http://www.collembola.org/taxa/tomoceri.htm - Last updated on 2014.03.21 by Frans Janssens
Checklist of the Collembola: Tomoceridae

Familia Tomoceridae Schäffer, 1896 key
Genus Number
of species
Antennacyrtus1
Aphaenomurus1
Entomocerus1
Lasofinius2
Lepidophorella10
Lethemurus3
Monodontocerus6
Neophorella1
Novacerus3
Plutomurus29
Pogonognathellus17
Pseudolepidophorella1
Tomocerina12
Tomocerus78
Tomolonus1
Tritomurus3
Tomoceridae169
Genera with number of species
  Syn.: ... Tomocerinæ(sic) Schäffer, 1896 ISS
  Syn.: ... Tomocerinæ(sic) Börner, 1906 ISS
  Syn.: Collembola Tomocerinæ(sic) Folsom, JW, 1913:451 ISS
  Syn.: Collembola Tomoceridæ(sic) Dethier, M, 1998:57 ISS
Pogonognathellus sp. from the UK
Body covered with iridescent scales
2007.12.19 © Campbell, A.
 
Tomocerus minor from the UK
3rd abdominal segment superequal to 4th
2008.09.24 © Robertson, A.


Subfamilia Tomocerinae Schäffer, 1896
  Syn.: ... Tomocerinæ(sic) Schäffer, 1896 ISS
Tomocerus sp. from France
Long 3rd antennal segment
2007.02.23 © Ollivier, E.
Tomocerinae juv. from Belgium
2008.06.09 © Vuijlsteke, M.
 
Tomocerinae juv. from the UK
Second instar with short 3rd antennomere
2008.09.28 © Barton, T.
Tomocerinae juv. from the USA
Long 3rd antennal segment
2009.03.14 © Justis, S.
Tomocerus sp. from Czechia
Descaled specimen revealing body colour
2010.11.09 © Fiala, M.
Tomocerinae juv. from the UK
First instar with short 3rd antennomere
2011.12.01 © Nurcombe, E.
Tomocerus minor from the USA
3rd antennomere with very fine annulation
2012.02.18 © Carr, J.F.


Tribus Tomocerini Salmon JT, 1941:346

Genus Tomocerus Nicolet, H, 1842:67
  Syn.: Podurelles Temnourus Nicolet, H in Desor, E, 1841:384
  Syn.: Podures Macrotoma Bourlet, C, 1839:387
  Syn.: ... Palpiger Olfers, 1907
  Syn.: Isotomidae Architomocerura Denis, 1931
  Syn.: ... Eutomocerus Yosii, 1955
  Syn.: ... Monodontocerus Yosii, 1955
Remark by ICZN. 2007:821: Placed on the Official List of Names in Zoology. Opinion 239.

Tomocerus minor from the UK
Long 3rd antennal segment subcylindrical
2008.09.24 © Robertson, A.
Tomocerus minor from the UK
1st and 2nd antennal segments scaled
2008.12.22 © Valentine., B
Tomocerus minor from the UK
Long 3rd antennal segment subcylindrical
2009.01.26 © Robertson, A.
Tomocerus minor from Germany
Colour of 3rd antennal segment distinctly different from other segments
2009.01.26 © Peters, J.
Tomocerus sp. from the USA
Eyecluster with 6 visible ocelli
2009.02.26 © Cowen, R.
 
Tomocerus sp. juv. from the UK
4th ant. segment > 1st + 2nd ant. segment
2009.04.20 © Robertson, A.
 
Tomocerus sp. from Canada
Dental spines array
2011.11.16 © Hochhalter, R.
Tomocerus sp. juv. from the UK
4th ant. segment > 1st + 2nd ant. segment
2012.09.02 © Murray, A.
Tomocerus sp. from the UK
Composed eye with 6 ocelli
2012.09.19 © Murray, A.
Tomocerus sp. juv. from the UK
Annulated 4th antennal segment
2012.09.26 © Murray, A.

Species sp.1 Agrell, 1939
Species sp.2 Agrell, 1939
Species sp.3 Hopkin S, 1998
Tomocerus sp. from the UK
Mucro with one intermediate tooth
1998 © Hopkin, S.

Species asahinai Yosii, 1954
Species asiaticus Martynova, 1969
Species asoka Yosii & Ashraf, 1965
Species baibungensis Sun, Y, Liang, A-P et Huang, F-S, 2006:561
Species baicalensis Martynova, 1969
Species baudoti Denis, 1932
Species bimaculatus Sun, Liang & Huang, 2006
Species caputiviolaceus Lee, 1975
Species catalanus Denis, 1924
Species changbaishanensis Wang, 1999
Species cheni Ma, Y-T et Christiansen, KA, 1998:47
Species christianseni Stomp, N, 1969:695
Species conagensis Sun, Y, Liang, A-P et Huang, F-S, 2006:516
Species cucullatus Olfers, 1907, t.t.
Species cuspidatus Börner, C, 1909:114
Species denticulus Lee, 1975
Species deongyuensis Lee, 1975
            Syn.: Tomocerus (Tomocerus) deogyuensis(sic) Lee, B-H et Park, K-H, 1992:106 ISS
            Syn.: Tomocerus (Tomocerus) deogyuensis(sic) Ma, Y-T, Chen, J-X et Christiansen, K, 2004:47 ISS
Species emeicus Liu, Hou & Li, 1999
Species folsomi Denis, 1929
Species fopingensis Sun & Liang, 2008
Species hexipunctatus Sun, Liang & Huang, 2007
Species huoensis Sun, Y, Liang, A-P et Huang, F-S, 2006:804
Species ishibashii Yosii, 1954
            Syn.: Tomocerus ishabashii(sic) Hopkin, 1997 ISS
Species jesonicus Yosii, 1967
Species jezoensis Matsumura & Ishida in Matsumura, 1931:200
Species jilinensis Ma, 2011
Species jinyunensis Liu, Y-Q, 2006:300
            Syn.: Tomocerus (Tomocerus) jinyun(sic) Liu, Y-Q, 2006:301 ISS
Species jordanai Sun, Liang & Huang, 2006
Species kinoshitai Yosii, 1954
  Subspecies dentiferus Yosii, R, 1956:90
Species lamelliger (Börner, 1903) Gisin, 1960:254
Species laxalamellus Lee, 1975
            Syn.: Tomocerus laxalamella(sic) Lee, 1975 IOS
            Syn.: Tomocerus (Monolamellus) laxalamella(sic) Lee, 1983 ISS
            Syn.: Tomocerus (Monolamellus) laxalamella(sic) Lee, B-H et Park, K-H, 1992:107 ISS
Species lividus Tullberg, T, 1877:32
Species maculatus Sun, Liang & Huang, 2007
Species magadanicus Martynova, 1979
Species magnus Sun & Liang, 2008
Species maximus Liu, Hou & Li, 1999
Species minor (Lubbock, 1862) Schäffer, 1900, t.t., B
            Syn.: Tomocerus plumbeus Lubbock, J, 1873:138
Tomocerus plumbeus from the UK
After Lubbock, J., 1873 Pl.19.

