cheng, H.-J.
Vesicephalus sp. from Taiwan
2022.05.06 © Cheng, H.-J.

Species bellingeri Bretfeld, 2002

Species crossleyi Snider, RJ, 1985:567

Vesicephalus crossleyi from the USA
Female, type I
After Snider R, 1985 Fig.1
Vesicephalus crossleyi from the USA
Female, type I
After Snider R, 1985 Fig.2
Vesicephalus crossleyi from the USA
Female, type II
After Snider R, 1985 Fig.3
Vesicephalus crossleyi from the USA
Female, type II
After Snider R, 1985 Fig.4
Vesicephalus crossleyi from the USA
Male
After Snider R, 1985 Fig.5
Vesicephalus crossleyi from the USA
Male
After Snider R, 1985 Fig.6
Vesicephalus crossleyi ♀ type II from the USA
2017.12.23 © Abbott, J.C.
Vesicephalus crossleyi ♂ from the USA
2018.01.13 © Abbott, J.C.
Vesicephalus crossleyi ♀ from the USA
2018.06.04 © Caterino, M.
Vesicephalus crossleyi ♂ from the USA
2018.08.15 © Caterino, M.

Species europaeus Ardanaz & Pozo, 1985

             Syn.: Vesicephalus europeus(sic) Jordana, R, Baquero, E et Montuenga, LM, 2001:289 ISS


Vesicephalus europaeus
2002 © Baquero, E. & Jordana, R.
Vesicephalus europaeus from Portugal
2011.02.01 © Almeida, J.
Vesicephalus europaeus from Portugal
2011.02.02 © Almeida, J.
Vesicephalus europaeus male from Portugal
2012.02.02 © Almeida, J.

Species longisetis (Guthrie, JE, 1903:51) Delamare Deboutteville, C & Massoud, Z, 1964:73, t.t.

Species occidentalis (Mills, HB, 1935:138) Delamare Deboutteville, C & Massoud, Z, 1964:75

Vesicephalus occidentalis from the USA
Female
After Snider R, 1985 Fig.48
Vesicephalus occidentalis from the USA
Female
After Snider R, 1985 Fig.47
Vesicephalus occidentalis from the USA
2008.11.22 © Hall, K.
Vesicephalus occidentalis from the USA
2008.11.22 © Hall, K.
Vesicephalus occidentalis from the USA
2010.01.10 © Ten Eyck, C.
Vesicephalus occidentalis from the USA
2010.01.10 © Ten Eyck, C.
Vesicephalus occidentalis from the USA
2010.01.10 © Ten Eyck, C.
Vesicephalus occidentalis from the USA
Pale colour form
2016.10.28 © Dobbs, Br.
Vesicephalus occidentalis from the USA
2016.11.06 © Johnson, S.T.
Vesicephalus occidentalis from the USA
2016.11.06 © Johnson, S.T.
Vesicephalus occidentalis from the USA
2016.12.03 © Johnson, S.T.
Vesicephalus occidentalis from the USA
Foot with many tenent setae
2016.12.03 © Johnson, S.T.
Vesicephalus occidentalis from the USA
Head with long curved blunt macrosetae
2016.12.03 © Johnson, S.T.
Vesicephalus occidentalis from the USA
Face
2016.12.03 © Johnson, S.T.
Vesicephalus occidentalis juv. from the USA
2017.11.10 © Dobbs, Br.
Vesicephalus occidentalis from the USA
2017.11.10 © Dobbs, Br.
Vesicephalus occidentalis from the USA
2017.11.10 © Dobbs, Br.
Vesicephalus occidentalis from the USA
2017.11.29 © Dobbs, Br.
Vesicephalus occidentalis from Canada
2017.12.05 © Barbin, T.
Vesicephalus occidentalis from Canada
2017.12.05 © Barbin, T.
Vesicephalus occidentalis from Canada
2017.12.05 © Barbin, T.
Vesicephalus occidentalis from Canada
Strong frontal macrosetae
2017.12.05 © Barbin, T.
Vesicephalus occidentalis from Canada
2018.01.12 © Barbin, T.
Vesicephalus occidentalis from the USA
2018.01.28 © Johnson, S.T.
Vesicephalus occidentalis from the USA
2018.12.28 © Pearson, R.R.
Vesicephalus occidentalis from the USA
2018.12.28 © Pearson, R.R.
Vesicephalus occidentalis from the USA
2020.11.07 © Gerth, Bi.
Vesicephalus occidentalis from the USA
2021.01.11 © Pearson, R.R.
Vesicephalus occidentalis juv. from the USA
2021.01.11 © Pearson, R.R.
Vesicephalus occidentalis from the USA
2021.03.06 © Parker, H.S.
Vesicephalus occidentalis from the USA
2022.01.03 © Parker, H.S.
Vesicephalus occidentalis from the USA
Strong frontal macrosetae
2022.01.03 © Parker, H.S.
Vesicephalus occidentalis from the USA
With distinctly large eyebrows
2022.12.03 © G, S.
Vesicephalus occidentalis from the USA
2022.12.03 © OakandSaga.
Vesicephalus occidentalis from the USA
2024.01.21 © OakandSaga.

References