            Syn.: Podura gigas Fabricius O, 1783
            Syn.: Podura plumbea von Linné, C, 1767:1013
            Syn.: Podura plumbea Fabricius, 1775
            Syn.: Podura plumbea Müller OF, 1776
            Syn.: Podura plumbea Schrank, 1781
            Syn.: Podura plumbæa(sic) de Fourcroy, AF, 1785:524 ISS
            Syn.: Podura plumbea Latreille, 1796
            Syn.: Podura plumbea Goring, CR, 1831:246
            Syn.: Choreutes plumbeus Burmeister, 1835
            Syn.: Podura plumbea Boisduval & Lacordaire, 1835
            Syn.: Podura plumbea Bourlet, C, 1839:384
Macrotoma plumbea from the UK
After McIntire SJ, 1867 Fig.51
After McIntire SJ, 1868


            Syn.: Choreutes plumbea(sic) Lubbock, J, 1870:284 ISS
Tomocerus plumbeus from the UK
After Beck J in Lubbock J, 1873 Pl.68

            Syn.: Tomocerus norvegicus Lie-Pettersen, 1897
            Syn.: Macrotoma tridentifera Tullberg, T, 1872:37
Tomocerus minor from Belgium
Tridentate dental spines
Legit Leruth, R., 1933.07.16
2008.05.28 © Janssens, F.
Tomocerus minor from Belgium
Unscaled 3rd antennal segment
Legit Leruth, R, 1933.10.03
2008.05.28 © Janssens, F.
Tomocerus minor from Belgium
2004 © Vuijlsteke, M.
Tomocerus minor from the UK
2005 © Hopkin, S.P.
Tomocerus minor from the UK
2006 © Cornwall, N.J.
Tomocerus minor from Belgium
2006 © Vuijlsteke, M.
Tomocerus minor from Germany
2006 © Peters, J.
Tomocerus minor from the UK
2007 © Barton, T.
Tomocerus minor from Germany
2007.02.15 © Peters, J.
Tomocerus minor from the USA
2007.03.03 © Murray, T.
Tomocerus minor from the USA
2007.03.03 © Murray, T.
Tomocerus minor from the USA
2007.03.11 © Murray, T.
Tomocerus minor from the USA
2007.05.04 © Moorehead, C.
Tomocerus minor from Belgium
2007.07.07 © Vuijlsteke, M.
Tomocerus minor from the UK
2007.07.22 © Valentine., B
Tomocerus minor from the USA
2007.09.21 © Moorehead, C.
Tomocerus minor from the UK
2007.12.14 © Botting, J.
Tomocerus minor from the Netherlands
2008.01.24 © Baas, A.H.
Tomocerus minor from the UK
2008.02.03 © Birch, P.
Tomocerus minor from the Netherlands
2008.02.03 © Baas, A.H.
Tomocerus minor from the UK
2008.02.27 © Spears, G.
Tomocerus minor from Belgium
2008.03.15 © Oger, P.
Tomocerus minor from the UK
2008.03.17 © Robertson, A.
Tomocerus minor from Belgium
2008.03.22 © Vuijlsteke, M.
Tomocerus minor from Belgium
2008.03.22 © Vuijlsteke, M.
Tomocerus minor from Belgium
2008.03.24 © Vuijlsteke, M.
Tomocerus minor from the USA
2008.05.29 © Nanz, S.
Tomocerus minor from the USA
2008.09.13 © Robertson, A.
Tomocerus minor from the UK
2008.09.24 © Robertson, A.
Tomocerus minor juvenile from the UK
2008.09.24 © Robertson, A.
Tomocerus minor from the UK
2008.10.21 © Spears, G.
Tomocerus minor from the UK
2009.01.14 © Valentine., B
Tomocerus minor from Germany
2009.02.18 © Peters, J.
Tomocerus minor from the UK
2009.02.27 © Knight., B
Tomocerus minor from Belgium
2009.03.15 © Janssens, F.
Tomocerus minor from the UK
2009.04.29 © Robertson, A.
Tomocerus-minor from Belgium
2009.09.13 © Wallays, H.
Tomocerus minor, juv. from the UK
2009.10.07 © Robertson, A.
Tomocerus minor from the UK
Body scales bluish iridisation
2010.01.02 © Robertson, A.
Tomocerus minor from the UK
Macroseta with blunt apex
2010.01.02 © Robertson, A.
Tomocerus minor from the UK
Microsetae on 3rd antennal segment
2010.01.02 © Robertson, A.
Tomocerus minor from Canada
Abdominal bothriotricha
2011.09.18 © Hochhalter, R.
Tomocerus minor from Belgium
Descaled specimen with dark thorax
2011.12.28 © Menten, J.
Tomocerus minor from Belgium
Mucro with teeth and setae
2011.12.28 © Menten, J.
Tomocerus minor from Belgium
Footcomplex: unguiculus, unguis and tenant hair
2011.12.28 © Menten, J.
Tomocerus minor from Germany
Descaled specimen, revealing dark pigmented thorax
2012.03.15 © Rose, A.
Tomocerus minor from Belgium
2013.04.06 © Declercq., B

Species mitrai Prabhoo & Muraleedharan, 1980
Species montanus (Oudemans, 1890:74) Yayuk, 1989:9
Species monticolus Huang & Yin, 1981
Species multisetus Sun, Liang & Huang, 2007
Species nepalicus Yosii, 1971
Species nigrofasciatus Sun, Y, Liang, A-P et Huang, F-S, 2006:559
Species nigromaculatus Sun, Y, Liang, A-P et Huang, F-S, 2006:513
Tomocerus (Tomocerus) nigromaculatus from Tibet
After Sun & al, 2006 Fig.1A

Species nigrus Sun, Y, Liang, A-P et Huang, F-S, 2006:803
Species nodentalis Nguyen, 1995
Species noricus Latzel, 1918
Species obscurus Huang & Yin, 1981
            Syn.: Tomocerus (Tomocerus) obsculus(sic) Chen, J-X et Ma, Y-T, 1997:157 ISS
            Syn.: Tomocerus (Tomocerus) obsculus(sic) Ma, Y-T et Christiansen, KA, 1998:47 ISS
            Syn.: Tomocerus (Tomocerus) obsculus(sic) Chen, J-X et Christiansen, KA, 1998:51 ISS
Species ocreatus Denis, 1948
            Syn.: Tomocerus kawamurai Yosii, 1954
            Syn.: Tomocerus kawamurai f. depicta Yosii, 1955
            Syn.: Tomocerus minor Uchida, 1954
            Syn.: Tomocerus ochreatus(sic) f. kawamurai Yosii, R, 1956:90 ISS
Species orcinus Absolon, K, 1901:88
Species orientalis Martynova, 1979
Species parvus Huang & Yin, 1981
Species petalospinus Salmon, 1969
Species problematicus Cassagnau, P, 1964:741
Species punctatus Yosii, 1967
Species purpuratus Sun, Liang & Huang, 2006
Species sachalinensis Martynova, 1979
Species serratospinus Salmon, 1969
Species setoserratus Salmon, JT, 1941:347
Species shuense Liu, 2003
Species sibiricus Reuter, 1891
Species similis Chen, J-X et Ma, Y-T, 1997:157
Species spinistriatus Lee, 1975
Species spinulus Chen, J-X et Christiansen, KA, 1998:51
Species steinbocki Yosii, 1971
Species teldanicus Gruia, 1995
Species terrestralis (Stach, 1922) Cassagnau, P, 1958:121
            Syn.: Tomocerus terrestrialis(sic) Christiansen, K, 1964:656 ISS
Species varius Folsom, JW, 1899:267
Species vasconius Bonet, 1928
Species violaceus Yosii, R, 1956:91
Species viridescens Wankel, 1860
            Syn.: Macrotoma viridiscens(sic) Absolon, K, 1899:317 ISS
Species viridis Yosii, 1967
Species vulgaris (Tullberg, 1871) Brook, 1883, B
            Syn.: Macrotoma ferruginosa Bourlet, C, 1839:390
            Syn.: Macrotoma nigra Bourlet, C, 1839:390
            Syn.: Tomocerus celer Nicolet, H, 1842:69
            Syn.: Tomocerus plumbeus Packard, 1873, partim
            Syn.: Tomocerus niger Guthrie, JE, 1903:80
            Syn.: Tomocerus minutus Hammer, 1953
            Syn.: Tomocerus botanicus Cassagnau, 1962
            Syn.: Tomocerus vulgaris var. kurilensis Uchida, H, 1965:207
Tomocerus vulgaris from Germany
2005 © Menzel, E.
Tomocerus vulgaris from Germany
2005 © Kohl, F.
Tomocerus vulgaris from Germany
2005 © Peters, J.
Tomocerus vulgaris from the Netherlands
2007 © Hautus, M.A.
Tomocerus vulgaris from Belgium
2007.07.12 © Janssens, F.
Tomocerus vulgaris from Germany
2007.04.07 © Kohl, F.
Tomocerus vulgaris from the Netherlands
2007.09.18 © Arp, X.
Tomocerus vulgaris from Germany
2008.01.25 © Peters, J.
Tomocerus vulgaris from the UK
2008.01.27 © Barton, T.
Tomocerus vulgaris from Belgium
2008.08.28 © Janssens, F.
Tomocerus vulgaris from the Netherlands
2008.10.06 © Baas, A.H.
Tomocerus vulgaris from the Netherlands
2008.10.07 © Baas, A.H.
Tomocerus vulgaris from the Netherlands
2008.10.08 © Baas, A.H.
Tomocerus vulgaris from the Netherlands
2008.10.10 © Baas, A.H.
Tomocerus vulgaris from France
2008.11.03 © Ramel, A.
Tomocerus vulgaris from Germany
2008.12.07 © Peters, J.
Tomocerus vulgaris from Belgium
2009.02.21 © Vuijlsteke, M.
Tomocerus vulgaris from the UK
2009.03.04 © Ransom, T.
Tomocerus vulgaris from Belgium
2009.03.07 © Janssens, F.
Tomocerus vulgaris from France
2009.04.19 © Lefebvre, V.
Det. Ponge, J.-F.
Tomocerus vulgaris from Belgium
2009.04.19 © Doornaert, A.
Tomocerus vulgaris from Germany
2009.11.07 © Buonvolere, D.
Tomocerus vulgaris from Belgium
Dental spines
2010.01.13 © De Bruyn, L.
Tomocerus vulgaris from Slovenia
2010.04.15 © Miran, C.
Tomocerus vulgaris from Holland
2010.04.30 © van Duinen, J.
Tomocerus vulgaris from Holland
Typical dorsal banding
2011.08.17 © van Duinen, J.
Tomocerus vulgaris from Germany
2011.10.26 © Lenke, M.
Tomocerus vulgaris from Germany
Metathoracic botriotrichum: mechanoreceptive sensilla in cup-like depression of skin
2011.10.26 © Lenke, M.
Tomocerus vulgaris from the UK
2012.02.03 © Dargie, S.
Tomocerus vulgaris from Belgium
2012.02.26 © Wallays, H.
Tomocerus vulgaris from the UK
2012.05.23 © Nicholls, D.
Tomocerus vulgaris from the UK
Dark pigmented clypeus
2012.12.09 © Shaw, P.
Tomocerus vulgaris from the UK
Aberrant mucro with one (optional) intermediate tooth
2012.12.09 © Shaw, P.
Tomocerus vulgaris from the UK
Dental spines
2012.12.09 © Shaw, P.
Tomocerus vulgaris from the UK
Foot complex
2012.12.09 © Shaw, P.
Tomocerus vulgaris from Belgium
The "golden Tomocerus"
2013.08.11 © Wallays, H.

Species wushanensis Sun, Liang & Huang, 2007
Species zayuensis Huang & Yin, 1981
            Syn.: Tomocerus (Tomocerus) zayensis(sic) Ma, Y-T et Christiansen, KA, 1998:47 ISS
            Syn.: Tomocerus (Tomocerus) zayensis(sic) Chen, J-X et Christiansen, KA, 1998:51 ISS

Genus Tritomurus Frauenfeld, 1854 key
Species falcifer Cassagnau, P, 1958:121
Tritomurus falcifer
After Thibaud J-M & D'Haese C, 2010 Fig.19

Species scutellatus Frauenfeld, 1854:15, t.t.
Tritomurus scutellatus from Croatia
2010.11.19 © Lukic, M.

Species veles Lukic, M, Houssin, C et Deharveng, L, 2010:3
Tritomurus veles from Croatia
2010.11.23 © Houssin, C.


Genus Pogonognathellus Paclt, 1944
Pogonognathellus sp. from Japan
2005.09.07 © Keresztes, G.
Pogonognathellus sp. from France
Furca with short dental spines array
2007.04.10 © Lebeaux, P.
Pogonognathellus flavescens
from the Netherlands
2008.11.03 © Baas, A.H.
Pogonognathellus sp. from Taiwan
Tapering antenna
2010.06.23 © Chien, H.-C.
Pogonognathellus sp. from Taiwan
2010.06.30 © Chien, H.-C.
Pogonognathellus longicornis from Sweden
Scaled 3rd antennal segment
2011.11.25 © Hallmén, J.
Pogonognathellus sp. from the UK
Eyepatch with 6 visible ocelli
2012.01.23 © Kent, J.J.

Species sp.1 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475
Species beckeri (Börner, C, 1909:115)
Pogonognathellus beckeri from Japan
2008 © Tsukiji, T.

Species belmontorum Park, K-H, Bernard, EC et Moulton, JK, 2011:2
Pogonognathellus belmontorum from the USA
After Park K-H & al., 2011 Fig.1A
            Syn.: Pogonognathellus flavescens Nearctic records partim
Species bidentatus (Folsom, JW, 1913:463) Massoud, Z & Ellis, WN, 1974:292
            Syn.: Tomocerus bidentatus ssp. Jeanneli(sic) Bonet, 1934
            Syn.: Tritomurus henroti Delamare Deboutteville, 1949
            Syn.: Tritomurus Henroti(sic) Cassagnau, P, 1958:121 ISS
            Syn.: Tomocerus (Pogonognathellus) dibentatus(sic) Christiansen, K, 1964:668 ISS
Pogonognathellus bidentatus from the USA
After Felderhoff KL et al., 2010 Fig.2A
Antennae typically curved backwards in capricorn fashion
Pogonognathellus bidentatus from the USA
After Felderhoff KL et al., 2010 Fig.5A
Descaled; cuticular pattern

Species borealis Yosii, 1967
Species brevifulvus Park, K-H, Bernard, EC et Moulton, JK, 2011:6
Pogonognathellus brevifulvus from the USA
After Park K-H & al., 2011 Fig.3A
Pogonognathellus brevifulvus from the USA
2013.07.04 © Liberta, A.

Species celsus (Christiansen, K, 1964:670) Massoud, Z & Ellis, WN, 1974:293
Species danieli Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:488
Pogonognathellus danieli from the USA
After Felderhoff KL et al., 2010 Fig.5B
Descaled; cuticular pattern

Species dubius (Christiansen, K, 1964:671) Felderhoff, KL, Bernard, EC & Moulton, JK, 2010:488
            Syn.: Pogonognathellus grayspeciescomplex Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:488
Remark by Felderhoff, K.L. et al. 2010:488: The taxa included in this group are united by cuticular pigmentation, scale color, and alignment of the basal and lateral mucronal teeth, but the sequenced specimens do not seem to form a monophyletic group. Instead, they clustered in four separate groups scattered among members of the "pale complex." Additional collections, including specimens from the P. dubius type locality, will be necessary to begin to satisfactorily resolve these taxa.

            Syn.: Pogonognathellus sp.4 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475
            Syn.: Pogonognathellus sp.9 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475
Pogonognathellus dubius s.l. from the USA
2005 © DiTerlizzi, T.
Pogonognathellus dubius s.l. from the USA
2005 © Murray, T.
Pogonognathellus dubius s.l. from the USA
2006 © Cresswell, S.
Pogonognathellus dubius s.l. from the USA
2007 © Murray, T.
Pogonognathellus dubius s.l. from Canada
2007.12.17 © McCann, S.
Pogonognathellus dubius s.l. from the USA
2008.09.09 © Bradford, A.
Pogonognathellus dubius s.l. from the USA
2008.10.08 © Bradford, A.
Pogonognathellus dubius s.l. from the USA
2008.12.06 © Houston, S.
Pogonognathellus dubius s.l. from the USA
2009.06.18 © Bradford, A.
Pogonognathellus dubius s.l. from the USA
2009.10.25 © Bergen, L.
Pogonognathellus dubius s.l. from the USA
2010.06.26 © Ueda, K.

Species elongatus (Maynard, EA, 1951:126) Felderhoff, KL, Bernard, EC & Moulton, JK, 2010:482
Pogonognathellus elongatus from the USA
2006 © Cresswell, S.
Pogonognathellus elongatus from the USA
Body scales removed
2008.08.12 © Trently, D.
Pogonognathellus elongatus from the USA
After Felderhoff KL et al., 2010 Fig.3A
 
Pogonognathellus elongatus from the USA
After Felderhoff KL et al., 2010 Fig.3B
Posterior dorsal macrochaetotaxy
Pogonognathellus elongatus from the USA
After Felderhoff KL et al., 2010 Fig.3B
Specimen with projected mesotergum
Pogonognathellus elongatus from the USA
After Felderhoff KL et al., 2010 Fig.3D
Specimen with projected mesotergum
Pogonognathellus elongatus from the USA
After Felderhoff KL et al., 2010 Fig.5C
Descaled; cuticular pattern
Pogonognathellus elongatus from the USA
2012.07.01 © Cresswell, S.

Species flavescens (Tullberg, 1871) Stach, 1929, B
            Syn.: Podura plumbea Linnæus, C, 1758:609
            Syn.: Podura plumbea Templeton, R in Templeton, R et Westwood, JO, 1836:93
Podura plumbea from Ireland
After Templeton, R, 1836 Pl.XI Fig.4,4.1

            Syn.: ... plumbea Ågren, 1903, nec Templeton, 1835
            Syn.: Podura violacea Geoffroy
            Syn.: Podura longicornis Müller OF, 1776
            Syn.: Podura gigas Fabricius O, 1783
            Syn.: Podura communis Fourcroy, 1785
            Syn.: Macrotoma ferruginosa Bourlet, C, 1839:390
            Syn.: Macrotoma nigra Bourlet, C, 1839:390
            Syn.: Tomocerus celer Nicolet, H, 1842:69
            Syn.: Macrotoma lepida Gervais in Walckenaër, 1844
            Syn.: Tomocerus lepida(sic) Folsom, JW, 1902:97 ISS
            Syn.: Pogonognathus beckeri Börner, 1909
            Syn.: Architomocerura crassicauda Denis, 1931,tt
            Syn.: Paratomocerus pierantonii Tarsia in Curia, 1938,tt
            Syn.: Tomocerus flavescens ssp. arcticus Yosii, 1940, nec Schött

Pogonognathellus flavescens from Belgium
2004 © Vuijlsteke, M.

            Syn.: Tomocerus Flavencens(sic) Palacios-Vargas, JG in Sánchez-Ramos, G, Reyes-Castillo, P et Dirzo, R, 2005:365 ISS
            Syn.: Tomocerus flavencens(sic) Palacios-Vargas, JG in Sánchez-Ramos, G, Reyes-Castillo, P et Dirzo, R, 2005:366 ISS
Pogonognathellus flavescens from the UK
2006.02.17 © Valentine., B
Pogonognathellus flavescens from Belgium
2006.03.19 © Vuijlsteke, M.
Pogonognathellus flavescens from the UK
2007.02.13 © Jones, D.
Pogonognathellus flavescens from the UK
2007.02.18 © Barton, T.
Pogonognathellus flavescens from Czechia
2007.06.23 © Krásenský, P.
Pogonognathellus flavescens from Czechia
2007.07.22 © Krásenský, P.
Pogonognathellus flavescens from the Netherlands
2007.11.05 © Baas, A.H.
Pogonognathellus flavescens from the UK
2007.11.14 © Campbell, A.
 
Pogonognathellus flavescens from the UK
2007.11.18 © Campbell, A.
 
Pogonognathellus flavescens from the UK
2007.11.25 © Warner, P.
Pogonognathellus flavescens from the Netherlands
2007.12.08 © Heemskerk, R.
Pogonognathellus flavescens from the Netherlands
2008.02.10 © Baas, A.H.
Pogonognathellus flavescens from the UK
2008.02.10 © Campbell, A.
Pogonognathellus flavescens from France
2008.02.11 © Lavoué., B
Pogonognathellus flavescens from the UK
Juvenile, descaled
2008.03.31 © Williams., B
Pogonognathellus flavescens from the UK
2008.05.01 © Farrell, S.
Pogonognathellus flavescens from Belgium
Scaled 3rd antennal segment
Legit Leruth, R, 1933.09.30
2008.05.28 © Janssens, F.
Pogonognathellus flavescens juv.
from the Netherlands
2008.10.15 © Baas, A.H.
 
Pogonognathellus flavescens juv.
from the Netherlands
2008.10.15 © Baas, A.H.
Pogonognathellus flavescens
from the Netherlands
2008.11.03 © Baas, A.H.
Pogonognathellus flavescens from France
2008.11.08 © Jacquemin, G.
 
Pogonognathellus flavescens from the UK
2009.01.31 © Ransom, T.
Pogonognathellus flavescens from Belgium
2009.02.21 © Vuijlsteke, M.
Pogonognathellus flavescens (bar=2µm)
Hexagonal epicuticular ultrastructure
After Nickerl J & al, 2012 Fig.2A'
Pogonognathellus flavescens from the UK
2012.09.12 © Jones, A.
Pogonognathellus flavescens from Belgium
2013.04.06 © Declercq., B
Pogonognathellus flavescens from Holland
Decsaled specimen revealing intrinsic yellow body colour
2013.10.26 © Baas, A.H.
Pogonognathellus flavescens from Holland
Moulting, final phase: redrawing furca from old cuticula
2013.10.26 © Baas, A.H.
Pogonognathellus flavescens from Holland
Just moulted, dorsal aspect
2013.10.26 © Baas, A.H.
Pogonognathellus flavescens from Holland
Just moulted, lateral aspect: thorax bulged
2013.10.26 © Baas, A.H.
Pogonognathellus flavescens from Holland
Moulting complete
2013.10.26 © Baas, A.H.
Pogonognathellus flavescens from the UK
Descaled specimen with partly recovered damaged left antenna
2013.12.24 © Evans, M.

Species longicornis (Müller OF, 1776) Paclt, 1944, t.t., B
            Syn.: Podura plumbea Linnæus, 1761
            Syn.: Podura plumbea von Linné, C, 1767:1013,tt
            Syn.: Podura plumbea Villers, 1789
            Syn.: Podura plumbea Templeton, 1835
            Syn.: Tomocerus plombeus(sic) Nicolet, H, 1847:380 ISS
Tomocerus plombeus(sic) from Europe
After Nicolet, H, 1847 Pl.6 Fig.12.

            Syn.: Tomocerus plombeus(sic) Frauenfeld, 1854 ISS
            Syn.: Podura bicolor Say, 1821
            Syn.: Podura iricolor Say, 1821
            Syn.: Macrotoma ferruginosa Bourlet, C, 1839:390
            Syn.: Macrotoma spiricornis Bourlet, 1843
            Syn.: Macrotoma Spiricornis(sic) Bourlet, C, 1845:67 ISS
            Syn.: Macrotoma rufescens Tullberg, 1871
            Syn.: Pogonognathus plumbeus Boldori, 1938
            Syn.: Architomocerura litsteriana Bagnall, 1940
Tomocerus longicornis from the UK
After Beck J in Lubbock J, 1873 Pl.67
Tomocerus longicornis from the UK
After Lubbock J, 1873 Pl.17

            Syn.: Tomocerus longicornes(sic) Noble-Nesbitt, J, 1963:253 ISS
Pogonognathellus longicornis from the UK
2002 © Hopkin, S.P.
Pogonognathellus longicornis
2003 © Pettersson., B
Pogonognathellus longicornis from the UK
Large leaf-like basal dental scales
2004 © Hopkin, S.P.
Pogonognathellus longicornis from Sweden
2005 © Hall, K.
Pogonognathellus longicornis from the UK
2006 © Cornwall, N.J.
Pogonognathellus longicornis from the UK
2006 © Cornwall, N.J.
Pogonognathellus longicornis from Russia
2006 © Macroclub.ru.
Pogonognathellus longicornis from Germany
2006.09.30 © Kohl, F.
Pogonognathellus longicornis from Germany
2007 © Peters, J.
Pogonognathellus longicornis from Belgium
2007.02.07 © Lindsley, J.
Pogonognathellus longicornis from France
2007.06.09 © Lebeaux, P.
Pogonognathellus longicornis from Belgium
2007.09.08 © Oger, P.
Pogonognathellus longicornis from the UK
2007.10.13 © Brian.
Pogonognathellus longicornis from the UK
2007.10.13 © Brian.
Pogonognathellus longicornis from the UK
2007.11.10 © Campbell, A.
Pogonognathellus longicornis from Belgium
2007.11.24 © Carlet, G.
Pogonognathellus longicornis from Sweden
2007.12.04 © Hallmén, J.
Pogonognathellus longicornis from the Netherlands
2007.12.08 © Heemskerk, R.
Pogonognathellus longicornis from the Netherlands
2007.12.08 © Heemskerk, R.
Pogonognathellus longicornis from the UK
2008.04.03 © Spears, G.
Pogonognathellus longicornis from the UK
2008.07.27 © Michaelson, J.
Pogonognathellus longicornis from the UK
2008.09.01 © Talbot, M.
Pogonognathellus longicornis from the UK
2009 © Chaplin, M.
Cleaning the antenna with maxillae
Pogonognathellus longicornis from the UK
2009 © Chaplin, M.
Everting vesicles of collophore
Pogonognathellus longicornis from the UK
2009.02.17 © Ransom, T.
Pogonognathellus longicornis from the UK
Unguis (U) with lateral teeth (LT)
Unguicular filament (uf) longer than unguis
2009.04.03 © Robertson, A.
Pogonognathellus longicornis from Sweden
5th and 6th abdominal segments
2011.11.25 © Hallmén, J.
Pogonognathellus longicornis from Sweden
2011.11.25 © Hallmén, J.
Pogonognathellus longicornis from the UK
Spirally curled antennae, ready to jump
2012.08.09 © Lindsay., B
Pogonognathellus longicornis from Germany
Eyepatch with 6 ocelli
2012.09.12 © Schmidt, R.
Pogonognathellus longicornis from the UK
Abdominal dorsal bothriotricha
2012.11.06 © Nurcombe, E.

Species magnibrunneus Park, K-H, Bernard, EC et Moulton, JK, 2011:9
Pogonognathellus magnibrunneus from the USA
After Park K-H & al., 2011 Fig.5A

Species mystax Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:488
Pogonognathellus n. sp. from the USA
2009.09.01 © Hodge, K.
Pogonognathellus mystax from the USA
After Felderhoff KL et al., 2010 Fig.2B
Pogonognathellus mystax from the USA
After Felderhoff KL et al., 2010 Fig.2C
Pogonognathellus mystax from the USA
After Felderhoff KL et al., 2010 Fig.2D
Patterned clypeus
Pogonognathellus mystax from the USA
After Felderhoff KL et al., 2010 Fig.5E
Descaled; cuticular pattern

Species nigritus (Maynard, EA, 1951:130) Felderhoff, KL, Bernard, EC & Moulton, JK, 2010:484
Pogonognathellus nigritus from the USA
2009.12.05 © Guzman, D.
Pogonognathellus nigritus from the USA
After Felderhoff KL et al., 2010 Fig.4A
Pogonognathellus nigritus from the USA
After Felderhoff KL et al., 2010 Fig.4C
Dorsal macrochaetotaxy
Pogonognathellus nigritus from the USA
After Felderhoff KL et al., 2010 Fig.5G
Descaled; cuticular pattern

            Syn.: Tomocerus (Pogonognathellus) elongatus Christiansen, K, 1964:672
Pogonognathellus nigritus from the USA
2011.10.09 © Bradford, A.
Pogonognathellus nigritus from the USA
2011.10.09 © Bradford, A.

            Syn.: Pogonognathellus nigirtus(sic) Soto-Adames, FN et Taylor, SJ, 2013:158 ISS
Species palespeciesgroup Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:487
Remark by Felderhoff, K.L. et al. 2010:488: We prefer the term "pale species group" to denote this group for convenience only, rather than "flavescens complex," to separate these North American taxa from the European species.
            Syn.: Tomocerus plumbea(sic) Packard, 1871 IOS
            Syn.: Tomocerus alba(sic) Packard, 1877 IOS
            Syn.: Tomocerus pallidus Packard, 1888
            Syn.: Tomocerus arcticus Schött, H, 1893:43
            Syn.: Tomocerus americanus Schött, 1896
            Syn.: Tomocerus niger Folsom, JW, 1902:97
            Syn.: Tomocerus flavescens var. separatus Folsom, JW, 1913:460
            Syn.: Tomocerus (Pogonognathellus) flavescens Christiansen, K, 1964:673
Remark by Christiansen, K. 1964:675: It is apparent that the species here classified as T. flavescens represents a vast complex of forms.

            Syn.: Pogonognathellus sp.2 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475
            Syn.: Pogonognathellus sp.3 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475
            Syn.: Pogonognathellus sp.5 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475
            Syn.: Pogonognathellus sp.6 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475
            Syn.: Pogonognathellus sp.7 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475
            Syn.: Pogonognathellus sp.8 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475
Pogonognathellus "flavescens" from the USA
2006 © Murray, T.
Pogonognathellus "flavescens" from the USA
2009 © Bernard, E.C.
Pogonognathellus "flavescens" from the USA
2010.05.24 © Taft, M.
2+4 frontal setae

Species taenitus (Koch, CL & Berendt, GC in Berendt, GC, 1845:119) -
Podura taenita from Baltic amber
After Koch CL & Berendt GC, 1845 Tab.XV Fig.126
            Syn.: Tomocerus taeniatus(sic) Weitschat, W et Wichard, W, 2002:86,- ISS

Genus Tomolonus Mills, 1949

Tomolonus reductus from the USA
2006 © Murray, T.
Species reductus Mills, 1949, t.t.


Genus Monodontocerus Yosii, 1955
Species absens Yu, Deharveng & Zhang, 2014
Species leqingensis Sun & Liang, 2009
Species modificatus (Yosii, 1955) Yosii, R, 1956:88, t.t.
  Subspecies satsumensis Yosii, R, 1956:89
Species mulunensis Yu, Deharveng & Zhang, 2014
Species odongnyeoensis Park, K-H et Lee, B-H, 1995:437
Species trigrandis Yu, Deharveng & Zhang, 2014

Genus Tomocerina Yosii, 1955 key
Species aokii (Yosii, R, 1972:94)
Species calcea Liu, Hou & Li, 1999
Species curta (Christiansen, 1964:662)
Species lamellifera (Mills, HB, 1934:84)
Species liliputana Yosii, 1967
            Syn.: Tomocerus (Tomocerina) lilliputanus(sic) Ma, Y-T, Chen, J-X et Christiansen, K, 2003:516 ISS
Species minuta (Tullberg, T, 1877:32) Yosii, R, 1956:91
Species purpurithora Liu, Hou & Li, 1999
Species simplex Yosii, 1966
Species teres (Christiansen, K, 1964:653)
Species tianshanensis Ma, Y-T, Chen, J-X et Christiansen, K, 2003:512
Species tridentatus Sun, Liang & Huang, 2007
Species yiliensis Ma, Y-T, Chen, J-X et Christiansen, K, 2004:43

Genus Aphaenomurus Yosii, R, 1956:86
Species interpositus Yosii, R, 1956:86, t.t.
  Subspecies denticulatus Yosii, R, 1956:87

Genus Plutomurus Yosii, R, 1956:78 key
Species sp.1 Pomorski, RJ et Steinmann, D, 2004:20
Species sp.2 Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:894
Species abchasicus Martynova, 1969
Species baschkiricus (Skorikov, 1900)
Species belozerovi Martynova, 1979
Species birsteini Djanashvili, R et Barjadze, S, 2011:28
Species borealis Suma, Y, 1981:502
Species brevimucronatus (Denis, 1928)
            Syn.: Tomocerus (Plutomurus) brevimucromatus(sic) Skidmore, 1995 ISS
Species californicus (Folsom, JW, 1913:469)
            Syn.: Tritomurus oregonensis Denis, 1928
Species carpaticus Rusek, J et Weiner, WM, 1978:742
            Syn.: Tomocerus terrestralis Paclt, 1957, nec Stach, 1922
Species diversispinus (Yosii, R, 1966:555) Jordana R & Baquero E in Jordana, R, Baquero, E, Reboleira, S & Sendra, A, 2012:78
Species edaphicus Yosii, 1967
Species ehimensis Yosii, R, 1956:83
Species grahami (Christiansen, 1980:121) Jordana R & Baquero E in Jordana, R, Baquero, E, Reboleira, S & Sendra, A, 2012:78
Species gul (Yosii, R, 1966:552) Jordana R & Baquero E in Jordana, R, Baquero, E, Reboleira, S & Sendra, A, 2012:78
Species iwatensis Yoshii, 1991
Species jeleznovodski Kniss & Thibaud, 1999
            Syn.: Plutomurus jeleznovozski(sic) Djanashvili, R et Barjadze, S, 2011:29 ISS
Species kawasawai Yosii, R, 1956:85
Species kelasuricus Martynova, 1969
Species leei (Yosii, R, 1966:554) Jordana R & Baquero E in Jordana, R, Baquero, E, Reboleira, S & Sendra, A, 2012:79
Species marmorarius Yosii, 1967
Species ortobalaganensis Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:77
Plutomurus ortobalaganensis from Abkazia
The deepest terrestrial animal living about 2 km below surface
2012 © Jordana R & Baquero E.

Species riugadoensis (Yosii, 1939) Yosii, R, 1956:78, t.t.
Species sorosi Kniss & Thibaud, 1999
Species suzukaensis (Yosii, 1938) Yosii, R, 1956:81
  Subspecies naikaiensis Yosii, R, 1956:83
  Subspecies ohminensis Yosii, R, 1956:82
Species unidentatus (Börner, 1901) Dobat, 1979, B
            Syn.: Pogonognathellus flavescens Dethier, M et Hubart, J-M, 2000:29,B
            Syn.: Plutomurus unidentaus(sic) Stomp, N et Weiner, WM, 1994:359 ISS
Species vigintiferispina Lee, 1974
Species wilkeyi (Christiansen, K, 1964:659) Jordana R & Baquero E in Jordana, R, Baquero, E, Reboleira, S & Sendra, A, 2012:79
Species yamatensis Yosii, R, 1956:84
  Forma albida Yosii R, 1956:85
  Subspecies kyushuensis Yosii, R, 1956:85

Genus Lethemurus Yosii, 1970
Species sp.1 Pomorski, RJ et Steinmann, D, 2004:20
Species finitimus Yosii, 1970
Species missus (Mills, 1949), t.t.
            Syn.: Tomocerus misus(sic) Cutz-Pool, LQ, Palacios-Vargas, JG et Vázquez, M, 2003:97 ISS
            Syn.: Tomocerus missus(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:110,ca ISS

Genus Entomocerus Christiansen, KA et Pike, E, 2002:170, -
Species mirus (Christiansen, KA & Pike, E, 2002:170, -) Christiansen, KA & Pike, E, 2002:170, -, t.t.

Subfamilia Lepidophorellinae Absolon K, 1903
Lepidophorellinae from Tasmania
2013.05.25 © Zosterops, K.
Lepidophorellinae from Tasmania
2013.05.25 © Zosterops, K.
Lepidophorellinae from Tasmania
2013.06.19 © Zosterops, K.
Lepidophorellinae from Tasmania
2013.06.19 © Zosterops, K.


Tribus Lepidophorellini Womersley, 1939

Genus Lepidophorella Schäffer, 1897
Species australis Carpenter, GH, 1925:11
Species brachycephala (Moniez, 1894) Denis, 1923
            Syn.: Lepidophorella brachycephalla(sic) Womersley, H, 1936:321 ISS
Species communis Salmon, JT, 1937:353


Lepidophorella communis from New Zealand
After Salmon, J.T. 1937 Pl.49 Fig.d.
Lepidophorella communis from New Zealand
After Salmon, J.T. 1941 Pl.54 Fig.246.

Species flavescens (Nicolet, H, 1847:383) Schäffer, 1897
            Syn.: Cyphoderus flaveus(sic) Nicolet, H, 1847:383 IOS
            Syn.: Cyphoderus giganteus Nicolet, H in Gay, C, 1849:92
Cyphoderus giganteus from Chile
After Nicolet H in Gay C, 1854 Lam.1 Fig.7.

            Syn.: Lepidophorella flava Schäffer, 1897,tt
            Syn.: Lepidocyrtus flaveus(sic) Lubbock, J, 1873:157 ISS
            Syn.: Cyphoderus flaveus(sic) Salmon, JT, 1964:536 ISS
Remark by Ellis, W.N. & Bellinger, P.F. 1973:27-28: Cyphoderus flaveus Nicolet, 1847 is an incorrect original spelling (art.32a(ii)). Nicolet, 1847:383 described the new species Cyphoderus giganteus and C. flaveus. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]:92-93. In the last paper however "flavescens" is used in stead of "flaveus". But in the first paper Nicolet cites as author of both species Nicol. in Gay, evidently the publications appeared in a reversed order. This, and the fact that "flaveus" is bad latin, brings us to the conclusion that "flaveus" is an incorrect original spelling.


Species fusca Salmon, JT, 1941:340

Lepidophorella nigra from New Zealand
After Salmon, J.T. 1943 Pl.42 Fig.48.
Species nigra Salmon, JT, 1943:383
Species rubicunda Salmon, JT, 1941:342
Species spadica Salmon, JT, 1944:152
Species tiegsi (Womersley, H, 1942:26)
Millsia tiegsi from Australia
After Womersley, H., 1942 Fig.3A
Remark by Greenslade, P. in Dallai, R. 1989:110: The type series consists of three specimens, all probably early instars. Millsia is similar to Lepidophorella and differs only in the lack of scales and the shorter, more cylindrical dens. It is possible that Millsia is a synonym of Lepidophorella.

Species unadentata Salmon, JT, 1941:341

Genus Pseudolepidophorella Salmon, JT, 1941:342

Species longiterga (Salmon, JT, 1937:354) Salmon, JT, 1941:343, t.t.


Lepidophorella longiterga from New Zealand
After Salmon, J.T. 1937 Pl.49 Fig.h.
Pseudolepidophorella longiterga from New Zealand
After Salmon, J.T. 1941 Pl.54 Fig.259.


Genus Antennacyrtus Salmon, JT, 1941:343

Antennacyrtus insolitus from New Zealand
After Salmon, J.T. 1941 Pl.55 Fig.261.
Species insolitus Salmon, JT, 1941:344, t.t.


Genus Neophorella Womersley, 1934
Species dubia Womersley, 1934, t.t.

Genus Lasofinius Ireson & Greenslade P, 1990
  Syn.: Lepidophorellinae Lasofinus(sic) Ma, Y-T, Chen, J-X et Christiansen, K, 2004:41 ISS
Species gemini Ireson & Greenslade P, 1990:211
Species willi Ireson & Greenslade P, 1990:209, t.t.

Tribus Novacerini Salmon JT, 1941:345

Genus Novacerus Salmon, JT, 1942:259
Remark by Salmon, J.T. 1942:259: Novacerus Salmon JT, 1942 nomen novum to replace the name Neocerus Salmon, 1941, pre-occupied by Wasmann, 1893.

  Syn.: Lepidophorellinae Novocerus(sic) Ma, Y-T, Chen, J-X et Christiansen, K, 2004:41 ISS
Species insolitatus (Salmon, JT, 1941:346) Salmon, JT, 1942:259

Neocerus spinosus from New Zealand
After Salmon, J.T. 1941 Pl.55 Fig.269.
Species spinosus (Salmon, JT, 1941:345) Salmon, JT, 1942:259, t.t.

Species tasmanicus (Womersley, H, 1937:154) Yosii, 1967
Tomocerus tasmanicus from Tasmania
After Womersley H, 1937 Fig.1g




References